59

28.09
98

1.

..

1.

1.
:

1.
2.
3.
4.
5.

.. ,

..

.
.
.
.

. .

7. .
8. .
9. .
10. .
11. .

..
-
:

Microleptinae

Ichneumonidae) 1 .. ;
, 2003.- 175 .; .
ISBN 5-02-00338-

2.
1.

-
Ichneumonidae (Hymenoptera).
- Micro\eptinae s. 1.

MICROLEPTES Gravenhorst, 1829

75
7

77
78
79
79
80
81
81
82
82

CUSHMANIA Dasch, 1992

psarevi Kasparyan et Huma1a,

3.

MICROLEPTINI Townes, 1958

aquisgranensis (Foerster, 1871) ......................................................

belokobylskii Humala, sp. n.............................................................
grandis Humala, sp. n......................................................................
minor Humala, sp. n. .......................................................................
obenbergeri Gregor, 1938 ...............................................................
orientalis Humala, sp. n...................................................................
rectangulus (Thomson, 1888) ........................................................
salisburgensis Schwarz, 1991..........................................................
spasskii Humala, sp. n. .....................................................................
splendidulus Gravenhorst, 1829 .......................................................
tibialis Humala, sp. n. ........ ......... .................... ...... .... .... ...... .... ...... ...

Oxytorinae (Hymenoptera:

[. . .. ]- .:

MICROLEPTINAE Townes, 1958

199

..................................................

84

HYPERACMUS I Holmgren, 1856

1. . crassicornis (Gravenhorst, 1829) .......................................................

85

- .

Oxyrorinae

2.

Ichneumonidae,

. Helictinae, Cyl\ocerinae
Diacritinae Microleptinae, Diacritini
. Pimplinae Cylloceriini.
; : .
J1J5:

29 220 -
1 ;
1 .

, .

2003(1)-

ISBN

12

5-02338-

1.

200J

""
( ),

2003

.
.
.
.
.

CYLLOCERIINI Wahl, 1990

CYLLOCERIA

Scfii~dte,

1838

aino (Uchida, 1928)..........................................................................

alpigena (Stro, 1901) ....................................................................
boealis (Roman, 1924).....................................................................
brachycera Humala, 2002................................................................
fusciventris (Hellen, 1940) ...... ....... ...... .. .... .. .. ..... ......... ............ ..... ...
. . impespicua Rossem, 1987 ..............................................................
7. . invicta Rossem, 1987 .......................................................................
8. . melancholica (Gravenhorst, 1820) ...................................................
9. . occupato (Gravenhorst, 1829) ........................................................
10. . oientalis Humala, 2002 ...................................................................
11. [. sueinensis (Brauns, 1905)] ............................................................
12. . sylvestris (Gravenhorst, 1829)..........................................................
1.
2.
3.
4.
5.

91
91
91
91
92
92
92
92
93
93
93
93
3

13. . tenuicornis Humala, 2002 ............................................................... .

14. . ussuriensis Humala, 2002 ................................................................ .

93
93

2. ALLOMACRUS Foerster, 188

1. . arcticus (Holmgren, 1881) ............................................................... ..
2. . jakuticus Humala, 2002 ..................................................................... .
3. . subtilis Humala, 2002 ........................................................................ .
4. . volcanus Humala, 2002 ..................................................................... .

94
95
95
96

3. ROSSEMIA Humala, 1997

1. R. longithorax Humala, 1997 ................................................................ ..

96

4. ENTYPOMA Foerster, 18897

1.
2.
3.
4.
5.
6.

. jel'ale Rossem, 1988 ......................................................................... ..

E.frontosum Rossem, 1988 .................................................................. ..
. prominens Kolarov, 1985 ................................................................. ..
. robustator Aubert, 1968 .................................................................... .
. robustum Foerster, 1871 .................................................................... .
. suspiciosum (Foerster, 1871) ............................................................. .

97
97
98
98
98
98

5. . fulvicornis Foerster, 1871.................................................................

6. P.fulvipectus Foerster, 1871 .................................................................
7. . paganus (Haliday, 1838) ..................................................................
8. . praetor (Haliday, 1838)....................................................................
9. . ,-udis Foerster, 1871..........................................................................
10. . subsulcatus Foerster, 1871 ........................................... ;...................
11. . zonatus (Gravenhorst, 1829) ....................... ~ .....................................

5.
99

3. HELICTINI Gupta, 1987

1.

ANISERES Foerster, 1871

1. . caudatus Humala, 1997 ....... .... .. .. .. .. .... .... .. . .. . .. ......... ........... .... ... .... .. .. 103
2. . pallipes Foerster, 1871 ....................................................................... 103
3. . paadoxus Rossem, 1991.... .. .. .. .. .. ... ..... ... . .......... . .. .................... ....... .. 103

dispa Rossem, 1980 ....... ... ... .... ... .... ....... .. .......... .. ....... ......... ....... ......
gracilis Rossem, 1987 ........................................................................
lubricus (Foerster, 1871) ....................................................................
luridus Foerster, 1871.........................................................................
rudepunctatus Strol, 1903.................................. .. .. ...... ... .. ..... ..... ......

.
.
.
.

ardentis Rossem, 1987 .....................................................................

attentus Foerster, 1871 .................................................................... :
comes (Haliday, 1838)......................................................................
edwardsi Roman, 1923 .....................................................................

HEMIPHANES Foerster,

188

7.

MEGASTYLUS

Schidte,

115
- 116
116
116
116
116
116

1838

1. MEGASTYLUS Schi0dte, 1838

104
104
104
105
105

3. PROCLITUS Foerster, 188

1.
2.
3.
4.

111
112
112
112
112
112
112
112
113
113
113

PANTOMIMA Rossem, 1990

1. . flavipes Foerster, 1871 .......................................................................

2. . gravator Foerster, 1871 ......................................................................
3. . l10rtense Rossem, 1987 ......................................................................
4. . inusitatum Rossem, 1987 ...................................................................
5. . montanum Rossem, 1987 ...................................................................
6. . performidatum Rossem, 1988 ............................................................
7. . townesi Rossem, 1980 ........................................................................

2. PANTISARTHRUS Foerster, 1871

1. .
2. .
3. .
4. .
5. .

99
99

189

1. P.festata Rossem, 1990 .......................................................................... 114

APOCLIMA Foerster, 1871

1. . haeselbarthi Rossem, 1987.................................................................

2. . signaticorne Foerster, 1871 ................................................................

APERILEPTUS Foerster,

1. . alpalpus (Gravenhorst, 1829) ........................................................

2. . flavus Foerster, 1871 ........................................................................
3. . imp-us Foerster, 1871...................................................... ...............
4. . infuscatus Foerster, 1871..................................................................
5. . melanopsis Foerster, 1871................................................................
6. . microspilus Foerster, 1871 ........................................................... ....
7. . plagiatus Foerster, 1871 ...................................................................
8. . ossemi Jussila, 1994. ..... ... ........ ... .. .. .. .. .. .. ........ ... ..... .... .. ... ... ... ... . .. ...
9. . tricinctus Foerster, 1871...................................................................
10. . vanus Foerster, 1871.........................................................................
11 . . viduatus Foerster, 1871 .....................................................................

5. KENTROTRYPHON Strol, 1902

1. . longecaudatus Strol, 1902. .... .. .......... ... ......... .......... .... .. ... ... ... .. .... .. ..

4.

109
109
109
109
110
110
110

107
107
108
109

1.
2.
3.
4.

. (.) cruentator Schipdte, 1838 .........................................................

. (.) flavopictus Gravenhorst, 1929 ...................................................
. (.) ortator Schipdte, 1839 ............................................................
. (.) suecicus Rossem, 1983 ..............................................................

2.
5. .
. .
7. .
8. .
9. .

119
120
120
120

DICOLUS Foerster, 1868

(D.) elegans Rossem, 1983 ................................................................

(D.) excuitor Foerster, 1871.............................................................
(D.) impresso Schipdte, 1838...........................................................
(D.) pectoalis Foerster, 1871 ............................................................
(D.) tenellus Rossem, 1983................................................................

121
121
121
121
121
5

8.
.
.
.
.
.
. .
7. .

1.
2.
3.
4.
5.

HELICTES Haliday, 1838

borealis (Holmgren, 1855) ...... .. .... ...... .. .. .. .... .............. ...... ...... ...........
carinata Humala, sp. n. ......................... .. ...........................................
conspicua (Foerster, 1871) .................................................................
erythrostoma (Gmelin, 1790).... ..... .. .... .. .. .. .... .. .. ... ...... .. ..... .. .. .... ... .....
fabularis Rossem, 1987.. ... ...... .............. ..... .... ... .... .. ... ... .. .... .... .... ..... .. ..
incongruens Rossem, 1987 .. ........ ............... .............. .... .. .. .... .. .... ...... .
karelica Humala, sp. n ...................................................... :................

9.

122
123
123
123
123
123
123

FETIALIS Rossem, 1990

33. .
34. .
35. .
36. .
37. .
38. .
39. .
40. .
41. .
42. .

(.)

prognathor Aubert, 198.................................. .. .......................

spuria Rossem, 1991 ................... ;.............................................
(.) subangulata (Foerster, 1871).....................................................
(.) substantiva Rossem, 1987 ................ .........................................
(.) subteres (Thomson, 1888) ......................................... ................

(.)

(.) tene (Foerster, 1871) ................................................................

(.) tenuicornis (Foerster, 1871) ......................................................
(.) terebrator (Foerster, 1871) ........................................................
( . ) vagator (Foerster, 1871) ............................................................
(.) ventosa Rossem, 1987 ...............................................................

10.
1.
1. .
2. .
3. .
4. .
5. .

(.)

7. .
8. .
9. .
10. .
11. .
12. .
13. .
14. .
15. .

(.)
(.)
( . )
( . )
(.)
(.)
(.)
(.)

agitator (Foerster, 1871) ........................................................... 129

(.) amicalis (FoersteF; 1871)........................................................... 130
(.) aquilonia Humala, sp. n. ........................................................... 130

cinctula (Foerster, 1871) ...........................................................

collaris (Gravenhorst, 1829) .....................................................
conjuncta (Foerster, 1871) ........................................................
connexa (Foerster, 1871) ...........................................................
crassicornis (Foerster, 1871) .............................................. .. .....
deterior (Foerster, 1871) ...........................................................
erythropyga (Foerster, 1871) .....................................................
eurystigma (Foerster, 1871).................................... ...................
(P.)foersteri Rossem, 1987 ..............................................................
1. . (.) fraterna (Foerster, 1871) ...........................................................
17. . (P.)fuscifemur Humala, sp. n ................................................ ...........
18. . (.) helleni Humala, sp. n .................................................................
19. . (.) helvola (Foerster, 1871).............................................................
20. . (.) humeralis (Foerster, 1871) ........................................................
21 . . (.) indomita Rossem, 1987 .............................................................
22. . (.) koponeni Humala, sp. n. ............................................................
23. . (.) melanocera (Foerster, 1871) .....................................................
24. . (.) mendica (Foerster, 1871) ...........................................................
25. . (.) mesoxantha (Foerster, 1871) ...... ...............................................
2. . ( . ) moerens (Foerster, 1871) ...........................................................
27. . (.) monticola (Foerster, 1871) ........................................................
28. . (.) nava (Foerster, 1871)..................................................................
29. . (.) nemorensis Rossem, 1987 .........................................................
30. . (.) obscura Rossem, 1991 ............................................................ ;.
31. . (.) parvula (Foerster, 1871) ............................................................
32. . (.) posticata (Foerster, 1871)..........................................................

11.

PLECTISCIDEA Viereck, 1914

(.) istriata (Thomson, 1888).........................................................

(.) landita Rossem, 1987 ..............................................................
. . (.) canaliculata (Foerster, 1871) ....................................................

FUGATRIX Rossem, 1987

43 . . (F.) communis (Foerster, 1871) ............................................. ........... 134

PLECTISCIDEA Viereck, 1914

2.

1. F. alacris Rossem, 1990 ......................................................................... 124

133
133
134
134
134
134
134
134
134
134

130
130
130
130
130
131
131
131
131
131
131
131
131
131
131
132
132
132
132
132
132
132
132
133
133
133
133
133
133

DIALIPSIS Foerster, 1868

1. D. exilis Foerster, 1871 ........................................ ................................... 135

12.

ATABULUS Rossem, 1988

1. A.faustus Rossem, 1988 .......................... ~ .............................................. 13

13.

CATASTENUS Foerster, 1868

1. C.femoralis Foerster, 1871 .....................................................................

14.

SYMPLECIS Foerster, 1868

1. S. beaumontor Aubert, 1968 ............................... ....................................

2. S. icingulata (Gravenhorst, 1829) .........................................................
3. S. breviuscula Roman, 1923 ...................................................................
4. S. invisitata Rossem, 1980 ............................. .. .......................................
5. S. paradoxa Rossem, 1988 ......................................................................

15.

137
137
138
138
138

PHOSPHORIANA Rossem, 1988

1. . rugosissima (Strol, 1903) ................................................................. 139

16. GNATHOCHORISIS Foerster, 1868

1. G. crassula (homson, 1888) ............................... .'.................................
2. G. dentifer (Thomson, 1888)...................................................................
3. G.flavipes Foerster, 1871 .......................................................................
4. G. meridionato Aubert, 1980.................................................................
5. G. restricta-(Rossem, 1980) ..... ............ ... .. .... ..... .. ... .. ... ............ ........ ...... .
. G. xanthocephala (Strol, 1903).............................................................

17.
1.

139
139
140
140
140
140

EUSTERINX Foerster, 1868

EUSTERINX Foerster, 1868

1. . (.) argutula Foerster, 1871 ...... ....................................................... 143

2. . (E.)fabulosa Rossem, 1990 ............................................................. 143
7

.
.
.
. .
7. .
8. .

3.
4.
5.

(.) fleischei Gregor, 1941..............................................................

(.) jugoum Strol, 1900.................................................................
(.) obscuella Foerster, 1871..........................................................
(.) oligomea Foerster, 1871 ..........................................................
(.) pseudoligomera Gregor, 1941 ...................................................
(.) subdola Foerster, 1871..............................................................

2.

143
143
143
143
144
144

ISCHYRACIS Foerster, 1868

9. . (1.) blspinosa Strol, 1900 ................................................................ 144

10. .(!.) permil-anda Rossem, 1988.......................................................... 145

3.
11. .
12. .
13. .

.
.

(.)
(.)

Rossem, 1982 .............................................................. 145

tifasciata (Aschmead, 1899) .................................................... 145
tichops (Thomson, 1888) ....................................... .................. 145

(.)

(.)
(.)
(.)
(.)
(.)

DALLATORREA Ashmead, 1902

HOLOMERISTUS Foerster, 1868

aquilonigena Rossem, 1982......................................................

minima (Strol, 1903) ...............................................................
refactaia (Rossem, 1982) .......................................................
similis Rossem, 1991 ................................................................
tenuicincta (Foerster, 1871) ......................................................
truculenta Rossem, 1991...........................................................

18.
1.
2.

DIVINATRIX Rossem, 1987

(D.) circaea Rossem, 1982 ............................................................... 146

6.
17. .
18. .
19. .
20. .
21 . .
22. .

(D.) inaequalis Rossem, 1980 .......................................................... 145

(D.) inaspicua Rossem, 1988 ........................................................... 145

5.
16.

TRESTIS Foerster, 1868

(.) tart-ea

4.
14.
15.

14

14

147
147
147
147

PROELIATOR Rossem, 1982

invictus Rossem, 1987 ........................................................................ 148

Rossem, 1982....................................................................... 148

. propius

4.

DIACRITINI Townes, 1965

1. DIACRITUS Foerster, 1868

1. D. aciculatus (Vollenhoven, 1878) .... .... .. .. .. .. .... ...... .... .... .. ......... .... .. ... ... 149
2. D. incompletus Momoi, 1966.................................................................. 149

2.
1.

ORTHOLABA Towiles, 1969

tenuis Townes, 1969 ........................................................................... 151

11.

1.
1.
2.
3.
4.
5.

.
.
.
.
.

. .
.
.
.

7.
8.
9.
10.

OXYTORINAE Thomson, 1883

OXYTORUS Foerster, 1868

armatus Thomson, 1883 .. .. .. .... .. .. .... ... ... ..... .. .. .... .. ..... .... .... ...... .. .... ..
canalis (Momoi, 195)................... ... .. .. .. .. .. ... ... .. ... .. .. .. .. .. .. .. ..... .. ... ..
confusus Humala, sp. n. ...................................................................
corniger (Momoi, 195) ..................................................................
kamikochianus (Momoi, 195) ........................................................
luridator (Gravenhorst, 1820)..........................................................
montanus (Momoi, 195).................................................................
nikkoensis (Momoi, 1965)................................................................
norikuranus (Momoi, 1965).............................................................
obtusus (Momoi, 1965) ....................................................................

154
154
155
15
15

15

157
157
157
157

Micro1eptinae sensu 1ato

..

( 1905) .
.. (1934, 1936-), ,
21 , ,

(Schmiedeknecht, 1930)

~ . "

" (,

Micro1eptinae s. 1. (= Micro1eptinae sensu Townes,

Oxytorinae sensu Rossem) ,

Ichneumonidae.

- .

, .

(Diptera): Nematocera

Brachycera

Orthohapha

-,

.
.

,

Ichneumonidae.

Scutellator

1995).

Plectiscidea

(,

(Wahl, 1990; Gau1d, 1991; Wah1, Gau1d, 1998).

(Townes, 1944, 1945, 1969, 1970, 1971; Townes
et 1., 1961, 1965; Townes ., Townes, 1966, 1973; Yu, Horstmann,

1997). .
(Rossem, 1974, 1980-1991)
.

, ~,

"wastebasket group" (
), . (Townes, 1971).

10

1981)

Microleptinae

, .

Oxytorus,

Oxytorinae.

(Townes, 1971),

. ,

29

200

Diacritinae
Micro1eptinae.

(Diptera, Mycetophilidae s. 1.),

,
.

70

21

, ,

8000

~ .

,
- .

. .

( 1984) .
-10, ZEISS Leica
(WILD-8, WILD-10).
.

,

,
(

. )

11

,
)
(

. ), )

(Nordic Counsil of Ministers).

Microleptinae

... .. ( ).

Oxytorinae

. ..
.. ( )

Mycetophilidae),

(Diptera,

2-15

, ,

3-5

Cylloceria, Hyperacmus

Oxytorus (6-15 ) Megastylus Eusterinx (

1 ). )

),

Microleptini,

,
)
,

. )
, ,

, ,
) , ,

), )
,

, ,
.

. , ,
). . ,

) .

Trestis

Eusterinx Helictes
Microleptini

(OOL)-
)

(POL)

13

 .
.

.
;

.
.

, , ,
, ,

Apoclima, Hemiphanes, Aperileptus

Hyperacmus, Rossemia,
Microleptes.

Microleptes

2.
1 - Eusterinx (Holomeristus) sp., 2- Dialipsis exilis

Apeileptus

, .

Allomacrus

Dallatoea)

3 , Cylloceia, Entypoma,
Poeliator, Eusterinx, Proclitus
. ,
(. . 2, 1)

Eusteinx:

Eusterinx

Symplecis, Catastenus,
Divinatrix, Trestis, Ischyacis,

Microleptini (Microleptes, Cushmania,

Hyperacmus) Megastylus
(. 1).
, , ,

Kentrotryphon,

Gnathochorisis

-.

Microleptes rectangulus

, ;

, Allomacus

Proclitus

volcanus

.
, -

. ,

D ial ips is

, ,

.
: Microleptes,
Proclitus, Gnathochorisis, Pantisarthrus . ,

(. .

(.

2, 2),

2, 1),
Cushmania, Cylloceria,
Allomacrus, Rossemia, Helictes, Atabulus, Plectiscidea,
Proclitus). , Entypoma, Plectiscidea,
Aperileptus, ,

1.

(:

1-

Microleptinae,
Townes, 1971)

2 - ; 3 5 - ; -

14

Microleptini,

Cylloceria

, ,

4-

( antisarthrus)

7- ; 8- ; 9-
; 1 -
; 11 - ,

.
,

Diacritus
Ortholaba

15

3.

()

.
,
,
.

14-30 .

Microleptini Eusterinx

3.

Eusterinx tenuicincta).

Microleptinae

1 - Apoclima sp., 2 -. signaticorne, 3 - Cylloceria borealis, 4 - Helictes karelica sp. n., 5- Helictes carinata sp. n., 6- Microleptes splendidulus, 7- Microleptes orientalis sp. n., 8- Hemiphanesflavipes; : 9- Apoclima sp., 10Helictes karelica sp. n.

(Cylloceria borealis,

MicJ-oleptes, Cushmania, Eusterinx Cylloceria.

Anisees Pantisathus- ,
-

'

Eusterinx, Megastylus

Plectiscidea.

(. .

Helictes,

Cylloceria,

3, 6-8)

(. .

16

Microleptini,

3, 1-5).

17

 , ,

()

- (.

, ,

- . ,

(Plectiscidea,
Aperileptus, Atabulus).
Cylloceria, Allomacrus, Apoclima, Pantomima, Entypoma, Symplecis,
Gnathochorisis, Phosphoriana Eusterinx.

Cushmania ( Microleptini), Allomacrus (

Cylloceriini), Diacritus, Ortholaba ( Diacritini), Aniseres,
Pantisarthrus,
Aperileptus,
Proclitus,
Fetialis,
Megastylus,
Phosphoriana, Symplecis, Gnathochorisis ( Helictini).

Cylloceriini-

3-

4-5, ).
Microleptini.

( Rossemia 5-9- , Kentrotryphon

); Helictini - 5- -

Eusterinx, Cyllocel'ia).

8-9-.

Eusterinx (Holomeristus) - 6-9(10)(11)(12) Hemiphanes (8)(9)10(11)(12). Microleptini Microleptes

1-2(3)- 5-8- (. . 3, 6),
Hyperacmus - 4-5- .

Cylloceria, Apoclima ( Cylloceriini), Hyperacmus (
Microleptini) Eusterinx, Helictes ( Helictini);

11-12-

(.

3, 1-5).

Cylloceria

Apoclima

Pantomima

..

Aniseres,

Proclitus

Eustel'inx

Pantomima
(Pimplinae),

Polysphinctini

, -, .

Pl-oclitus

, ,

.
,
, (

; 4- (. . 3, 1-3).
Cylloceria Apoclima

Microleptini, Proeliator).
4, ) ,
Rossemia, Atabulus Aperileptus, '. vanus,

( ) .

3- ,

(.

, ,

(. .

3, 7).

Helictes

(, ,

1996).
Megastylus, Helictes Hemiphanes

.
{. .

5)

() ,

( , )

. ,

Cylloceriini,

Orthocentrinae, -
Gnathochorisis ( Helictini).

. ,

()

1-3- . ,
,

; -
,

1-3- .

, ,

18

Eusterinx
.

19

n .

Aperileptus

nn, .

(.

Gnathochorisis, Eusterinx,

4-5,

Helictini;
): Catastenus, Symplecis,

, .

Microleptini

Microleptes,

.
(),

Eusterinx ( Dallatorrea, Ischyracis, Trestis),

Schizopyga Oxytorus armatus. Microleptini
Microleptes,

.
,
, ,

Helictes, Eusterinx, Plectiscidea.

Helictes

Oxytorus

. ,
, .

Microleptini,
Gnathochorisis,
Eusterinx, Cylloceria orientalis Catastenus femoralis
. Proclitus,
antisarthrus .
, .

(. .

. 4-5. Microleptinae. (4 - ; 5 - )

1-33 -, a-n- : 1-3 - : 1 - ,

2- , 3- () ; 4, 5-
: 4 -' ~ , 5 - ; 6- ;
7- ; 8- ; 9-15- (): 9- ,
1 - , 11 - , 12 - () , 13 -
, 14- , 15- ; 16- ; 17-21: /7- ; 18- (
); 19- () ; 20- ; 21-
; 22 - ; 23 - ; 24-33 - : 24 - 1-
, 25 - 2- , 26- 3- , 27- 1-
, 28- 2- , 29- 3- , 30-
, 31 - , 32 -, 33- . -
; - ; - ; d- ; -
; ~ - ~ ; g - ; h -

28, 3, 4),
Hyperacmus. egastylus
2/3,
(. . 28, 3). Pantomima
. Entypoma Catastenus
femoralis .

Cylloceria

, ,

(. 6), ,

; t - ;

20

j- ; k-
; l-

n-

21

(Gauld, Fitton, 1987).

.
: ,

, Microleptes, Hemiphanes Megastylus

( Dicolus) - .

Oxytorinae

Helictini

. 6. Microleptinae ( - , - )
(1-9): 1- , 2- , 3- , 4-
, 5 - 2- , 6 - , 7 - , 8 - ,

9 - ; : - , Sc + R - , Pt -
, R - , Rs- , (bas) - , - , 1rm, 2rm,

r- , + - , 1 - 1- ,
2 - 2- , - , (par)- , nrv-
, pnrv - , 1 - 1- , hm - , nrl - , (: Townes, 1969 )

),
. ( Microleptini, Cylloceria,
Oxytorus) ,
. Oxytorinae

.

, .

- -.

Cylloceria

Hyperacmus

, Helictini), ,
, _, .

Oxytorinae ,
,

Oxytorus , . !urida-

tor

,
;
Allomacrus, Entypoma, Symplecis, Gnathochorisis
Eusterinx.
(nn)
( Megastylus, Hemiphanes, Eusterinx)~
( Helictini)- .
Microleptini , Cushmania
.

22

, 2-
,
.

Microleptini,

,
4-6- ,
.

, ,
.

2-

3-

.
-

23

Cylloceria, Rossemia, Allomacrus, Apoclima,

Kentrotryphon, Entypoma, Aniseres, PantisarthJ-us, Aperileptus,
Pantomima , Hyperacmus
, .. ,

, .
.

Plectiscidea

Atabulus

, .

Orthopelmatinae,

, .

.

,
;

Aperileptus, Proclitus, Aniseres,

- Plectiscidea , Dialipsis,

Gnathoch-isis .

, ,
,

(
).

Plectiscidea,

Proclitus.

7.

1 - Aperileptus a!ipalpus, 2 - Cylloceria melancholica, 3 - Po clitus comes, 4 Apeileptus ossemi

.
(Hemiphanes, Pantisarthrus, Helictes, Megastylus,

Symplecis)

. ,

. -,
, ,
. ,
,

(.

(Helictes, Megastylus, Pantisarthrus,

Eusterinx)

7).

Aperileptus

lectiscidea.

Proclitus

(. . 7, 1 ,4). Microleptini, Aperileptus,

Proclitus Plectiscidea,
.
Helictes Megastylus, ,

, ,

:.

3-4

. ,

(Cylloceria sylvestris,

Aniseres caudatus, Proclitus comes, PlectisCidea communis, Aperileptus

rossemi .); , ,
24

Eusterinx, Helictes, Megastylus

, -

25

Oxytorus,

, ,
,
(

Ctenopelmatinae);
Oxytorus
(Quicke et al., 1994).

, . (Quicke et al., 1994),

Orthocentrinae sensu lato

Ichneumonidae (. 8). Microleptinae s.l., Diplazontinae
Orthopelmatinae,
Pimplinae.

( ),
(Microleptes, Cylloceria)
(Eusterinx, Aperileptus).
,

Orthopelmatinae
Mesochorus).

(Orthopelma)

Oxytorinae

Microleptinae

Orthocentrinae,
Mesochorinae (Cidaphus,

.

,,

11

12

13

. 8. (: Quicke et al., 1994)

1 - Oxytorus (Oxytorinae), 2 - Microleptes, 3 - Cylloceria, 4 - Diacritus, 5 A~erileptus, 6- Orthocentrinae, 7- Eusterinx, 8- Mesochorus (Mesochorinae), 9Czdaphus (Mesochorinae), 10- Genus indet. (Microleptinae s. 1.), 11 - Genus indet.
(~icroleptinae s. 1.), 12 , 13 - Orthopelma (Orthopelmatinae), 14 - Diplazon
(Dtplazontinae)

Microleptinae.
(Megastylus, Gnathochorisis,
Symplecis), (Proclitus, Helictes, Megastylus),
(Gnathochorisis), , ,
(Cylloceria, Plectiscidea), .

,
,
,

26

27

Cylloceria

.
,

Megastylus, Eusterinx, Hemiphanes, Gnathochorisis,

Symplecis, Cylloceria.
Divinatrix Eusterinx
2-3- ,

.
;

Diplazontinae

9.

1 - Symplecis blcingulata, 2 - Cylloceria melancholica

,
.

Microleptini , ,
.

Eusterinx, Hemiphanes, Megastylus,

Cylloceria, Gnathochorisis, Symplecis ,
(. 9).

28

Pimplinae.

n, .

(Holmgren,
1881; Thomson, 1888; Viereck, 1914; Perkins, 1962, .).
Plectiscoidae
: (Foerster, 1868).: ,
, ,

, , -
,

.
, , ,

XVIII -

XIX

, , , ,

ey

(Gmelin in Linnaeus, 1790), Cylloceia accusato (= melancholica) (Fabricius, 1793), Oxytorus luidator, Cylloceia melancholica
(Gravenhorst, 1820) ( Ichneumon), s

cassiconis (

Helictes

thostoma

Exochus), Cylloceia caligata (

Phytodietus), . sylvestris ( Tryphon), Microleptes splendidulus
( Micoleptes), Poclitus zonatus, Plectiscidea collaris,
Apeileptus alipalpus, Megastylus flavopictus ( Plectiscus),
Symplecis icingulata ( Mesoleptus) (Gravenhorst, 1829),
Megastylus cruentato, . impesso, . oitator (
Megastylus) (Schiodte, 1838), Eusteinx pusilla (= tichops) (
Tryphon) (Zetterstedt, 1838), Proclitus comes, . paganus,
. t ( Cryptus (Clepticus)) (Haliday, 1838).
, ,

Allomacrus

Ischyacis

(=

Eusteinx)

Hemiteloidae, Symplecis Gnathochoisis Campoplegoidae,

Microleptes - Ichnemonoidae, Hypeacmus - Exochoidae,
Lamponota (= Cylloceria) Lissonotoidae,
Diacitus- Phaeogenoidae.

Symplecis

Gnathochorisis

: Anisees, antisathus,

Entelechia (= Entypoma)

Symphylus (= Acrodactyla) (Foerster, 1871).

Cylloceria, Proclitus, Megastylus,

~
, .

(Gravenhorst, 1829; Schiodte, 1838) -

Plectiscus, Cylloceria, Megastylus,-

Micoleptes,

:, ,

(Foerster, 1868, 1871;

Thomson, 1888; Schmiedeknecht, 1907, 1911, 1925, 1930 .).
(

Plectiscoidae,

Microleptinae

25)
(Foerster, 1868, 1871).

,
. , ,
. ,

,

30

.).

Hypeacmus. , ,

Micoleptes,

Helictes,

(Ate leute, Polyaulon, Symphylus, Adelognathus

Plectiscides Plectiscina
Tryphonidae (Thomson, 1888), ,
12 . : Allomacus hidias

(= Diacitus) .
.
1 Allomacus,

Symplecis,
Blapticus Catastenus,
,

,
.

Testis, Eusteinx

Holomeistus (= Eusteinx),

, ,

, . 3
n Plectiscus, ,

. 4

Megastylus, Miomeris (= Micoleptes) Hemiphanes,

31

.
Blapticus (= Entypoma, Entelechia,

Megastylus (= ldioxenus, Myriarthrus) ,

Aperileptus, Dialipsis, Proclitus, Pantisarthrus
Plectiscus Helictes, Dicolus Megastylus.
Ateleute Polyaulon,

Gnathochorisus)

Microleptes s. str.
Exochini Exochides
n Tryphoninae s. 1. (Morley, 1911; Schmiedeknecht, 1925,
1930). Microleptes
(= Miomeris) (Thomson,
1888; Dalla , 1902; Schmiedeknecht, 1911; Morley, 1914).
Hyperacmus

, ,

Symphylus (recte Acrodactyla) -

Polysphincta - ,

n .

Diacritus

(Foerster, 1868).

n (

, ).

Pimplinae (Vollenhoven, 1878),

(Thomson, 1888),
, Plectiscides
Allomacrus. (Morley, 1908)
Xoridides . Pimplinae,. ,

Orthocentrinae

, ,

Blapticus

Phidias),

Plectiscides.

(= Gnathochorisis).

(Schmiedeknecht, 1925, 1930)

Plectiscini . Ophioninae.

(Dalla , 1901-1902).

Plectiscinae

, ,

, .
,

. "Opuscula Ichneumonologica"
(Schmiedeknecht, 1911),
Allomacrus Phidias Plectiscina. ,
, ,

Adelognathina ,
: Adelognathus (Adelognathinae), Ateleute, Polyaulon (Cryptinae)
Symphylus = Acrodactyla degener (Pimplinae, Polysphinctini).
Cylloceria

Lissonotides (Foerster, 1868; Thomson, 1881; Dalla , 1902;

Morley, 1908, .) Lissonotini (Schmiedeknecht, 1907, 1930).
Allomacrus , ,

(Thomson, 1888),
Hymenoptera (Dalla ,' 1902),
.

32

(Wahl, Gauld, 1998).

Hymenoptera .
Diacritus Poemeniini Pimplinae
(Muesebeck et al., 1951) Poemenia Neoxorides,
(W alkley, 1967)
() Microleptinae.
Ortholaba, Diacritus
Diacritini Pimplinae (Townes, 1969),
Hymenoptera
(Carlson et al., 1979).
Oxytorus (= Callidiotes),
,

(Townes, 1971 ). Oxytorus luridator 1820 . ,

. armatus- 1883 .- . ,
Oxytorus Callidiotes, Mesoleptoidae
(Foerster, 1868). (Thomson, 1883, 1888)
(Schmiedeknecht, 1925, 1930)
Oxytorides (Oxytorina) ,
Mesoleptina (Mesoleptini),
.

2 ..

33

Callidiotes

Oxytorus

(, 1936).

Microleptinae,

1971

, , , ,

.
.

Cryptinae (Gupta, 1987), ,

Oxytorus

Atractodes),

,
, , -

Oxytorinae,

(Wahl, 1990; Gauld, 1991; Wahl, Gauld, 1998)

25

Hyperacmus Ophioninae (Schmiedeknecht, 1930) -

(Townes,

1958).

Microleptes

(Muesebeck et al.,'

1951; Walkley, 1967),

(Townes

et al., 1965; Townes, 1969).

(Dasch, 1992),

1991),
(Wahl, 1986, 1990; Wahl, Gauld, 1998).

n ,

(Woldstedt, 1881),
(1905), (Roman, 1921,
1936-); '-- (Ozols, 1941,

(Holmgren, 1881, 1883),

193 1), (1934,

1961); :~ - ~, , .
(1975), Jont1s (1983), , (2000), Jonais,
Rimsaite (2000). .
~ ,
,

(Townes,

Eusterinx (Aubert
Aperileptus Plectiscidea (Aubert, 1976), Proclitus
Pantisarthrus, Aniseres Helictes (Aubert, 1977)
1968),

,
,

Microleptes (Schwarz, 1991),

1998), Cushmania -
(Kasparyan, Humala, 1996),

(,

1994).

2*

34

(Hellen, 1915,

1937,

1937,

1953, 1961). 90-

(
, 1991, 1997, 2002; Kasparyan, Humala, 1995; Hurhala, 1997).

1939, 1940, 1942, 1946,

(Jussila,

Oxytorus.
Diacritus
D. aciculatus (Momoi, 1966).

(Uchida, 1928, 1930).

(Momoi, 1965),

( )

(Rossem, 1974-1991),

(Yu, Horstmann, 1997).

. ! ..

Microleptinae s. 1.
(Gauld,

(Townes, 1971).

1971)

(Keich,

1939), , ,

MICROLEPTINAE

Ichneumonidae

36

,
Pimpliform: Microleptinae (1 ), Cylloceriinae
(2 ), Orthocentrinae s. str.

(Gauld, 1991)

Helictinae,
Diacritinae (2 ) Oxytorinae (1 ) -
.

. , ,
,

Microleptinae

(Townes, 1971).
Oxytorus, Microleptes Cylloceria.

(Dasch, 1992),

Microleptinae
Hyperacmus Cushmania.
Cushmania 9

; , ,
(Kasparyan, Humala, 1996). Cylloceriinae Oxytorinae
( )

Orthocentrinae s. str.

Helictinae,

Microleptinae
1958) ,
(Plectiscus impurator Gravenhorst)
- Orthocentrinae.
Plectiscus

(Wahl, Gauld, 1998),

Pimpliformes, Cressonia,
Helictinae (Dasch, 1992), Dacritinae.

Orthocentrinae s. 1.,

, , -,

(ownes,

Plectiscus auct.-

Plectiscidea (Viereck, 1914).

,

(Fitton, Gauld, 1976, 1978),

Oxytorinae Thomson, 1883
Oxytorus Foerster.

(Wahl, 1986, 1990; Wahl, Gauld, 1998).

icroleptes

Microleptinae,
(Wahl, 1986).
Cylloceria Allomacrus
Cylloceriinae (Wahl, 1990),
, Oxytorus,

Oxytorinae

Orthocentrinae s. 1.
36

(Townes, 1971; Dasch,

1992),
Orthocentrinae.
Microleptinae, Cylloceriinae Helictinae sensu
Dasch (1992) ,
Microleptinae. Microleptini,
, . Cylloceria
Allomacrus,
(, 2002),
, Cylloceriini
Helictini
(== Plectiscini auct.). ,
Diacritini, Pimplinae
~ (Gauld, 1991; Yu, Horstmann, 1997;
ahl, Gauld, 1998).
(Wahl, Gauld, 1998),

Impliformes.
37

MICROLEPTINI Townes, 1958

:
, ,

. Microleptes ,
n , Eusterinx.
Cylloceria Eusterinx

, Micoleptes
;
.

; ;

Cylloceria

, .

; n
;

Microleptes

Eusterinx.
,

Microleptes,
Cylloceriini, Helictini Diacritini,

; , 1-

, ,

Cylloceria,

Hyperacmus);

Microleptes-

Hyperacmus,

Helictini),

(Dasch, 1992; Yu, Horstmann, 1997)

Microleptes, Hyperacmus Cushmania ,

Microleptes),

-
(Diptera, Stratyomyiidae ), ,
, Microleptes Hyperacmus (Wahl, 1986;
Dasch, 1992). Diptera Orthohapha,
, , Nematocera,
Microleptinae sensu lato, ,

, ,

,
.
Hyperacmus Cushmania (Wahl,
Gauld, 1998), .
Cushmania ,

Hyperacmus,

(Wahl, 1986)

,
. ,

, (Dasch, 1992): Microleptes,

Cushmania Hyperacmus,
Cylloceriini, Helictini Diacritini
Microleptinae.
Microleptini

,
.
, ,

(Cushmania),

. ,

, .

, , Helictini.
, ,
: , ,
,

Hyperacmus 4-5-
Cushmania, ,

1-

Cushmania

Hyperacmus

.
Cushmania.

Microleptes,

, , ,

Cylloceria

Eusterinx,

, -

38

r , ,
.

, ,
, , .
39

CYLLOCERIINI Wahl, 1990

'

Pimpliformes
Cylloceria Allomacrus

Cylloceriinae (Wahl, 1990).

Cylloceria

Allomacrus )
Microleptes, Helictes, Megastylus, Eusterinx,

Cylloceria, Allomacrus Rossemia -

Cylloceriinii Microleptinae s. l.

(, 2002).

.
,

Cylloceria,

DIACRITINI Townes, 1965

Microleptinae, Microleptini, Cylloceriini Helictini,

Diacritini Townes 1965,
Diacritus Ortholaba.

. ,

, ; ,

; ,

, 1-

, ; 1-

Cylloceria,

(Tipulidae),

. Cylloceria,
,

Microleptinae,

(Tipulidae).

(Townes, 1969; Wahl, 1993). Cressonia,

Helictinae (Dasch, 1992),

(Wahl, Gauld, 1998),

Pimpliformes.

Cyl1oceriinae Sweatel'ella (Wahl,

Gauld, 1998),
Rossemia (Humala, 1997).
Cylloceria, Allomacrus Rossemia

-, , ,

Helictini

Eusterinx

Symplecis.
, (
),

,
, ,

. ,

Pantisathrus .

(Thomson, 1888), -
Plectiscides (Schmiedeknecht, 1911)
(Perkins, 1940). Microleptinae.

(Wahl, 1986, 1990; Wahl, Gauld, 1998),

Microleptinae
s. 1., , : Apoclima, Entypoma, Kentrotryphon, Aniseres,
" "

( ),

Cylloceriinii
Helictini (Apoclima, Entypoma ,
Kentrotryphon),

40

Microleptinae s. 1.,

HELICTINI Gupta, 1987

.
m , ,
,
.

(Wahl, 1990),
41

1-

~
~

i:ij

<t:

~
,_:j

'"'
'"'

u
<t:

~
~
..... u::::

-<

<1.)

~
,_:j

~::

>..

Aperileptus

;::! <1.)
~

r::
!) ..:: -<
ut:"-'

.80
t: 11
0'-'

Kentrotryphon.

: (

;::!

), ,

~;::!

~::

i:ij

Pantisarthrus

, ,

Pantomima,

Cylloceriini
Aniseres,

, ,
.

Aperileptus,

, .

. ,

ORTHOCENRINAE

(lncludes remaining
former oxytorines)

(. .

(. .

8, 5),

7, 1)
Aperileptus

, -,
.

( ) Proclitus,
Fetialis, Dialipsis, Plectiscidea, Atabulus, Hemiphanes, Megastylus,
Helictes, Proeliator, Phosphoriana.
,

1- .

n
.

10.

Pimpliformes

(:

Wahl, 1990)

- ; - .

Aniseres, Aperileptus, Atabulus, Entypoma, Kentrotryphon

Pantisarthrus; Dialipsis, Helictes, Hemiphanes, Megastyl
Phosphoriana, Plectiscidea, Proclitus , Proeliator; Eusterinx
Catastenus, Eusteinx, Gnathochorisis, Sphingozona, Symplecis

(Eusterinx group ),
Eusterinx, ( ),
Catastenus, Symplecis, Gnathochorisis, Sphingozona,
,

1-

.
.

egastylus

1992), ,
Helictinae

(D

. ,

10),

(
) 1- , ,

42

, Eusterinx,
.

Proeliator.
Proeliator

, ,

( .

Eusterinx
- Keroplatidae.

Entypoma, Apoclima, Aniseres, Pantisarthrus

),
( Eusterinx

Proeliator) ()
(), J
-

43

 - ;
, ,

;

Orthopelma

, ;
, , .
( ) Eusterinx

n ,

, ,

Eusterinx s. str.,
,

Microleptinae (Aubert, 1980),

i .

Orthocentrinae (sensu Townes).

(Wahl, 1990; Wahl, Gauld, 1998).

(Sciaroidea). Orthocentrinae,
Gnathochorisis, Hemiphanes, Megastylus (
Helictini). ,
, ,

, ,

emiphanes :

(Holomeristus),

Proeliator

Eusteri

6-8-

Gnathochorisis.

Megastylus

, : ,

Holomeristus

(. .

2, 1),

Viereck,
.

, .

Dialipsis, Fetialis,

Proeliator.
lectiscidea

Atabulus, Pantisarthrus).

(,

Plectiscidea

Microleptini.

Pantomima (Rossem, 1990).

, ,

Acrodactyla Polysphinctini
(Pimplinae); .

al., 1994).
Orthopelma,
Pimplinae (. . 9). ,

Orthopelmatinae (Quicke et
Microleptinae s. 1.,

Helictini,

Orthopelmatinae.

, . ,

,
, ,
, -,

44

OXYTORINAE Thomson, 1883

, ,

Oxytorus
Mesoleptoidea (Foerster, 1868)
(==Ctenopelmatinae), Mesoleptina (Thomson, 1883, 1888)
Mesoleptini (Schmiedeknecht, 1925, 1930)
Oxytorides (Oxytorina).
Ctenopelmatinae
( Euryproctini), 1- :

45

Oxytorus ,
(Wahl,
1990;
Gauld, 1991; Wahl, Gauld, 1998),

Ichneumonidae (Yu, Horstmann, 1997),

(0. luridator), ,

Oxytorinae,
..........,,........ "_. .,.._
, , ,

. ,

Ctenopelmatinae Oxyt

Oxytorus)

I""'..n:r.-,n:r~.-

4-6- .

Ctenopelmatinae -
Oxytm-us ,
Symphyta,
Ctenopelmatinae.
Oxytorus
Cryptinae
- Atractodes. ,
. ,

( ,

1- ,

), q , LHJ.,.U.

Oxytorus - . Zuridator Grav. ''I"l7l...

. antennatus Cresson
( Mesoleptus) (Gravenhors
1829, .);
Mesatractodes Morley, 1907. Oxyto
Cryptinae (. . Gupta, 1987).
Oxytorus
( ,_.,,.,,,_,_

Cryptinae),
(W ahl, 1990)

Cryptinae) Oxytorus Cryptinae.

Mioreleptinae Oxytorus ,..,..,."''""- 1-. ( ,

j - ,
(",

0 ;' , .v
~ .::;

Microleptes-

- .

Oxytorus

, ,
.,............ 1_

Oxytorus

(. .

46

8).

, Microleptinae. ,
. -,
Cryptinae (
"lchneumoniformes" (Wahl, Gauld, 1998)) Ctenopelmatinae,
"Ophionoid subfamilies".
Epitropus,
(Rossem, 1990),
, Oxytorinae.

, Epitropus
Oxytorus, ,
, 7-8-
.

....

1,

, .
, ,
,

(.

1).

"

Diptera

) ... " ,

, .
,

, --

J!,~. . .

Hymenoptera, t

. ,

, , ,

(Shaw, Askew:

1979).
. -

r-j

(Wahl, 1986, 1990, 1993;

Hanson, Gauld, 1995; , 1996; Wahl, Gauld, 1998).
, -

L-

-
, (Roman, 1923). ClJc 1

: Morley, 1914; Edwards, 1924;
Mansbridge, 1933; Madwar, 1937; Seguy, 1940; Szelenyi, 1942;
F~rriere, 1955; Aubert, 1966, 1968; Pielou, Matthewman, 1966;
P1e1ou, Verma, 1968; Lastovka, 1971; , 1977; Trifourkis,
1. .977; Kolarov, Bechev, 1995; Jonaitis, Rimsaite, 2000; Rimsaite, 2000;
, , 2000.

Micro1eptinae Sciaroidea.

25

1977-2000 .

(~ .. )

(
),
- . ,

.

-
.

Microleptinae

,
(Diptera,
Sciaroidea).
Microleptini Cylloceria Cylloceriini.
- (. Stratyomyiidae). ,
Microleptes obenbergeri Chloromyia formosa Scopoli
(Schwarz, 1991), Microleptes sp.

(Wahl,
1986) Allognosta fuscitarsis Say
(Stratyomyiidae). Cushmania (Microleptini),
, Stratyomyiidae,
,

..

(1980).

11 - Microleptinae.
34 (26 )
33 .
,

48

(Rozkosny, 1982).
Cylloceria,

, - - .
Tipulidae.
. aino ,
Tipulidae; - . tipulivora,

, -

49

'1

.1

Microleptinae,
,

Diptera
1

( ;

.~

Microleptinae

* Entypoma robustum

- -

) -

* *

Anisaes pallipes,'
2JX.1978
. pallipes;
24.XII.1991
.

Dynatosoma fuscicome
D,fuscicome
18.YIII.l978
Mycetophila ichneumonea, Sciaridae;
24.YIII.l991

pallipes; 23.IX.1991

. fungoum

pallipes;

22.1.1991

A. pallipes; 16.IX.1991
.

pallipes;
20.VII.1982
. pallipes; 13.IX.l992
. pallipes; 1.IV.1975
*Pantisarthrus lubricus

7,XI ,1997

*Apeileptus

Gnathochmisis
flavipe s; 14.II, 1979
*Symplecis viuscula ; .1918
*Symplecis matilei

Roman, 1923

Apeileptus a!palpus ;
8 . .1991
.

vanus; 24.XII .1985

vanus; 27.XI .1987

vanus; 29.XII .1986

GB; VIII.l918
1830
Agrocybe praecox;
22.YI.1991

Ramaria flava;
21.IX.1985
R.flava; 1987
Leccinum vesipelle ;

*Eusterinx

+ .

ruficollis 23.IX.l991
Mycetophila fungoum

*Eusterinx

vanus; 15.XII.1981

vanus 30.IX.2000

vanus 30.IX.2000

vanus 5.11.2001

Lactaius

resimus;
13.YIII.1981
Phyllodon tomentosus;
Ks; 27.VIII.2000
Ph. tomentosus; Ks;
27.YIII.2000
Ph. tomentosus; Ks;

vanus 5.II.2001

* . a!palpus

Ph. tomentosus; Ks;

27.YIII.2000
GB

*. a!palpus

GB;

*.

Roman, 1923

Aubert, 1966

hetschkoi

hetschkoi

hetschkoi

hetschkoi

50

*.

caseyi
amoenus
*Megastylus sp.

Poc!itus adentis;

2. ! .1991

'
r

(-

**

M,fungorum
Sciophila rufa

Telephoa

Neoempheia stiata

terrestris;
14,VIII, 1978

. pa etor;

ll.IX.1990

Jonaitis, Rimsaite, 2000

Kolarov ,
Bechev, 1995

Diadocidia feruginosa

Roman, 1923

Neoemplia

Delobel, Matile,
1976
Rossem, 1987

GB
-

ombrop11ila

Mycetophi1idae,
Limoniidae, Muscidae

1983
NA

erotelion

Polypous

squamosus;
VII-IX.

johanseni
Orfelia fultoni
Orfelia sp.

GB

Orfelia sp.

GB

GB
NA
NA
GB, YI.1974
Lycopedon sp. ; ;
4.VIII.l991

stigma
Orfelia inops
. inops
r sp.
? eoplatus testaceus
Bmcl1ypeza signata;

Dasch, 1992
"

Mansbridge,
1933
Mansbridge,
1933
"

Dasch, 1992
"
"

Wahl , 1986

2.1 . 1991

Lycopedon if;

Allodia pseudodomesti-

; 2. . 1990

+ Bachypeza

sig-

nata
.

subsulcatus; 1.1990

.- ,

P.paganus;
26.IX.1991
Poclitus sp.;
24.VIII.1982
* . ardentis;

17. . 1997

*.

1,

'
;1
1

1
1

Phronia? forcipata +
Mycetophila alea?
Mycetophila pictula

Roman, 1923

Thompson,
1957; Dasch,
1992

Morley, 1914

Pleurotus pulmonaius;

Bachypeza t

; 22.1.1989

'

+ M.fungoum

pectoralis

*.

alea

Diptera

Amanita muscaria;
; 26JX, 1997
; 22JY ,83

* . o1tator
1

8. . 1908
* . alpalpus

) -

27.VI.2000
.

;
* . cuentator

1".

Epicypta atarima
Mycetophila fungoum +
Allodia ornaticollis;
8.YII.1991
. hetschkoi;
24.XII.1985
. hetschkoi; 27.XI.l987
. signatoides?

armata

*Megastylus impressor

M.fungorum + Allodia
omaticollis; 16.IX.1991
Allodia pyxidifomis +
Exechia seriata
Allodia lgens (?)

. /

* . ispinosa

.'

Phronia flavicollis

tenuicincta

1.

23.1.1986

*. lubricus

sp, ;

- -

7 1983

GB
Fomitopsis pinicola; ;
5.YII.1978
Entoloma clypeatum;
; 7.VIII.1991
saundersi?; ;
29.VIII.l991
Hebeloma crustulineforme; ; IX. l991
. mesophaeum; ;
2JX.1991
Kuehneromyces mutalis; ; 23. VI.1982
Russula sp.; ; 1992
Lactarius helvus; -

Micro1eptinae

- * , a !ipalpus

,1

Diptera (-

1 ()

attentus;
19.VII.1998
* . attentus
* . attentus
* . comes
28 .VIII.1997
* . comes 4.VIII.1997
*Proclitus edwadsi;
YIII .1918

* . edwadsi
* . edwardsi

Fomitopsis pinicola; ;
1991
Russula aeruginea; ;
1982
Polyporus ;
; 25.IX.1997

Mycetophila laeta?

? ;

llodia

Mycetophila

stoi

szernyi

Jonaitis, Rimsaite, 2000

"

sp.

La~tovka, 1971
Rossem , 1983
Jonaitis, Rimsaite, 2000
"

Mycetophila laeta

30. VII.1998

A1millaia

mellea

Lactarius pipeatus;
14. .1997
Russula sp.; ;
20.VII.1997
P/eurotus cornucopiae,
Pl. spp.; GB

Pleurotus cornucopiae, Pl. spp.; GB

Pleurotus osteatus

Codyla

"

Roman, 1923;
Edwards, 1924;
Madwa, 1937
Chandler, 1973

"

Trifourkis, 1977

Brachypeza radiata

51

Microleptinae

- -

) -

* *

A1millaria mellea
Fomitopsis pinicola;

23.1.1998

* . praetor;

? ;

29.1.1998

praetor; 4.XI.1998

Diptera (n-

* . fulvicornis
*. fulvicornis
17.XI.I998
* . praetor;

*.

1 ()
Microleptinae

Allodiopsis ustica

collaris;
11 .XII.1984
. collaris;
10.XII.1984
. eythropyga ;

arnania

JI.IX.1990
. spuria; 2 1.1.1992

tarnanii

. spuia; 16.IX.1991

28.1.1998
*.

?paganus; IX.1922

*P. paganus
Dialipsis exilis
26.XI.I982
*D. exilis 27.VII .I998
*D. exilis 19.VIII.I998
*D. exilis 25.VIII.1997

*D. exilis

26. VIII.1997

Clitocybe nebularis;
GB
Entoloma sp.; ;
2. VIII.1982
Russula sp.; ;
14.VII.I998
Russula sp.; ;
1. VIII.I998
Russula sp.; ;
13. VIII.1997
?

Bolitophila (Cliopisa)
glabrata
Mycetophilidae

Roman, 1923

Dasch, 1992

Jonaitis, Rimsaite, 2000

Cordyla sp.

Exechia

seiata

; ;

13.VIII.1997
Russula decolorans;
; 25.VI.1998
Russula sp.; ;
*D. exilis 6.VII.1997
23.VI.1997
*D . exilis 4.VIII.1997
Russula sp.; ;
20. VII.1997
*D. exilis 11.VIII. 1997 Russula sp.; ;
23.VI.1997
*D. exilis 28. VIII. 1997 Russula sp.; ;
l . VIII. 1997
Lactius pipeatus;
*Dialipsis exilis
28.VIII.1997
; 14.VIII.1997
Lyophyllum decastes;
Plectiscidea canaliculata; 6.VIII.1991
; 22.VI.1991

*D. exilis 13.VII. 1998

cinctula; 16.IX.1991

Melanoleuca brevipes;
28.VIII.1991
Inocybe geophylla; ;
29.VIII.1991
Bankera fuligineoalba ;
; 10.VIII.1985
B.fuligineoalba; ;

cinctula; 23.IX.1991

collaris; 13.1.1986

collaris; 11.1992

Cordyla sp.

Codyla

sp.

52

B.fuligineoalba;
6.IX.1990
B. fuligineoalba;
11.1X.l981

;
;

tereb1ator? ;
16.X.I991
Plectiscidea. sp.;
29.XII.I990
Plectiscidea sp.;
1O.XII.1990
Plectiscidea sp.;
16.IX.1991
Plectiscidea sp.;
22.VI.1991
Plectiscidea sp.;
1\.IX.1990
Plectiscidea sp.;
23.VIII.1983
Plectiscidea sp.;
16.VII.1991
Plectiscidea sp.

Plectiscidea sp.

sp.

Mycetophila fungoum +
Allodia sp. + Bolitophila
(Cliopisa?) hybrida
Allodia onaticollis +
Botanophila sylvatica;
. ornaticollis +
Mycetophila ruficollis
Mycetophila hetschkoi;
13.1.1986
. hetschkoi; 11.1992

26.1.1991
. collaris;
29.XII.\990
. collaris; 20 . . 1981

spuria

Plectiscidea sp.

Codyla

- -

26. VIII.1998

26.VIII .1998
Tricholoma flavovirens; ;

Dasch, 1992
Jonaitis, Rimsaite, 2000

; 25.1.1998
? ;

. hetschkoi;
29.XII.1990
. hetschkoi; 20..1981

Jonaitis, Rimsaite, 2000

1 ()

**

Sarcodon imbricatum;
; 22.VIII.1984
Cortinarius
(Telamonia) sp.; ;

Diptera (-

) -

hetschkoi;
11.XII.1984
. ruficollis stroi? ;
1O.XII.1984

20.1.1984

Clitocybe gibba; ;
2.VIII .1990
Hebeloma mesophaeum; ; 1992
Tricholoma inocyboides; ; 27.VIII.1991
. inocyboides; 199091?
Sarcodon imbricatum;
1991
S. imbricatum; ;

Allodia sp.+ Nlodiopsis

rustica
Exechia fusca + Allodia
lugens
Mycetophila fungoum +
Rymosia connexa
Exechia separata? +
Exechiopsis indecisa
Mycetophila hetschkoi
.

hetschkoi

hetschkoi

11.1.1990

Ramaria flava; ;
11.IX.1990
Russula sp.; ;
27.VIII.1991
g praecox; ;
1991
Laccaria icolour; ;
2.VIII.1990
Boletus pinicola; 1983

M.fungorum
M. fungorum +Allodia
ornaticollis
Rondaniella dimidiata?
Mycetophilafungorum +
Codyla breviconis

Mycena niveipes?; ;
20.VI.1991
Kuehneromyces mutabilis; V; 28. VI.1978
Inocybe lacera; V;
1978
Entoloma sp.; V;
28.VI.1978

Mycetophila dentata
Exechia seriata;
16.VII.1978
. lucidula + Exechia
nitidicollis
Suillia atricornis + Pego geniculata
22..1978

Plectiscidea sp.;
20. VII.1 982
Plectiscidea sp.;
12.IX.1991
Plectiscidea sp.;
16.IX.1991
Plectiscidea sp.;
l . IX.\ 991

Plectiscidea sp.;
9.VII.l991
Plectiscidea sp.
P!ectiscidea sp.;
1..1977

P!ectiscidea sp.;
19.X.l 992

Kuehneromyces mutailis; ; 22. VI.1982

Lactarius rufus;
22.VIII.1991
Omphalina pyxidata;
28.VIII.1991
Hebeloma mesophaeum?; ; 1991
Cortinarius sp.; ;
19.VI.1991
Agrocybe praecox
Suillus luteus ;
2.IX.1977
Phaeolepiota ;
22.IX.1992

Exechia seriata + Allodia

ornaticollis; 20.VII.1982
Mycetophila stroli
M.fungoum

M. fungorum + Allodia
ornaticollis
. uninotata
M. fungorum
Bolitophila ossica;
16. . 1977

Allodiopsis cristata +
Mycetophila fungorum;
19 . . 1992

53

Microleptinae

1 ()

- -

**

Boletus pinicola;
20.1X.I984

Plectiscidea sp.;
20.X.I981

Suillus variegatus?;

* .

Amanita muscaia;
; 17 .IX. 1964
PleuJotus ostreatus
Russula sp.; ;
7.VII.1998
Russula sp.; ;
14. VII.1998
Collyia sp.; ;

* .
* .

collaris
collaris;
23.YII.1998
*. collais;

30. YII.1998
*. collais; 1..1997

*.

collaris; 15. . 1998

* .

collaris;
17.YII.1998
*. colla1is;
19.VII.1998
* . monticola
17.VIII.1998
*. nava 25 . . 1998

) -

10..1981

Exechia sepaata +
Mycetophila fungoum;
II.I985
Mycetophila signatoides + Boletina sp. +
20..1981

11.1.1997
? ;
; 21.IX.1998
Lepista invesa;
; 25.VI.1998
Lepista sp.; ;

M.fungorum

Aubert, 1968

M.fungorum

Dasch, 1992
Jonaitis, Rimsaite, 2000

M.fungorum
Exechia

dosalis

tener

* .

melanocera

GB

* .

blstriata

Schizophora paradoxa;
GB
Lepista sp., Collyi sp., Cystoderma
amianthinum
Clitocybe sp., AmaniAllodia omaticollis
ta sp., Cortinaius sp.,
Russula spp.
Russula spp.,Boletus , Codyla fus ca
sp.
,
Tipulidae
17.VI.I979
; Tipulidae
; Guizhou; Y-VI.
1976
NA, :t~
Tipula (Tiplici:tipula) sp.
25.XI.1982
Chloromyia formosa

*Plectis:cidea sp.
CyllGceia aino
24..1979
*. apUvora

*Cyllocaia sp.;
25.1.1983
*Microleptes obenbergeri
*Micoleptes sp.;
25.XI.1982

NA, ;
26.III. 1981

Helictini

53 (Sciaroidea) 26
, : Mycetophilidae, Bolitophilidae
Keroplatidae, 15 28
.
Mycetophilini Exechiini
Mycetophilinae (Mycetophilidae)
Bolitophilidae,
. ,
,
- Sciophilinae, Leiinae,
Mycomyinae Gnoristoinae (Mycetophilidae).
Keroplatidae Keroplatinae
Macrocerinae egastylus

M.fungorum
Exechia fusca + tophila sp. (ruficollis
group)
Exechia fusca + tophila sp. (ruficollis
group)
Apolephthisa suinca + T1ichonta vitta
Allodiopsis domestica

llo gnos.ta fcitarsis

Roman,1923

..

,
(l,

1977

, ,
,

.
- ( )

. ,
lectiscidea,
Chao Hsiu-fu,
1994
Wahl, 1986
Schw.arz, 1991
Wahl, 1986

); - ; - ; N- ; Ks-

GB- ; NA- .

.
. , ,

: - ""; - (

54

n (

, , ,

Mycetophila fungoum

** - , .

Eusterinx .

Allodiopsis ustica

* .

*Plectiscidea sp.

(Chao, 1994);

Cylloceria sp. ()
Tipula (Triplicitipula) (Wahl, 1986).

Cylloceriini)

25.VI.1998
Rozites caperata;
; 1. YIII. 1998
Russula sp.; ;
12.VIII.1998
GB

*Plectiscidea sp.

Plectiscidea sp.;
Il.1985

agitator

Diptera (-

Mycetophilidae,
-

Mycetophilinae

Mycetophila. Plectiscidea
Allodia, Exechia,

Allodiopsis, Cordyla, Exechiopsis (Mycetophilinae, Exechiini),

Rondaniella (Leiinae), Apolephthisa (Gnoristinae) Bolitophila

(Bolitophilidae) (. 2).

Aperileptus, ,
I Mycetophila (. alea,
. fun gorum, . hetschkoi, . signatoides, . finlandica),
(Morley,
55

19

(Diptera)

--------------=-----.------___j

Macrocerinae
Keroplatinae

Keroplatidae

Keroplatinae
Diadocidiidae

Macrocera
Oifelia
Keroplatus
Cerotelion
Neoditomyia

Megastylus

Orfelia
Cerotelion

Eusteinx

Diadocidia

Symplecis

f-------------< Gnathochoisis

Mycomya

Mycetophila

Mycetophilini
Mycetophilidae

antisarthus

Mycetophilini

,
.

caudatus

Aniseres pallipes

F omitopsis pinicola (Humala, 1997),

Mycetophila Dynatosoma.

Proclitus

Mycetophilinae.

Pleuotus

pulmonaius

(Polyporales) ( Brachypeza),
(Trifourkis, 1977).
, ,

Russula aeuginea (Russulaceae) Lycoperdon pyriforme

(Gasteromycetes: Lycoperdales). Proclitus

Dynatosoma
Mycetophila
Allodia

Clavariadelphus, Ramaria, Bankera Sarcodon (, 1994).

Aniseres pallipes
, Entoloma clypeatum, . saundersi, Kuehneromyces mutailis
(Strophariaceae), Hebeloma crustulineforme, . mesophaeum
(Cortinariaceae) Russula spp. (Russulaceae).

Sciophilinae
Sciophila
1 - - - - - - - - , - - - - - - - - - + - - - - - - - - - J Aperileptus

Phronia
Epicypta

vanus,
hetschkoi,

Mycetophila Allodia.

Neoempheia

Mycomyinae

14; Roman, 1923; Dasch, 1992; Jonaitis, Rimsaite, 2000).

, Aperileptus
.

,
Anisees

,
.

Brachypeza
Exechiini

Mycetophilini
Bolitophilidae
Tipulidae

Allodia
Exechia
Allodiopsis
Cordyla
Exechiopsis

- Proclitus

Plectiscidea

testaceus (Wahl, 1986).

Keroplatidae,

Megastylus

Macrocera

(Macrocerinae), Ofelia, Ceotelion, Neoditomyia

(Keroplatinae).
, , , -

Bolitophila
(Cliopisa)

Poclitus

ipula

Cylloceria

Allognosta

Mycetophila

Chlomyia

sp.

Hyperacmus
Stratyomyiidae

egastylus

Keoplatus

Micoleptes

, - - .. ,
(Keroplatus, Cerotelion),

(r, Orfelia),
CMansbridge, 1933; Matile, 1990) (. . 2).

. , egastylus

,
.

56

57

Gnathochorisis flavipes
Neoempheria striata (Mycetophilidae, Mycomyinae),
Thelephora terrestris (Thelephoraceae ).

-.

Symplecis breviuscula (Roman, 1923)

Diadocidia ferruginosa (Diadocidiidae),

; ,

, -

Mycomyinae - Neoempheria ombrophila (Delobel, Matile

197).
.
,

Helictini

~v .., .., . V ! V - .

Sciaroideae; Microleptini
Cylloceria - Tipulidae.

(.

Stratyomyiidae,

3), (Townes et al., 191,

195; Townes ., Townes ., 19, 1973; Gauld, 1984, 1991; Gupta,
1987; Dasch, 1992; Yu, Horstmann, 1997)

Oxytorinae

,
,

Oxytorus 1

Microleptinae s. l. ,
328 33 .
: Microleptini
3 11 , Cylloceriini - 45 ,
Diacritini - 3 5 , Helictini - 21 27
(Yu, Horstmann, 1997; , 2002). .

.
. ,

50

Microleptini

Helictini), , .
Oxytorinae (
, . ); Microleptini
; Diacritini - ,
lloceriini - 14 , Helictini - 14
101 , ..
36% . : Cressonia
(Diacritini) Neoproclitus (Helictini) ,
(Dasch, 1992).
, ,

Microleptinae ; -

59

7'

.,

1 1

' 1

11
1
1

Microleptes
Cushmania
Hyperacmus
Cylloceria
Rossemia

13
1
4
1
3

Allomacus
entotyphon

Apoclima
Entypoma

Aniseres
antisarthrus
Proclitus
Pantomima
Helictes
Megastylus
Hemiphanes
Neop1oclitus
Sphingozona
Fetialis
Plectiscidea
Dialipsis
Atabulus
Epitropus
Aperileptus
Proeliator
Eusterinx

10
1
4

2
1
1

3
4

4
1
4
2
10

'

1
1

30
1
1

20
1

4
14

5
1
24
8

2
1

2
1

Oxytous

10

255

15

12

+
.+

5
1

+
3

2
+

2
1

+
2
+

,!

1
1
1
1

+
+

\'

.::
11

Ortholaba
Cressonia

5
4
11
1
5
10
4

11
2
27
7
4
1

Diacitus

5
5
15
1
7
15
7

Phosphoiana

Gnathochoisis

.,

14
2
31
9
7
1
1

Symplecis
Catastenus

10
1
1

1.

13
2
5
25
1
4
1
7

..

8
11
24
1
10
34
7
1
2
1
95
2
1
1
18
3
48
15
14
2
1

3
1
1

21

13

12

>4

>14

380

'

-;

'
.~

1,

60

61

 70 .
,
40 ;
Cylloceria (6 ), Pantisarthrus (5 ),

Plectiscidea, Helictes, Megastylus, Proclitus, Sphingozona, Symplec

Catastenus, Gnathochorisis Eusterinx (Townes ., Townes ., 196
Gauld, 1991).
(Sphingozona)
(Townes, 1971), .,..,.,..,.,,..-.1-

,
,
. (Owen D., Owen J.,
1974; Gauld, 1987, 1991),
,

,
.

, , ,

, .

13
(Townes ., Townes ., 1973; Matile, Delobel, 1976).
egastylus - 5 ;
Proclitus - 3 , Symplecis Plectiscidea 2 , Aperileptus 1 (. 3).

Microleptinae

,
, .

; Microleptes, Hyperacmu
Cylloceria, Apoclima, Entypoma, Pantisathus, Hemiphanes
Megastylus, Helictes, Proclitus, Apeileptus, Plectiscidea, Symplecis

Gnathochorisis, Eusterinx (Townes et al., 1961; Gau1d, 1984;

1987; , 1998).
(Gauld, 1984).

, .

, ,

(91%)
.

_.

65% ;
: Microleptini (7 )- 64%, Cylloceri
(27 )- 60%, Diacritini (3 )- 60%, Helictini (176 )
66% . ,
, Microleptinae

( , ,

, ,

.

(213

)-

, , ,..,....,,, .... ,.., __

,
..

(1984).

, , ),

.


.

1. (5 ).
1.1.
: egastylus cruentator, Hyperacmus
crassicornis, Cylloceria melancholica.
1.2. - Microleptes
rectangulus emiphanes townesi.

Mycetophilidae-

. 1'"'\,-,.
,

.
-

62

63

2.

(32 ).

, -, ( ).
.

nava,

Aperileptu~ flavus,

nemorensis, . subangulata, . substantiva,

impurus, _. inf~s~atus, . . melanopsis,
. micospzlus, . plagzatus, . rossemz, . trzcznctus, . vzduatus.
3.6.2. (13 ) Entypoma
robustator, . prominens, Symplecis paradoxa, Phosphoriana rugosissima, Eusterinx tartarea, Proclitus albldipes, . fulvipectus,
. edwardsi, . rudis, Plectiscidea prognathor, . humeralis,
. moerens, . tenuicornis.
3.6.3. (6 ) Kentrotryphon longecauadatus,
Pantisarthrus rudepunctatus, Hemiphanes montanum, Fetialis alacris,
Plectiscidea indomita, Epitropus insolitus.
3.6.4. (
- 76 ).
3.7. (1 : Cushmania psarevi).
3.8. (7 ):
Cylloceria brachycera, Allomacrus jakuticus, Entypoma frontosum,
Aniseres paradoxus, Plectiscidea obscura, Eusterinx truculenta,
. similis.
3.9. - ( 21
). 17
neni sp.n.,

2.1. - 19
Cylloceria borealis, Allomacrus arcticus, Pantisarthrus lubr
Megastylus pectoralis, . impressor, . orbltator, Helictes borealz
. erythrostoma, Eusterinx oligomera, . blspinosa, . trifasc
Gnathochorisis crassula, G. dentifer, Symplecis blcingulata, Cataste
femoralis, Plectiscidea collaris, Proclitus fulvicornis, . paga
Aperileptus alblpalpus.
2.2. 13
: Microleptes splendidulus, Entypoma robustum, Aniseres pallipes
Aperileptus vanus, Eusterinx subdola, . inaequalis, . refractar
. tenuicincta, Symplecis breviuscula, S. invisitata, Megastylusflavopz
tus, Proclitus praetor, Plectiscidea communis.
3. (204 ).
3.1. (23 : Microleptes aqui
granensis, . salisburgensis, Cylloceria fusciventris, . invict
Rossemia longithorax, Apoclima signaticorne, Entypoma suspicios
Pantisarthrus gracilis, . luridus, Helictes incongruens, Eusterz
,
obscurella, . argutula, . trichops , Proeliator proprius, Procl"
ardentis, . comes, Hemiphanes flavipes, . gravator, Dialipsis exilis .
Gnathochorisis flavipes, G. xanthocephala, Plectiscidea agitator .

Diacritus aciculatus).

3.2.

. lchneumonidae
(Apoclima haeselbarthi emiphanes

, ,
jo rmidatum).

(91% )
3.3. - (14 ). Cylloceri

.
Mlcroleptinae,
sylvestris, . tenuicornis, Allomacrus subtilis, Proclitus attentus

(
Plectiscidea spuria, . subteres, . melanocera, . monticola, . parvu

),
la, . terebrator, Eusterinx pseudoligomera, Eusterinx aquilonigena

15%,

. minima, Helictes carinata sp.n.

Tryphonini,
3.4. --

7%.
, : Megasty

excubltor, Eusterinx circaea, Plectiscidea vagator, Helictes fabularis.
,
(87 , 35%
3.5. - (4 : Plectiscide
n
)
canaliculata, . helvola, . posticata, Oxytorus luridator).
,

3.6. (128 ).
~,

3.6.1.
D1plazontinae38%
(Manukyan,
1995).
(33 ). Oxytorus armatus, Aniseres caudatus, Pantisarthrus dispar,

(23 ),
Proclitus subsulcatus, . zonatus, Helictes conspicua, . kar-elica sp.n.
n

(2
)

Eusterinx jugorum, Atabulus faustus, lectiscidea amicalis, . aquilo

(14
)
.

sp.n ., . blstriata, . cinctula, . conjuncta, . crassicornis, . ery , thropyga, . fuscifemu sp.n., . helleni sp.n., . hyperborea, . ko
:

64

3 ..

65

', ,
( ).

, .

, Cylloceria invicta, . fuscive

. tenuicornis, Aniseres paradoxus, Proclitus paganus, Megastylus
toralis, . impressor, Eusterinx oligomera, . trifasciata, . tric
. aquilonigena, Gnathochorisis dentifer, Catastenus femora
lectiscidea

Plectiscidea

collaris, Aperileptus vanus,

n Helictes , Pantisarthrus, Proclitus, Eusterinx,

Apel'ileptus; :
Megasty lus, Cylloceria, Pantisarthrus, Aperileptus, Proclitus ,
Eusterinx, Symplecis; - Entypoma ,
Eusteinx, Microleptes.

Entypoma robustator, Proclitus

. tenuicornis ,
u.,. . ...._

Microleptinae

( )

Microleptinae

a-

Plectiscidea grossepunctata,

. , -

, , -

)

,
, '
. ,
(Scaramozzino, 199
( )
,

(Perkins, 1939).
30%
, (Gauld,
1991). (, 1991, 1997) (Humala,
1994, 2001).
, . -

( )

1989-1991 . "". -

,
.
~ 30
250 , - 1 10 .
21 10 ,
13 , , ,

Ichneumonidae,
,
9% ( ) 22% ( ) 26% ( ) (. 11).
" " , ,

: Cylloceria aino .._.,u. u..JJ.-".Ij

(Uchida; 1928), Diacritus incompletus 7 Oxytorus
(Momoi, 1965, 1966), Ortholaba tenuis (
1969), Eusterinx (Divinatrix) inaspicua , Entyp
ferale Eusterinx (lschyracis) pel'miranda

30% . -
50-65 %, Mycetophilidae
(Russell-Smith, 1979; , 2~ .).

(Rossem, 1988). , 6
1

, ,

66

I .

''" (1997); -

(2000).

67

 , ,

1\1 Pimplinae

n .

Cryptinae

111 Campopleginae
8%

9%

18%

4%

Orthocentrinae

18%

(
) ,
.,

111 Cryptinae

6%

111 Microleptinae

111 Tryphoninae

10%

" " (

6%

nn, . . .

1!1 Ichneumoninae

4%

111 Tryphoninae
111 Ctenopelmatinae (8%)

20%

. .

Ctenopelmatinae (1

1994) : egastylus
cruentator, . orbitator, Symplecis icingulata, Gnathochorisis
fiavipes, Aperileptus vanus, . infuscatus, Pantisarthrus lubricus,
. gracilis, Proclitus praetor, . paganus, . subsulcatus.
Megastylus oritator Symplecis
icingulata.
,

.
,

111 Campopleginae

(2000),

11 Ophioninae

, ,

lchneumoninae

,
.

1!1 Microleptinae

111

, , ,

gulata,

S.

ic in

, ,

, ,

6%

4%

6%

20%

12%

111 Pimplinae

. . oritator,

111 Tryphoninae

- ,

, .

Cryptinae

111 Banchinae

. ,

1!1 Campopleginae
111 Microleptinae

, .

35%
.

11.

20

75

Microleptinae
:

Ichneumonidae

( , "",

1989

.)

Plectiscidea.
, ,
,
(, 1991, 1997),

r - .

68

69

;
. ,
,

, Proclitus, Pantisarthrus
Aperileptus.

Cylloceria, - (Tipulidae).

, 0

- .

Aperileptus lectiscidea :
- - .

OXYTORINAE

1(2).

.
, ,

MICROLEP/NAE

,
; ,
Oxytorus luridator;
; ,

. ,
. 1-
, 0,6-0,7 , ........... ..

2,4-3,6

.. ..... .. .. ... ... ......... ...... ............. ................ ........... .. .......... .. .... ... ......... ...Oxytorinae

2(1).

, ; ,

Microleptes,
Hemiphanes

, ,
,

Megastylus,

(1,1-2,1

( ) ..... Microleptinae

MICROLEPTINAE Townes, 1958

1(2).

;
; ;
;

; ................................................................................ 1. Microleptini
2(1). ;

;
; ;

;
.

3(4). ,
, .

.
; , , ,
.
; -

71

.....
.............................................................................. 2. Cylloceriini

4(3).

3(4). : 1-2(3)(4)- 5-8- ,

, , ,
,

, ,

; 14-18
, ,

, . (

Aniseres, Pantisarthrus, Aperileptus Pantomima).

, 1-

, , , .

.
, .......................................... .

5(6). . .
. .
.
. 1- . .

, ......................... .4. Diacritini

6(5). . (
,
. .)

1.

... 3. Helictini

.................................. ...................... 1. Microleptes Gravenhorst

4-5- ,

4(3).

, ,

, ;
;

MICROLEPTINI Townes, 1958

1.

, .
,
; ,

14-32 ;

1-3(4), 4-5

, 1-

4,1-8,1 ; 1,3-7 ;
, ,

26-32

5-8;

; ;
;

.................................... 3. Hyperacmus Holmgren

MICROLEPTES Gravenhorst, 1829

Foerster, 1868; Schulz, 1906; Schmiedeknecht, 1911, 1924; Walkley, 1967;

Rossem, 1980; Schwarz, 1991.
= Miomeris Foerster, 1868
= Minomeris Schulz, 1906
= Gnathoniella Schmiedeknecht, 1911
= Microcleptes Walkley, 1967

(.

Microleptes rectangulus (homson, 1888)

12).

; ,
; ,

; ;
;

; 1-
, ,

Hyperacmus );

; .
:

Microleptes, Hyperacmus

Cushmania.
MICROLEP/N!

1(2).

16-17

; ,

2,45-2,85 ,
.................................... .
..... ... ..... ........... ............................... .. .-................................. 2. Cushmania Dasch

1-

2(1 ). ;
, , , ,
,

72

1,65-1,95

, .

12. Microleptes rectangulus

(: Townes,

1971)

73

 .
, .
1-3(4)- 5-8- ,
;

14-18 .
. ;
, ;

;

, ;

; 1-
.

I_ MICROLEPTES

1(2).

(. 13, 5); ,
; ; 1-
, , 2- ,
. 5-8- .

........... .
........ ......... ....................................................... ...... rectangulus Thomson

2(1).

,
3 (. 13, 1-4);
,
. 1-2(3)(4)- .

3(6).

(.

13, 6);

, .

4(5).

,
;

(.

13, 2);

.....................

splendidulus Gravenhorst

;
(.

13, 1).

6(3).

.......................................... . obenbergeri Gregor

. . (. 13, 7); ,
, ,

7(8).

,
, ;
,

, (.

13, 9);

, 2-, ;

,
(.

74

13. Microleptes

1-5- : 1- . obenbergeri, 2- . splendidulus, 3 -. aquisgranensis, 4- . salisburgensis, 5- . ectangulus; 6-7-

: 6- . splendidulus, 7- . salisburgensis,
: 8 - . salisburgensis; , , 9 - . salisburgensis (:
Schwarz, 1991)

1-2(3)-

5(4).

13, 8). 1-3(4)- .......................................................................... . salisburgensis Schwarz

8(7). , ,
; V -
; 2-;

1-3 ........................... . aquisgranensis Foerster

1. Microleptes aquisgranensis (Foerster, 1871)

= Miomeis aquisganensis Foerster, 1871

. .

. 1 . . 22.VII.l975 ().
(Ozols, 1961).
.
75

2. Microleptes belokobylskii Humala, sp. n.

. .
(): 4.3 ;
3.6 ;

28.VII.1972

0.5

20

: 2-5.VIII.1993 (); .
().

. ,
, ,
; ,

(); ,

24.VI.1981 (); 30
1. VIII.1993 - (), 20

18.VII.1995

3. Microleptes grandis Humala, sp. n.

. .

. ,
. 17 ( 16) ;

2.0

. , ,

1-

, ;

1.15

(): 7.5 ;
;

5.3

. 1-3- .

. ,
.
- ,

. , .
18 ; ,

. , .

. ,

1.54

1.6

1- . 1-2-

OOL

. .

. ,

; ,

. .
, , ,

V- ,

, . .

. .

, .

4.0

( ,

); ,

; , .

. .

, ,

, .

.
, .

, , .

. ,

; , .

4.7

.
.

. .

, . -

. ,

0.6

0.5,

. .

. , , ,

, ;

. ,

, .
.

, ;

0.4

0.45,

. ,

10.IX.1970 (); : -

,. -

76

6.

77

.
. .
. , , ,
. , ,
.

0.58

. .

.
.
.

. : . : .

13.VII.1993 ().

. , , , , ,
, .
.

. : ~
().

4. Microleptes minor Humala, sp. n.

. .
(): 3.3 ;
2.9 . ,

1O.IX.1982

5. Microleptes obenbergeri Gregor, 1938

. ,

. .

, ;

6. Microleptes orientalis Humala, sp. n.

; ~.

. ,

. 14 , ;
, 2.1
,

1.26

1- .

. ,
.

, V- ,
. ,

. , .

. .
; ,

. .
, ,
..

5.2 ,
,

, ; , .

.
.

.
, .
,

.
, .

,
;

78

0.46,

. .

(): 4.5 ;
;
. , ,

3.8

; ;
,

. , , ,
.

14

; ,

2.2

1.45

1-

. . ,
.
,

V -

, . ,
.

. .
; ,

. .
, ;

.
.

4.7

; , .

. -

79

(), , ,

, .
, , ,

; .
.

, .

, .

8. Microleptes salisburgensis Schwarz, 1991

. .

, ;

0.46, 0.51 .
. ,

6-7.VII.1975

()

1. VII.1971 ();
10

9. Microleptes spasskii Humala, sp. n.

.
.

, .
. ,

3 .

. .

():

3.3

3.9

. ,

, , , , ,

. ,

, .

, ;

- .

. 4.8 ; 4.0 ;

16

1-2- (. 3, 9). ,
.

. 4 S?, 25 . : ,
, 10.IX.1982- (
), -, 16.VIII.1972, ,
21.VIII.1972 (),

19.VIII.1978, - , 40
25.VIII.1978, - ()
11-13.IX.1978, -, 14.IX.1978 (
); , , 16.IX.1978,
29.VIII.1982 (),
3.IX.1983 (), 20 31.VII.1991,
13.VII.1993 ().

, ,

-
.

, .
;

. , ,

. ,
.

1.9

0.8

1- . .
, .
,

V -

, .

, .

. .
; ,

. .
, ,
, .

3.6

; , .

7. Microleptes rectangulus (Thomson, 1888) (. . 12)

= Miomeris rectangulus Thomson, 1888

= Gnathoniella egregia Schmiedeknecht, 1924

. .
. 14 S?, 8. , ., -

80

.
, .

, , .
, . -

81

 ,
;

0.47,

0.52 . .

.
n, ,

3.5

. , .

; , .

, .

1. 1

. ,

, , , , ,

.
. , .
0.3, ,

, . -.
.

. ,

. 2 ~ . :

11.VII.l991

().

, , ;

0.5,

0.6

. .

10. Microleptes splendidulus Gravenhorst, 1829

= Miomeris glabriventris Thomson, 1888

. .

. 13 ~, 11 . .; , .
.

11. Microleptes tibialis Humala, sp. n.

. .
(): 5.0 ;
3.9 ;
. ,

, ; ;
,

. .

.
.

. , ,
; , ,

.
.

. : ~.

20

31.VII.1991

().

2.

CUSHMANIA Dasch, 1992

Dasch, 1992; Kasparyan et Humala, 1996.

Cushmania ignota Dasch, 1992,

,
. , , ,

~ ~. ~

14 ;

4.1-4.8

1.3-1.4

, 1.9 ,
, 1- . .

; , ,

, , .

, ;

16-17

2.9

V- ,
. , .

; ,

. ; ,

. .

; , ;

2.45-2.85

; ,

. . ,

; 1- ,

, , .

, .

82

2.6

83 .

4-5

,
, , ;

26-32

-;
, ;
,

1-

. .

; .
, -
. ,
.

1. Hyperacmus crassicornis (Gravenhorst, 1829)

(.

15)

Exochus crassicornis Gravenhorst, 1829

= Lamponota suerinensis (Brauns, 1905)
. - .

46 S?,

. .,

7
.

14. Cushmania psarevi

1 - , 2- , 3- , 4-
, , 5- , , 6- , , 7-
, (: Kasparyan, Humala, 1996)

1. Cushmania psarevi Kasparyan et Humala, 1996

(.

14)

. .
. 1 (). ,
; 15-19.VII.1989 ()
. .

3.

HYPERACMUS Holmgren, 1856

Holmgren, 1856; Cameron, 1902.

84

Hyperacmus crassicornis (Gravenhorst, 1829).

15. Hyperacmus crassicornis

(:

Townes, 1971)
85

., ., , ., ,

1.

., , ,

; ( ., ), ,

(), , , , ,
.

2. CYLLOCERIINI Wahl, 1990

CYLLOCERIINI

1(4). .
, , .
.

2(3). ,
. . 3.0-4.7
. 3-4
................................................ 6. Apoclima Foerster
3(2) .. . .
5-9 ............................................. 3. Rossemia Humala
4(1). .
5(10). .
6(7). . ,
, 1.8-2.5
. .

CYLLOCERIA Schi0dte, 1838

Schidte, 1838: 140; Foerster, 1868: 166; Woldstedt, 1881: 61; Hellen, 1915:
47-48; , 1934: 195-198; Townes, 1971: 192; Rossem, 1980: 98-107, 1987:
54-58; 1988: 106;Dasch, 1992:267-286
= Lampronota Thomson, 1877
= Chalinoceras Ratzeburg, 1852
=Asphragis Foerster, 1868

Cylloceria caligata (Gravenhorst).

15

(Rossem, 1980, 1987).

(2002).

Cylloceria -

14-15

6.5-10.5

. ,

Cylloceria

,
(. .

3, 3).

. 3-6- -

............................................................. 4. Entypoma Foerster ()

7(6). . (

. .

Kentotryphon ).

8(9). 2.5 .
...... 2. Allomacrus Foerster ()
9(8). 2.3 . .
, (
, 2000 ) ... 5. Kentrotryphon StroI
.

10(5).
11(12). , (
6.5-10.5 )

............................................................................. 1. Cylloceria Schi!Z)dte

12(11). , 3-5(6)
13(14). , , 2 , .
. .....
..................................................................... 2. Allomacrus Foerster ()
14(13). 1.3-1.7 .
. ,

......... 4. Entypoma Foerster

.
, .
. ~
, . .

, , .
(
) -- (Diptera, Tipulidae).

(.

Tortricidae) ~ ~~

Cylloceria

Lissonoti

(Banchinae).

86

()

CYLLOCERIA

1(2).

. ,

, .
1- . 2-4-
-, . ~-
. -

87

 . - . .................... .

................................................................................ ..... 9. . occupator Grav.

2(1). ,
.

3(4). ;
17-19 . ,

. ,

. .

................ ........ 3. . borealis Roman

4(3). .
. (

brachycera).

5(6). . ,

. .

-.
. 0.46

)(10

. - ......................................................2. . alpigena Strol

6(5). . .
7(10). .
8(9). ,
(. 16, 6). .

.
. 23-24 .

0.85 .................................. 7. . invicta Rossem

9(8). . ,
. , ,
( )
.

. 25-28 . 0.50-0.67

................................... 8.

9
4

melancholica Grav.

10(7). .
11(16). .
12(13). (. 16, 9).
19-21 ; .
. ,

. .
, ................................................. 14. . ussuriensis Humala
15(14). , 5.0-9.5 . , 0.62-0.81

J
{?7

3.5 (. 16, 1) .............. 10. . orientalis Humala

13(12). . 26
. , 4.2-4.5
(. 17, 2-3).
14(15). , 9.5-10.5 . , 0.41-0.52

11

14

16. Cylloceria
1-3- : 1 - . orientalis, 2- . ussuiensis, 3- . aino; 4-6-
: 4- . melancholica, 5- . tenuicornis, 6- . invicta; 7-9-
: 7- . brachycea, 8 - . ussuiensis, 9 - . oientalis; 10-11 Allomacrus: . jakuticus: 10- , 11-
, 12 - . volcanus: ; 13-14-
: 13- . acticus, 14- . subtilis (: , 2002)

. ,

............................................................................. 1. . aino Uchida

16(11). .

88
89

17(18). , 0.8
(0.35 0.28 ). 1-
7 . 20 ................................................... 4. . brachycera Humala
18(17). , , .
19(20). 2- , , . ............................. .
.......................................... ...........
5 fusciventris Hellen
...... ......................
20(19). .
21(22). 0.5-0.6 .
7-9.5 .
. .................................................................. 8. . melancholica Grav.

22(21). 0.9-1.1 .
, 9-11 .
23(24). 21 , 11
........................................................... 13. . tenuicornis Humala
24(23). ,
9.6 .
25(26). ;

10(7).
11(12).

3- ,
;

4- (. .

12(11).

. .

........................ 12.

sylvestris Grav.

1(2). . 2-4- -,
......................................................... 9. . occupator Grav.
2(1). .
3(4). ,
.............................................................................. 8. . melancholica Grav.
4(3). .
5(6). , 2- , -
. , ,

22-24

..................................................................................... 3. . borealis Roman

3- ,
; 4- .

13(14).

. , -

14(13).

............................................... 12.

sylvestris Grav.

. ,
.
.

15(16).

16(15).

25-28

7-8

.............................................................................. 8. . melancholica Grav.

23-24 , 8-9

, 3-4-

............................................................................ 13.

. .
. - ........................................... . . imperspicua Rossem

26(25). ;
,

3, 3).

. . ,

tenuicornis Humala

1. Cylloceria aino Uchida, 1928

. .

35 ~,

20. ,

, -, .
.

17.VI-24.VIII.1979,

Tipulidae

(.N'2 47,

, ). .

2. Cylloceria alpigena

(Strol,

1901)

. .
.

(Aubert,

1968) ,

Allomacrus arcticus

(Holmgren, 1881).

. ,

. 22-23 ........................... 5. . fusciventris Hellen

6(5). ,
( . ussuriensis,
).

7(10). .
8(9). . .......................................... 14. . ussuriensis Humala
9(8). ,
. , .................................................................................. 1. . aino Uchida

* . oientalis . b1achycaa .
90

3. Cylloceria borealis (Roman, 1924)

. .

120 ~,

152. : .,

, ., , . , , ,
, ; .

4. Cylloceria brachycera Humala, 2002

. .

1~

(). ., ,

. ~

4.VIII.1975

().

91

5. Cylloceria fusciventris (Hellen, 1940)

349 2, 883 .

., ,

. .

., ., .,

. 3 2, 12. : ., - ,

, ., , .,

; , .

, , . , , ., ,
., , , ,

Cylloceria imperspicua Rossem, 1987

., , , ; ,
, , , , , ,

. .

, , , , .

7. Cylloceria invicta Rossem, 1987

. .
. 2. - . . ,
19, 24. VII.1982 (); , ". .
. ." 29.VII.1897 (); - . "7-89",
VIII.1874 () (); [- ]
. ,

VIII

().

9. Cylloceria occupator (Gravenhorst, 1829)

. .
,

civentris Hellen,

. .

fus-

, ,
, .

1. Cylloceria orientalis Humala, 2002

8. Cylloceria melancholica (Gravenhorst, 1820) (. 17)

= Cyllocaia caligata (Gravenhorst, 1829)

(Rossem, 1987),

. -
.

. 7 2. , .

11. [Cylloceria suerinensis (Brauns, 1905)]

= Hyperacmus cassicornis Gravenhorst,

1929

12. Cylloceria sylvestris (Gravenhorst, 1829)

. .

. 4

2, 99 .

.,

., , , , , ,
., ., ,

, , ., ; ,
.

13. Cylloceria tenuicornis Humala, 2002

. .

9 2,

5. .,

(R. Frey);
29.VII.1908 ();
; , Al: Karislojo (W. Hellen); Ivalo, Tavastehus
(=Hameenlinna) (J. Sahlberg).
. -

14. Cylloceria ussuriensis Humala, 2002

.

. 17. Cylloceria melancholica (: Townes, 1971)

92

93

12?, 4. : -,
3, 6, 7.VII.1981 ();
13 15.VI.1993 (); 15
16. VI.1993 (); 26-27. VI.1996 (

, ,

).

2. ALLOMACRUS Foerster, 1868

Foerster, 1868: 177; Holmgren, 1881: 13; Thomson, 1888: 1281; Roman,
1910;Townes, 1971: 190; Rossem, 1980: 95-96, 1988: '106; Dasch, 1992: 286-290.
= Sibliakoffia Holmgren, 1881
= Ciptopimpla Roman, 1910

Allomacrus arcticus (Holmgren, 1881).

.
(Rossem, 1980, 1988).

(2002);

(Townes, 1971).
ALLOMACRUS

18. Allomacrus arcticus

(:

Townes, 1971)

1(2). , - ,
0.14 .
, ,

(. .

16, 11).

, -

(. .

16, 10) ................................. 2. A.jakuticus Humala

2(1). , ,
. , (

. subtilis ),
. ,

0.29 ,

. .

1. Allomacrus arcticus (Holmgren, 1881) (. 18)

SiiYiakoffia actica Holmgren, 1881

= Allomacrus pimplarius Thomson,

94

1888

32. ., ,

, , ., . , .,
, ., , , ; .
, (. ), ( ),
, .

. .

3(6). . ,
0.24 . .
4(5). . ,
(. . 16, 13). 1.7-2.0
. 4.5-7 .......................................................................... 1. . arcticus Holmgren
5(4). . (. . 16, 14).
1.2 .
9 ....... 3. . suilis Humala
6(3). ,
(. . 16, 12). , 0.19
. .... 4. . volcanus Humala

41?,

., ., , - , -

2. Allomacrus jakuticus Humala, 2002

,

. .

3?

( )

22 r3.

- 22, 26.VII.1990, 30-31.VII.1990 (),

, . - 4.VIII.1988 (), 1 .

3. Allomacrus subtilis Humala, 2002

. .
.

5.VIII.1970

, . . . ,
(), 1? (); , : Pieni

95

Kotavaara, , 22. 20-25.VIII.1997, 11-20.VIII.1998

(Tietavainen).

4. ENTYPOMA Foerster, 1868

Foerster, 1868, 1871; Thomson, 1888
= Entelechia Foerster, 1871
= Entypomus Thomson, 1888
= Entelechius Thomson, 1888

4. Allomacrus volcanus Humala, 2002

. -.

. , , 27.VII.1973 (
), 1 ().

1(6). .
2(3). .
. ( ) .............................. ..
......................................................................... 1. . ferale Rossem ()
3(2). .

3. ROSSEMIA Humala, 1997

Humala, 1997: 297-299; Wahl, Gauld, 1998: 292.

= Sweaterella Wahl et Gauld, 1998

4(5). .
-.
5.6 , - 4.4 .....

1. Rossemia longithorax Humala, 1997 (. 19)

......................................................................... :...... 6. . suspiciosum Foerster

5(4). .
2-5 . ........................... .
.............. ............... ......... ............................ 2. . frontosum Rossem ()

= Sweaterella shakeyi Wahl et Gauld, 1998

.
.

(!). .

. , "", 3 2,

7(8). .

(); , , 1 ,
26.VII.1973 (); , 15 ,
12, 29.VI.1996 ().

, .

2 ,
, , .

-,

3. . prominens Kolarov ()
8(7). .
.

.
.

9(1 0). 3-6(7)-

, .

3.7-4.6 ............... 5. . robustum Foerster

10(9). 3-6(7)- . , .

, , ,
. 3.1-3.3
, .................

4.

robustator Aubert

1. Entypoma ferale Rossem, 1988

. .

2. Entypoma frontosum Rossem, 1988

. .
. 19. Rossemia longithorax (: , 1997)

96

. 12 (). ., -
, . . . 5.VII.l977 ().
4.

..

97

5.

3. Entypoma proininens Kolarov, 1985

. ().

4. Entypoma robustator Aubert,

KENTROTRYPHON

1. Kentrotryphon longecaudatus

StroI,

1902

Strol,

1902

. ().

198

. .
. 5 ~, 14. . ();

6.

APOCLIMA Foerster, 1871

(.

21)

APOCLIMA

( .); .

1(2). ,

5. Entypoma robustum Foerster,

1~871 (.

. .
. 25 ~, 50 . .;
( ., ), , , .
.

20)

3- 2 (. .

.................................................................................. 2.

3, 2) .......... .
signaticorne Foerster

2(1 ). , , . ............... .
................................................................................... 1. . haeselbarthi Rossem

Entypoma suspiciosum (Foerster, 1871)

1. Apoclima haeselbarthi Rossem, 1987

=Bassus remotus Marshall, 1896

. ~.

. .

. ~,

21 . , .;

2. Apoclima signaticorne Foerster, 1871

(), .

(. .

3, 2)

. .

. ~, 22. , ., ,.
- . ., ..<;~ ,
, .

98

20. Entypoma robustum (: Townes, 1971)

4*

21. Apoclima sp.

(:

Townes, 1971)
99

3.

HELICTINI Gupta, 1987

15(11).
16(1 7).

HELICINJ

1(2).

2.7-4.3

. . (

0.21-0.94 ................................................................ .4. Aperileptus Foerster

. .

5(8).

1.1-2.1

................................................................................... 6.

emiphanes

21(24).
22(23).

1.8

. 7-8..: .

( .)

8(5).
9(10).

................................................ 19. Epitropus Rossem

23(22).

Catastenus.

0.5-0.7

1.4-3.8 .
1.5-2.2 . .............................................. 1. Aniseres Foerster

14(13).

, -
.

1.9-3.6

100

0.15

.......................................................... 2.

Pantisart~rus

Foerster

.................... 10. Plectiscidea Viereck

0.3

2-

2- 3- .

2, 2).

4.2-6.0
................................................. 10. Plectiscidea Viereck

, , .

26(27).

, .

0.10-0.28

. .
, .
, .

6-9(11) .............. .
.................................................................................... 17. Eusterinx Foerster

27(26).

, .
, .

2-

24(21). 2- 3- .
25(28). , , ,
80-90,

13(14).

, .

.
.

.............................. .

11(15).
12(20).

4 .....
...................................................................................... 11. Dialipsis Foe1ster

..................................................................................... 5. Pantomima Rossem

10(9).

. . ,

1.5--1.9

(. .

Foerster

. ,

2.7-3.5

. ,

. , .....

7(6).

4.0-5.5

.)

; ,

.. 12. Atabulus Rossem

( )

0.13-0.22
20(12).

6(7).

........................... 10. Plectiscidea Viereck

....

.................................................................................... 7. Megastylus Schi0dte

4(3).

4.0-6.5

. 1- ,

0.9-0.30

30)

. ( ,

, ,

. (

17(16).
18(19).
19(1 8).

3(4).

. , .

,
.

2(1).

, ,

.)

28(25).

......................................................................... 8. Helictes Haliday

, ,
, .

29(30) .

30(29).

............................ 3. Proclitus Foerster

, ,
.

31(32).

3.5

101

1.6 .
0.12 ............................................ 13. Catastenus Foerster

32(31). .
33(34). ( ). :I
, ~ . .
........................................................................... 16. Gnathochorisis Foerster
34(33). ( Fetialis alacris ).
.

35(40). ; .
36(37).
. ,

..... 15. Phosphoriana Rossem

37(36). . .
38(39). , 053 . ....
............................................................................................ 9. Fetialis Rossem
39(38). , 018-023 .
, (
)

4(3). . 0.44-0.74 .
18-20 . 0.5 ....................................................................................... 2. . pallipes Foerster

1. Aniseres caudatus Humala, 1997.

Humala, 1997 : 299
. .
. 15 ~, 3 . ; .

2. Aniseres pallipes Foerster, 1871

(.

22)

= Aniseres lapponicus Jussila, 1994. Syn. nov.

Jussila, 1994: 117;
. .
. 151 ~, 63 . ., ,
.; .

......................................... 14. Symplecis Foerster

40(35).
41(42). .
6.3-7 .
, .

3. Aniseres paradoxus Rossem, 1991

. .
. 3 ~ , 1 . .

................................. 14. Symplecis Foerster

42(41). .
43(44). , 7-9
. ,
.

........ .
............................................................................. 15. Phosphoriana Rossem
44(43). 7
( Plectiscidea helvola)
. .

45(46). , . . .............. ..
.................................................................................... 18. Proeliator Rossem
46(45). . . ,
......................................................................... 10. Plectiscidea Viereck

1.

ANISERES Foerster, 1871

Foerster, 1871; Rossem, 1980, 1991; Jussila, 1994: 117; Humala, 1997: 299

ANISERES

1(2). , 3.7-3.8
1.2-1.3 . 18
. ... 1. . caudatus Humala
2(1). .
3(4). , 0.35 . ............................................................. 3. . paradoxus Rossem
102

22. Aniseres pallipes (: Townes, 1971)

103

2. PANTISARTHRUS Foerster, 1871

= Pantisarthus pseudochropus Strol, 1903

= Aniseres subalpinus Strol, 1903

Foerster, 1871; Strol, 1903; Rossem, 1980, 1987; 1991.

. .

. 122 2, 71 . ., ,

Pantisarthrus lubricus Foerster, 1871

PANTISARTHRUS

., ; (); ,
.

1(2). , 2.7-3.0
; .
. ,
. 0.16 .... 2. . gracilis Rossem

2(1). 2.5
, .

3(4). ,
. ,
. ( , , -

.) ........................... 5. . rudepunctatus Strol

4(3). , .

5(6).
. 2.0-2.6 ..................................................................... 1. . dispar Rossem
6(5).

4. Pantisarthrus luridus Foerster, 1871 (. 23)

= Pantisarthrus ochropus Foerster, 1871

= Pantisarthrus pseudochropus Strol, 1903

= Pantisarthrus grossepunctatus Strol,

1903

. .

. 19 2, 22. , .,
, , .

5. Pantisarthrus rudepunctatus StroI, 1903

. .

.
1.4-2.3 .

7(8).

8(7).

.
1.8-2.3 ............................... 3. . lubricus Foerster


. 1.4-2.5 .......................................................... .4. . luridus Foerster

1. Pantisarthrus dispar Rossem, 1980

. .
. 18 2, 4. ., ,
.; .

2. Pantisarthrus gracilis Rossem, 1987

. .
. 2 2, 3 . , .,
; , .

3. Pantisarthrus lubricus (Foerster, 1871)

= Aniseres lubricus Foerster, 1871

= Pantisarthrus inaequalis Foerster,

1871
= Pantisarthrus ochropus Foerster, 1871

104

23. Pantisarthrus luridus

(: Townes, 1971)

105

3.

PROCLITUS Foerster, 1868

(.

24)

16(17).

, , -

=Cryptus (Clepticus) Haliday, 1838

= Plectiscus (Proclitus) Thomson, 1888

= Aclastoneura Kriechbaumer, 1896
= Mischoxorides Ashmead, 1900

17(16).

18(19).

................................................................................ 6.

Proclitus praetor Haliday, 1838

PROCLITUS

19(18).
,

2.5-3 .0
.................................................................... 3. . comes Haliday
, 0.6-0.7 ,
2.7-4.5

3.5-4.5
0.59-0.69
. .. ...8. . praetor Haliday
6(5). , 2.7-3.0 .
0.63-0.65 . ....................................................... 2. . ardentis Rossem
7(2). , 0.6 .
8(13). , , ,
5(6).

,
.

2.4

...................................................... 7. . paganus Haliday

0.37-0.48 .

0.37-0.47

... 5. . fulvicornis Foerster

12(11). 0.48 .
, .
. ,

13(8).

... .4.

, 0.26
. .
, .
. ,

15(14).

, ........ 11. . zonatus (Gravenhorst)

, 0.36-0.48
. .

106

0.50

....................................................... .9.

rudis Foerster

, ,

22(21).

2.5

... 7. .

paganus Haliday

23(24). ~ , 0.45 ,
.

3.0

, ,

............................................................................ 1.
24(23).
25(26).

.....
subsulcatus Foerster

.................................. .
......................................................................... 11. . zonatus (Gravenhorst)

26(25).

.
.

27(28).

4.0-4.7

. ...... 8. .

28(27).

4.0

praetor Haliday

3.5-3.7
..................................................................................... 1. . ardentis Rossem
30(29). 3.1-3.3
..................................................................................... 2. . attentus Foerster
29(30).

edwardsi Roman

, ,

14(15).

. ,

, .

2.0

0.27-0.31

10(9).
11(12).

0.42-0.48 .
2.7-3.0 .
( 0.54
) ....................................................................... 2. . attentus Foerster
0.39 .

20(1).
21(22).

9(10).

................. ..
fulvipectus Foerster

0.6-1.05 .
0.8-1.05

4(3).

, .

1(20). .
2(7).
3(4).

, ,

1. Proclitus ardentis Rossem, 1987

. .
. 27 S?, 2 . , ., .
, ; .

2. Proclitus attentus Foerster, 1871

= p,-oclitus quaesitorius Foerster, 1871

= Proclitus mesoxanthus Foerster, 1871
107

 (), . (. . ); ;
(); .

4. Proclitus edwardsi Roman, 1923

. .

52 .

., .;

Brachypeza radiata.

5. Proclitus fulvicornis Foerster, 1871

=Proclitus periculosus Foerster, 1871

= Proclitus inquietus Foerster, 1871
= Proclitus cupidus Foerster, 1871
= Proclitus evacuator Foerster, 1871

=Proclitus icarinatus Foerster, 1871

= Plectiscus (Proclitus) heterocerus Thomson, 1888
=? Proclitus ruficaudator Aubert, 1963; Foerster, 1871
. .
. 2, 2. , . (); ;
.
. 24.

Proclitus sp. (: Townes, 1971)

= Proclitus fossulatus Foerster, 1871

= Proclitus validus Foerster, 1871
1871
= P1oclitus gracilentus Foerster, 1871
= Proclitus procerulus Foerster, 1871
= Proclitus melanocephalus Foerster, 1871
=Proclitus substriatus Foerster, 1871
= Proclitus leptosomus Foerster, 1871
= Proclitus sinceus Foerster, 1871

_= Proclitus stenofaster Foerster,

. 2, 2. ., ,
., ., . , . ;
(); .

3. Proclitus comes (Haliday, 1838)

Cryptus (Clepticus) comes Haliday in Curtis, 1838
= Proclitus macrurus Foerster, 1871
= Proclitus perditorius Foerster, 1871
= Proclitus caudiger Foerster, 1871
= Proclitus pallens Foerster, 1871
. .

.
108

103 2, 2. , ., .

Proclitus fulvipectus Foerster, 1871

= Proclitus fulvocingulatus Strobl,

1903

. .
. 3 2 . , .

7. Proclitus paganus (Haliday, 1838)

Cryptus (Clepticus) paganus Haliday in Curtis, 1838
= Proclitus providus Foerster, 1871
= Proclitus autumnalis Foerster, 1871
= Proclitus curiosus Foerster, 1871
= Proclitus instigator Foerster, 1871
=Proclitus contrubator Foerster, 1871
= Proclitus clype-is Foerster, 1871
= Proclitus dimidiatus Foerster, 1871
= Plectiscus (Proclitus) longitarsis Thomson, 1888
. .

. 10

2, 49. ., ,

., ., , . (),

, ; (, , , .

8. Proclitus praetor (Haliday, 1838)

Cyptus (Clepticus) t Haliday in Curtis, 1838
= Proclitus grandis Foerster, 1871

109

2(1).

. .

. 79 S?, 71 3. , ., .
(); (), , .

9. Proclitus rudis F oerster, 1871

= ?Proclitus exilis Foerster, 1871

= Proclitus definitus Foerster, 1871

. .

10. Proclitus subsulcatus Foerster, 1871

. .
. 1. ().

11. Proclitus zonatus (Gravenhorst, 1829)

Plectiscus zonatus Gravenhorst, 1829
= Proclitus absconditus Foerster, 1871
= Proclitus contemptiilis Foerster, 1871
= Proclitus punctatus Foerster, 1871
= Proclitus inferior Foerster, 1871
= Proclitus displicitus Foerster, 1871
= Proclitus sordidus Foerster, 1871
=.Proclitus humilis Foerster, 1871
= Proclitus maginatus Foerster, 1871
= Proclitus inaestimailis Foerster, 1871
= Proclitus denticulatus Foerster, 1871

Aperileptus a!ipalpus (Gravenhorst, 1829).

APERILEPTUS

1(2). .
.
, ,

110

0.5

, ,
.

0.23-0.28 . ..
.................................................................................. 7. . plagiatus Foerster
7(6). , 0.33-0.46 ........... 4. . infuscatus Foerster
8(5). , 0.28 .
9(10).
0.32-0.35 .......................... 2. . flavus Foerster
10(9). 0.35 .
11 (12). , 0.26-0.28 .
0.41-0.43 ............. 5. . melanopsis Foerster
12(11). .
13(16). , 0.5 .
14(15). , 0.30-0.38 ........... 6. . microspilus Foerster
15(14). , 0.42-0.43 ............ 11. . viduatus Foerster
16(13). , 0.35-0.48 .
17(18). 0.41-0.43 .
0.35-0.37 ................ 9. . tricinctus Foerster
18(17). 0.31-0.37 .
19(20). , 0.40-0.48 . ......................... 1. . alipalpus Gravenhorst
20(19). , 0.37-0.39 .
......................................................... 3. . impurus Foerster

= Aperileptus penetrans Foerster, 1871

Gravenhorst, 1829; Foerster, 1871; Thomson, 1888; Rossem, 1985; Jussila,

1994
= Plectiscus Gravenhorst

4(3).
5(8).

0.88-0.91
.. 8. . rossemi Jussila

=Aperileptus a!ipalpus Foerster, 1871

APERILEPTUS Foerster, 1869

. ,
.

1. Aperileptus albipalpus (Gravenhorst, 1829)

. .
. 2 S?, 26. ; .

3(4).

6(7).

1871
= Proclitus visitator Foerster, 1871
= Proclitus cautus Foerster, 1871
= Poclitus unicinctus Foerster, 1871
= Pl-oclitus spectailis Foerster, 1871
= Proclitus litigiosus Foerster, 1871

. .

0.21-0.28

= Proclitus zelator Foerster,

4.

......................................................................... 10. . vanus Foerster

= Aperileptus fungicola Foerster, 1871

= Aperileptus placidus Foerster, 1871
= Aperileptus tutorius Foerster, 1871
= Aperileptus vacuus Foerster, 1871
= Aperileptus custoditor Foerster, 1871
= Aperileptus frontalis Foerster, 1871
= Aperileptus impactus Foerster, 1871
= Aperileptus extirpator Foerster, 1871
= Aperileptus euryzonus Foerster, 1871
= Aperileptus conformis Foerster, 1871
= Aperileptus nigricarpus StroI, 1904
. .
. 44 S?, 17 . ., ,

., - , .,
. , ., , ;
.

111

2. Aperileptus flavus Foerster, 1871

= Aperileptus advesarius Foerster,

1871

= Apeileptus vilis Foerster, 1871

= Apeileptus meritus Foerster, 1871
= Aperileptus rufus Strol, 1903

. .

. 12 S? , 4. , ., ; .

3. Aperileptus impurus Foerster, 1871

= Aperileptus immundus Foerster, 1871

= Aperileptus electus Foerster, 1871
= Aperileptus secretus Foerster, 1871
= Aperileptus inamoenus Foerster, 1871

. .
. 1 S?. .

4. Aperileptus infuscatus Foerster, 1871

. .

. 111 S? , 55 . , - ;

9. Aperileptus tricinctus Foerster, 1871

= Aperileptus subsignatus Foerster,

1871

. .

4 S?.

; (), .

10. Aperileptus vanus Foerster, 1871

= Plectiscus (Aperileptus) oliquus Thomson,

(.

25)

1888

. .

203 S? ,

142. ., ,

., , . (. ), . (
), ., ; (),
, , .

(Mycetophilidae ),
hyllodon tomentosus.

Mycetophila hetschkoi
Ramaria flava,

11. Aperileptus viduatus Foerster, 1871

. .

11 9

, .; .

5. Aperileptus melanopsis Foerster, 1871

. .

. 3 S?. , . ; ..
. Aperileptus microspilus Foerster, 1871
= Aperileptus spoliator Foerster, 1871

. .

. 7 S?. , .;

(), .

7. Aperileptus plagiatus Foerster, 1871

. .
. 3 S? . , .

8. Aperileptus rossemi Jussila, 1994

Jussila, 1994: 115; Humala, 1997: 299

. .
. 14 S? , 7 . , .;
.

112

25. Aperileptus vanus (: Townes, 1971)

113

5.

PANTOMIMA Rossem, 1990

16(17).

, 2-
3- . ,

Rossem, 1990: 314

2,3

1. Pantomima festata Rossem, 1990

. .

4,5

. -

1,

. ................... .. ............................................................... 7.

17(16). ,
. 5

HEMIPHANES Foerster, 1868

Hemiphanesflavipes Foerster, 1871.

- .

............ ..
townesi Rossem
1,4

1,3

. ........................................................................ .

18(15).

.................................... ..
performidatum Rossem

, , ,
,

HEMIPHANES

1(10). .
2(5). .
3(4). (8)9-12 .
. .

................................................................................... 2.

4(3).

gravator Foerster

. .
........................ ..

................................................................................... 3.

hortense Rossem

3-

..................................................................................... 1.

1.

emiphanes flavipes

14 S?

Foerster, 1871

(.

flavipes Foerster

26)

, . ., , ,

., , , ;
(- .), , .

5(2). .
6(7). ,
3 . .................................. ...................................... 4. . inusitatum Rossem
7(6). , 5,0-7,6
.

8(9).

, 10-11 .
, ,
. -

9(8).

...................................................................... 7. . townesi Rossem

(8)9-10(11) .
, , ,
, .

........................................................ ..
. flavipes Foerster

. ............ ... .............. ..... ..... .. ....... ........ ..... .... ......... ......... 1.

10(1). .
11(12). .

................................................................................... 2. . gravator Foerster

12(11). . .
13(14). .
..... 5. . montanum Rossem
14(13). .
.

15(18).

, ,
.

114

26. Hemiphanesflavipes

(: Towes,

1971)
115

2. Hemiphanes gravator Foerster, 1871

. .

. 1 ~ 8 ., ;

3. Hemiphanes hortense Rossem, 1987

.

4. emiphanes inusitatum Rossem, 1987

.

5. Hemiphanes montanum Rossem, 1987

.

. Hemiphanes performidatum Rossem, 1988

. 12 ~ , 9. .;
, .
.

7. Hemiphanes townesi Rossem, 1980

. .
. 7 ~ , 13 . , ;

27. egastylus
1-2 - . cuentator (1 - , 2 - ); 3-4 -
, 3 -. pectoalis, 4 - . impesso; 5-6 -
, 5- . excublto, 6- . impesso; 7-8- : 7- .
pectoralis, 8- . impesso (: Rossem, 1974)

7.

MEGASTYLUS

Schi~dte,

1838

=ldioxenus Foerster, 1868

= Dicolus Foerster, 1868

= egalostylus Schulz, 1906
= ldioxenus Foerster, 1868
=Myiathridea Viereck, 1914
= Miomeroides Kiss, 1924

Megastylus cruentator Schipdte, 1838.

MEGASTYLUS

1(14).

2(3). 2/3,
(. 27, 7). ............ ;........................... 8. . pectoralis Foerster
3(2). ,
(. 27, 8).
4(5). .
.

1,5 . 5 . ............................. 2. . flavopictus Foerster

116

5(4). .
6(9).
0,25-0,28 , . ( Dicolus).
7(8). 8-9
. 7,0-8,6 .
. .

. ,

. . ...................................................................... 7. . impressor Schi!i}dte.

8(7). 7,5
. 12,3 .
,

.
. ,

.
. .............................................................. 5. . elegans Rossem
9(6).
0,25--0,28 . ( Megastylus).
10(11). - . ...................... 3. . orbltatior Schi!iidte

117

11(10).

, .
-.

12(13).

,
.
-

13(12).

.... 1.

cruentator Schidte

, .

14(1). .
15(24).

................................ 3.

3,5-5,0

orhitator Schidte

16(17).

0,25-0,28 . ( Megastylus Schidte).

.

. -
, -

17(16).

. ....................... 2. . flavopictus Foerster

,
.

18(21).

26(25).
27(28).

0,7,
(. 27, 3). 3,3-5,5 .

..................... 8. . pectoralis Foerster

28(27). (. 27, 4).

29(30). 0,56
,
. ,

. (.

27, 5).

........................................................ 6. . excubltor Foerster

30(29). 0,33-0,43
, . ,

27, 6). ................... .

................................................................................ 7. . impressor Schidte

(.

1. MEGASTYLUS Schi0dte, 1838

Megastylus cruentator

Schi~dte, 1838.

19(20).

4,1-5,0 .
2,5-3,9 . ,

. ,
, -.

3,0-3,1 ,
5,0 .......................... 3. . orhitator Schidte
20(19). , 5,1-7,0 .
, ( 4,0-7,0 ).

1. Megastylus (Megastylus) cruentator Schi~dte, 1838 (. 28)

= mediator Schi!(Sdte, 1838
= nigriventris Foerster, 1871

3,0-3,1
21(18).

3,5-5,0

............... 1.

cruentator

Schidte

-
.

22(23).

(OOL : POL) = 3 : 2 3 : 3.

, .

suecicus Rossem,

-. ,

.................... .
.................................................... :............................ 3. . orbitator Schidte
23(22). OOL : POL = 4 : 3.
, , . rufipleuris Foerster.
,

24(15).

................................. .4. . suecicus Rossem

6,0-10,0

. (

25(26).

Dicolus

0,25-0,28

17,5

118

Schidte)

3,7
..................................................................... 9. . tenellus Rossem

28. l'vfegastylus cruentator

(: Townes, 1971)

119

= conformis Foerster, 1871

=fuscicornis Foerster,

1871
= Cryptus (Helictes) cruentatus Haliday, 1838
? = pleuralis Thomson, 1888

5. Megastylus (Dicolus) elegans Rossem, 1983

.

. 79 2 , 64. ., ,

., ., ,
, , ., .,

, ~ , ; (), ,
, , ; , , .

2. Megastylus (Megastylus) flavopictus Gravenhorst, 1929

= lectiscus flavopictus Gravenhorst, 1929
1838
= My1iarthrus cingulator Foerster, 1871
= Myriarthrus aemulus Foerster, 1871
= Myriarthrus flavopictus Foerster, 1871
= Myriarthridea cingulator Townes, 1971

= lineator Schidte,

. 19 2 , 14. , .,

~ , ., ,
; ( ., ), (),
.

3. Megastylus (Megastylus) orbitator Schi~dte, 1839

= pumilio Foerster, 1871
= retroligatus Foerster, 1871
= leptoderus Foerster, 1871
= pauxillus Foerster, 1871
=Myriarthrus rufipleuris Foerster, 1871
? =facialis Foerster, 1871
= esoleptus maderensis Wollaston, 1859

Dicolus

( .), . (. . ),
., ; , ,
, ; (); , ,
-.

Megastylus (Dicolus) excubltor Foerster, 1871

excuitor

. .

2. DICOLUS Foerster, 1868

Dicolus impressor Schi~dte, 1838.

112 , 1 .

, ; (), (

.); , .

7. Megastylus (Dicolus) impressor

Schi~dte,

1838

= Dicolus insectator Foerster, 1871

. .

82

, 22.

., ,

., ., ,
, ; (), .

8. Megastylus (Dicolus) pectoralis Foerster, 1871

= Dicolus subtiliventris Foerster, 1871
=? Dicolus hirticornis Strol, 1903
. .

98 2

, 90 .

, .,

., , . , ,
, ; () , ,
.

9. Megastylus (Dicolus) tenellus Rossem, 1983

. .

8.

HELICTES Haliday, 1838

(.

29)

= Myriarthrus Foerster, 1869

= Idioxenus Foerster, 1868

= egastylus Holmgren, 1855
= Megastylus (Helictes) Thomson, 1888

4. Megastylus (Megastylus) suecicus Rossem, 1983

Foerster, 1871

. .

. 88 2 , 34. ., ,
., ., .

Helictes erythrostoma (Gmelin, 1790).

1(4).
2(3).

HELICTES

5- .

5-7-

.................................. .
............................................................................ 4. . erythrostoma Gmelin
3(2). 5-8- ........................... .
.. .......................................................................... 6. . incongruens Rossem
4(1). 5- .

120
121

329 1710'

., , ,

., - , .,

., ., ,
(),

, , ;

; , , .

2. Helictes carinata Humala, sp. n.

(. .

3, 5)

. .
. 40'. ; .

3. Helictes conspicua (Foerster, 1871)

Idioxenus conspicuus Foerster, 1871
= Idioxenus inquilinus Foerster, 1871
= ldioxenus intricator Foerster, 1871
= Idioxenus tetraglyptus Foerster, 1871
=elictes nigricoxus StroI, 1903
. .

200'.

, .;

4. Helictes erythrostoma (Gmelin, 1790)

.

29. Helictes sp.

(:

Townes, 1971)

5(8). 6- .
6(7). 6- ..... 5. . fabularis Rossem
7(6). 6- (. . 3, 4). ............................................................... 7. . karelica sp. n.
8(5). .
9(10). 6-8- ........................... .
..... ........ ..... ................... .. ............ .. .... ........ ... ............. 1. . borealis Holmgren
10(9). 9- .
11(12). 6-9- .. 3. . conspicua Foerster
12(11). 6-10- , (. 3, 5) ..................................... 2. . carinata sp. n.

Ichneumon erythrostoma Gmelin in Linnaeus, 1790

Plectiscus erythrostoma: Gravenhorst, 1829
= Cryptus (Helictes) fulvicornis Haliday, 1838
= Idioxenus mediator (Schi~dte, 1838) sensu Foerster, 1871
= Idioxenus inaequalis Foerster, 1871
. .
. 31 . , .,
., , ; , .

5. Helictesfabularis Rossem, 1987

. .
.

160'.

, ,

, .

(Humala, 1997)

1. Helictes borealis (Holmgren, 1855)

Megastylus borealis Holmgren, 1855
= Idioxenus coxalis Foerster, 1871
= ldioxenus propinquus Foerster, 1871
= Idioxenus invalidus Foerster, 1871
=1dioxenus variator Foerster, 1871
= Megastylus (Helictes) pilicornis Thomson, 1888
. .

122

6.

elictes

incongruens Rossem, 1987

7. Helictes karelica Humala, sp. n.

.

59 , 260'.

(.

3, 4, 8)

; .

123

9.

FETIALIS Rossem, 1990

Fetialis alacris

15(16).

Rossem, 1990.

4,0-4,8 ] .......... ..
............................................................................. . (). suteres (Thomson)
16(15). 2,0-2,4

1. F etialis alacris Rossem, 1990

.

1,4-1,8
. [ 0,22-0,27 .

0,20-0,22 .
4,0-4,6 (4,8) .
2,0-2,3 ..
. (). indomita Rossem
18(17). 0,25-0,27 .
4,3-4,6 .
2,2-2,4 .......
.. ............................................................................ . (). moerens (Foerster)
19(14). 5,0-6,5 .
20(31). 5,0-5,6
17(18).

10. PLECTISCIDEA Viereck, 1914

Viereck, 1914: 118; Aubert, 1968: 40-41; Townes, 1971 :196-197; Rossem,
1987: 62-86.
= Plectiscus auct. 1914 .
Plectiscidea collaris (Gravenhorst, 1829).
PLECISCIDEA

1(2). 0,72-0,85 .
[ 3,6-4,5
. 1,8-2,5
.

21 (22).

2,3-3,6 .] Fugatrix Rossem ........................................ .

.......................................................................... . (F.) communis (Foerster)
2(1). 0,70 . (
Plectiscidea Viereck.)
3(40). , 0,09-0,30
.

5(4).
6(13).
7(10). 4,0-4,8 .
1,4-1,8
.

8(9). 4,8
............................................................................ . (). istriata (Thomson)
9(8). 4,3 ....
.................................................................................... (.) koponeni sp. n.
10(7). 5,8-6,3 .
2,1 -2,5

22(21).

124

2,1

0,41 . 0,21 ................................................................................ . (). tener Foerster

;
.

23(24).

0,21

5,5
24(23).

7,3

. [

.
.

...............

. ().

7,3

2.0
parvula (Foerster)

25(28).

1,4-1,8

0,19-0,22 .
5,0-5,6 .
1,4--1,7 .
................................................... . (). obscura Rossem
27(26). 0,23-0,28 .
5,0-5,5 .
1,4-1,9 .
- ....
..................................................................................... . (). spuria Rossem
28(15). 2,0-2:5
26(27).

11 (12). .
................................................................................... . (.) aquilonia sp. n.
12(11). -. ............................... . (.) fuscifemur sp. n.
13(6). , 0,18 ...................................... .
14(19). 4,0-4,8 .

5,0

4(5). 0,09 . [
, 6,0
, ,
. , ,
. 2,3
, ] ... . (). nemorensis Rossem
, 0,10 .
0,14-0,18 .

, . [

0,24-0,27 .
5,2-5,6 .
2,0-2,3 ........................... .
......................................................................... . (). tenuicornis (Foerster)
30(29). 0,28-0,30 .
5,0-5,6 .

29(30).

125

2,2-2,5

................................................................................

31(20).
32(35).
33(34).
34(33).

.......................... ..
cinctula (Foerster)

6,0-6,5 .
, 19-0,23 .
1,4-1,7
. 6,0-6,5 . [ ] .......................................... . (). obscura Rossem
2,0-2,6
. 6,0

...............................................................................

35(32).
36(37).

().

0,25-0,30

().

amicalis (Foerster)

1,6-2,0
0,25-0,30 .
6,0-6,5 ] ........................... .
. [

.......................................................................... (). melanocera (Foerster)

37(36). 2,4-2,7 .
38(39). 6,5-7,0 .
2,4-2,5 ......................................................................... . (). helvola (Foerster)
39(38). 5,6-6,0 .
2,7 .
................................................................................. (). vagator (Foerster)
40(3). , 0,30 .
41(62). .
42(45). , 3,1-3,7
.

0,3 .
5,2 .
3,1-3,5 ............................................ ..
......................................................................... (.) canaliculata (Foerster)
44(43). 0,5 .
6,6 .
3,7 ......... . (). prognathor Aubert
45(42). 3,0

43(44).

46(55).
47(48).

5,0-6,3 .
0,55 . [

6,0 .
2,7 . 4,8 .] ................................................. .
........................................................................ (). erythropyga (Foerster)
48(47). , 0,35-0,52 .
49(50). , 2,0
. [ 0,44
. 5,0
. 3,7 .] ........ . (). monticola (Foerster)
50(49). 2,3-2,9
,

51(52).

6,0 .
0,40 . [

126

4,5 ..................................................................... . (). agitator (Foerster)

52(51). 5,0-6,3 .
0,35-0,52 .
53(54). 0,14-0,16
. 0,36-0,38 .
5,2-5,3 .
2,2-2,4 ......... .
............................................................................. (). conjuncta (Foerster)
54(53). 0,17-0,19
. 0,35-0,52 .
5,0-6,3 .
2,3-2,9 ......... .
.......................................................................... (). collaris (Gravenhorst)
55(46). 5,0
2,3

56(57).

3,7

[ 0,44 .
2,4 . 4,7r
.] ................................................................ . (). foersteri Rossem

57(56). 4,2-4,6 .
58(59). 0,38 .
4,3 . [
2,3 ' . 3,7 ] ....
.. ................................................................................... (). nava (Foerster)
59(58). 0,41-0,48 .
4,0-4,6 .
60(61). 0,29-0,35 . 0,43-0,48
. 4,0-4,5
. 1- 2,4-2,7
. 4,2-5,2 ................. .
............................................................................. . (). substantiva Rossem
61(60). 0,41-0,42 . 0,41-0,43
. 4,2-4,6
. 2,3-2,6 . 3,7-4,5 .......... .
.. ....................................................................... (). crassicornis (Foerster)
62(35). .
63(64). , 7,0
. [ 0,33-0,35
. 2,7 ] ..................................................................... . (). posticata (Foerster)
64(63). , 5,8
.

0,40-0,47 .
5,0-5,7 .
1,8-2,8 .
66(67). 0,40 . [
5,3 . 0,40 .
2,7

65(78).

127

0,37 ] ......... .
.............................................................................. . (). connexa (Foerster)
67(66). 0,40 .
68(69). 1,8
, 0,32 . [
5,7 .
0,42 ] ............. . (). mesoxantha (Foerster)
69(68). 1,8

, .

, ,

, .

70(75).

0,40-0,50
71(72).

2,0
5,0 . 0,37 ] ......................... .
............................................................................. . (). mendica (Foerster)
(0,50).

72(71).

82(81).

83(84).

5,0

2,1

0,34-0,37
76(77).

1.

().

PLECTISCIDEA Viereck, 1914

2,0-2,6
...
humeralis (Foerster)

3,6-4,3

(.

30)

Plectiscidea collaris (Gravenhorst, 1829).

1. Plectiscidea (Plectiscidea) agitator (Foerster, 1871)

= lectiscus subang ulatus Foerster, 1871

=Plectiscus mesoxanthus Foerster, 1871

. .

~. . ().

2,5

0,42

, , -

77(76).

0,32-0,34 .

............................................................................

0,40

0,35 .......................................... .
.............................................................................. . (). deterior (Foerster)

75(70).

0,43

2,5
4,0 ........................ .
........................................................................ . (). eurystigma (Thomson)
84(83). 5,0 .
0,33 . 0,37-0,38

2,3
. 5,5 . 0,38 ...................................... .
.............................................................................. (). fraterna (Foerster)
0,43

3,5-4,3

74(73).

5,0 .
0,37 . 0,36
.

0,31-0,38

0,40-0,43
73(74).

.] 5,2 .................................... . (). subangulata (Foerster)

5,0-5,2 .

2,2

0,34 ........................................ .
0,33-0,35 .

2,1-2,3

(). landita

Rossem

5,0-6,6
0,42-0,47 . 3,5-4,5 ] .................................... .
............................................................................. (). terebrator (Foerster)
78(65). , 0,33-0,38 .
4,3-5,4 .
1,6-2,6 .
79(80). 4,3-4,6 .
[ 0,32-0,37 .
1,6-1,8 ] ........
................................................................................... . (). ventosa Rossem
80(79). 5,0--5,4 ~.
81(82). 5,4 .
0,36 . 0,31
0,34-0,37

. [

. [

128

5.

..

30. Plectiscidea sp.

(:

Townes, 1971)
129

9. Plectiscidea (Plectiscidea) conjuncta (Foerster, 1871)

2. Plectiscidea (Plectiscidea) amicalis (Foerster, 1871)

= Plectiscus sodalis Foerster, 1871

= ?Plectiscus flavicoxis Foerster, 1871

. .

. .

1 9.

().

. .

. .

4. Plectiscidea (Plectiscidea)
= Plectiscus

istriatus

istriata

16-19.VIII.1991
Turku, 712:234,

11. Plectiscidea (Plectiscidea) crassicornis (Foerster, 1871)

. .

(Thomson, 1888)

4 9, 1

., .

. .

. ; .

5. Plectiscidea (Plectiscidea)

andita

13. Plectiscidea (Plectiscidea) erythropyga (Foerster, 1871)

Rossem, 1987

. .

. .
.

9.

12. Plectiscidea (Plectiscidea) deterior (Foerster, 1871)

Thomson, 1888

. .

().

10. Plectiscidea (Plectiscidea) connexa (Foerster, 1871)

3. Plectiscidea (Plectiscidea) aquilonia Humala, sp. n.

. 2 9. , . "",
(), (. , ); , :
18.IX.1981 (. Koponen leg.) DAZHU.

1 9.

7 9.

, ., .

14. Plectiscidea (Plectiscidea) eurystigma (Foerster, 1871)

Plectiscidea (Plectiscidea) canaliculata (Foerster, 1871)

=Plectiscus subtilis Foerster, 1871

= lectiscus distinctus Foerster, 1871
= Plectiscus subcurvatus Foerster, 1871

. .

. .

. .

25 9, 23

15. Plectiscidea (Plectiscidea) foersteri Rossem, 1987

. , - ,

. , ; (), ;

1. Plectiscidea (Plectiscidea) fraterna (Foerster, 1871)

. .

7. Plectiscidea (Plectiscidea) cinctula (Foerster, 1871)

17. Plectiscidea (Plectiscidea) fuscifemur Humala, sp. n.

= Plectiscus determinatus Foerster, 1871

= Plectiscus curticauda Foerster, 1871
. .

3 9 . ;

8. Plectiscidea (Plectiscidea) collaris (Gravenhorst, 1829)

. .

. 3 9. , , Vihti, Siikajarvi, (8:3),

24.IX-1.X.1971 (light trap, . Petramaa leg.); N: Helsinki, 82:382,
1.I.1978; Nurmijarvi, 710:381, 23.IX.1993 (. Koponen leg.)
DAZHU.

= Plectiscus inodulus Foerster, 1871

18. Plectiscidea (Plectiscidea) helleni Humala, sp. n.

. .

40 9, 15

130

. .

. , .,

., ; (), ,

. 1 . , Le: Kilpisjarvi, Siilastupa

11.VII.1950 (W. Hellen leg.) ZMHU.
5*

131

19. Plectiscidea (Plectiscidea) helvola (Foerster, 1871)

= Plectiscus subsimilis Foerstei", 1871

= Plectiscus petiolatus Foerstei", 1871
. .

. 3 ~. .; , .

20. Plectiscidea (Plectiscidea) humeralis (Foerster, 1871)

= lectiscus fulvus Foerster, 1871

= Plectiscus hostilis Foerstei",

1871

. .

21. Plectiscidea (Plectiscidea) indomita Rossem, 1987

. .

22. Plectiscidea (Plectiscidea) koponeni Humala, sp. n.

.

. .
. 2 ~. , .

28. Plectiscidea (Plectiscidea) nava (Foerster, 1871)

= lectiscus flavizonus Foerster, 1871

= Plectiscidea navus Aubert, 1977
. .
. 3 ~. , (
).

29. Plectiscidea (Plectiscidea) nemorensis Rossem, 1987

= Ephalmator subsimilis Rossem, 1980 ( Foei"stei", 1871)
. .
. 1 ~ . .

Humala (in press).

. .
(.

27. Plectiscidea (Plectiscidea) monticola (Foerster, 1871)

. 1 ~., N: Vantaa, 668:39, 26.VIII.1977

Koponen leg.).
23. Plectiscidea (Plectiscidea) melanocera (Foerster, 1871)

= Plectiscus vagato Foerstei", 1871

= Plectiscus ambulator Foerstei", 1871
= Plectiscus proximus Foerstei", 1871
. .
. 4 ~. ., .,
.

30. Plectiscidea (Plectiscidea) obscura Rossem, 1991

. .
. 4 ~..

31. Plectiscidea (Plectiscidea) parvula (Foerster, 1871)

= Plectiscus coxator Foei"stei", 1871

= Plectiscus nuptialis Foerster, 1871
. .

~. .

32. Plectiscidea (Plectiscidea) posticata (Foerster, 1871)

= Plectiscus pungens Foerster, 1871

24. Plectiscidea (Plectiscidea) mendica (Foerster, 1871)

. .

25. Plectiscidea (Plectiscidea) mesoxantha (Foerster, 1871)

. .

26. Plectiscidea (Plectiscidea) moerens (Foerster, 1871)

= Plectiscus xanthoneuris Foerster, 1871

= Plectiscusflavizonus Foerstei", 1871

= Plectiscus eversorius Foerster, 1871

. .

~. , ().

33. Plectiscidea (Plectiscidea) prognathor Aubert, 1968

= Plectiscidea perfera Rossem,

1988

. .

34. Plectiscidea (Plectiscidea) spuria Rossem, 1991

. .
. 7 ~ . , .

. .
. 1 ~. .

132

133

35. Plectiscidea (Plectiscidea) subangulata (Foerster, 1871)

. .

36. Plectiscidea (Plectiscidea) substantiva Rossem, 1987

. .
. 2 S?. ().

. .
. 121 S? , 3 . ., ,
., . (), ;

(, ), , , .

11.

37. Plectiscidea (Plectiscidea) suhteres (Thomson, 1888)

= Plectiscus subteres Thomson, 1888

DIALIPSIS Foerster, 1868

Plectiscus (Dialipsis) Thomson, 1888

= Parentypoma Strol, 1901

. .
. 1 S? . .

Dialipsis exilis Foerster, 1871.

1. Dialipsis exilis Foerster, 1871

38. Plectiscidea (Plectiscidea) tener (Foerster, 1871)

. .

39. Plectiscidea (Plectiscidea) tenuicornis (Foerster, 1871)

= Plectiscus brachyurus Foerster, 1871
. .
. 1 S?. .

(. .

2, 2; 31)

= Plectiscus communis Foerster, 1871

= Plectiscus (Dialipsis) crassipes homson, 1888
= Parentypomafemoratum Strol, 1901
. .
. 91 S? , 51 . , .,
., . (), .
(. . ), ., ; .I,
, , .

40. Plectiscidea (Plectiscidea) terebrator (Foerster, 1871)

= lectiscus habllis Foerster, 1871
= Plectiscus praepositus Foerster, 1871
. .

2 S? .

.; .

41. Plectiscidea (Plectiscidea) vagator (Foerster, 1871)

= Plectiscus ambulator Foerster, 1871

~
"\!}~

. .
. 1 S?. .

42. Plectiscidea (Plectiscidea) ventosa Rossem, 1987

. .

2.

FUGATRIX Rossem,

1987

Plectiscus communis Foerster, 1871.

43. Plectiscidea (Fugatrix) communis (Foerster, 1871)

=Plectiscus nigritus Foerster, 1871

= Plectiscus gilvus Foerster, 1871
= Plectiscus infirmus Foerster, 1871
134

31. Dialipsis exilis

(:

Townes, 1971) '

135

12. ATABULUS Rossem, 1988

Atabulus faustus Rossem, 1988.

1(4).
2(3).

. .

1 2.

SYMPLECIS Foerster, 1868

Symplecis alpicola Foerster, 1871.

1. Atabulus faustus Rossem, 1988

. ()

14.

7-15.V.l895

().

.
3,5-6,5 .
,

13.

CATASTENUS Foerster, 1868

Catastenusfemoralis Foerster, 1871.

1. Catastenus femoralis Foerster, 1871

(.

1 2 , 43

1,6 . .................................................. ..
.. ......................................................................... 2. S. bicingulata Gravenhorst
3(2). 7 .
,

32)

0,3

. ,

. ., ,

., ., , -

, ., , , ;
(), .

0,41-0,45

. .

SYMPLECIS

2, 1 . ............................................................................ 1. S. beaumontor Aubert

4( 1).

5(6).

6-7

. ....................

6(5).
7(8).

3. S. breviuscula Roman

0,14-0,16

.
. -

............ ..
.. .. ........ .......... ...... ..... ..... .. ........ ....... .. ............... ..... ..... .... 4. S. invisitata Rossem
8(7). 0,27 .
,

. ,

. -

.............................................................................. 5. S. paradoxa Rossem

1. Symplecis beaumontor Aubert, 1968

. .

2. Symplecis bicingulata (Gravenhorst, 1829)

esoleptus icingulatus

(.

33)

Gravenhorst, 1829

= Symplecis alpicola Foerster, 1871

= Blapticus leucostomus Foerster, 1871
= Symplecis xanthostoma Foerster, 1871
= Symplecis zonaria Foerster, 1871
= Symplecis facialis Thomson, 1888
= Symplecis basalis Brischke, 1880
. .

159 2 , 124

. ., ,

., , , .,
.

136

32. Catastenus femOl'alis

(:

Townes, 1971)

., ., . , ,

137

15.

PHOSPHORIANA Rossem, 1988

= Phosphorus Rossem, 1980

Phosphoriana rugosissima (Strol, 1903).

1. Phosphoriana rugosissima

(Strol,

1903)

Entypoma mgosissimum Strol, 1903

= Phosphorus rugosissimus Rossem, 1980
. .

(Jonaitis, Rimsaite, 2000).

16.

GNATHOCHORISIS Foerster, 1868

= Blapticus Foerster, 1868

= Laepserus Foerster, 1868
= Acroblapticus Schmiedeknecht, 1911

Gnathochorisis flavipes Foerster, 1871.

GNATHOCHORISIS

1(2). . -

. 33. Symplecis icingulata (: Townes, 1971)

, ...........................

3(4).

., , , ,

, , ; ( .,
., , ), , ; ,

3. G. flavipes Foerster.

2(1). .

. .
(, , , ,
) .....................

4(3).

1. G. crassula (Thomson)

.
. .

.. .............. .
..................................................................................... 2. G. dentifer Thomson

3. Symplecis breviuscula Roman, 1923

= Symplecis infavorabilis Rossem, 1980

1. Gnathochorisis crassula (Thomson, 1888)

. .

. ~, 2 . , ; .

. .

4. Symplecis invisitata Rossem, 1980

. .

. 12 ~, 11 . .,

., , , , -

; .

5. Symplecis paradoxa Rossem, 1988

. .
138

= Blapticus crassulus Thomson, 1888

= Acroblapticus crassulus Schmiedeknecht, 1911

54

~,

57

. , .,

, ., . , . ,
, ., ; (, ),
(), . .

2. Gnathochorisis dentifer (Thomson, 1888)

(.

34)

= Laepserus dentifer Thomson, 1888

= Acroblapticus deilis Schmiedeknecht, 1911

139

 . .

. 8 . ., ,
; .

17.

EUSTERINX Foerster, 1868

= Holomeristus Foerster, 1868

= Catomicrus Thomson, 1888
= Divinatrix Rossem, 1987
= Dallatorrea Ashmead, 1902
= Trestis Foerster, 1868
= Stroia Shmiedeknecht, 1911
= Acanthostroia Roman, 1925
= lschyracis Foerste1, 1871

Eusterinx oligomera Foerster, 1871.

EUSTERINX

1(4).
2(3).

. 34. Gnathochorisis dentife (:

Townes, 1971)

. .

. 55 5?, 35 . , .,
, , ; Kurland; .

.................................................................................... 2. Ischyracis Foerster

7(6).

4. Gnathochorisis meridionator Aubert, 1980

.
-

............................................................... 4. Divinatrix Rossem
.
. , . , 0,12-0,14 .............. .

................................................................................ 5. Dallatorrea Ashmead

12(11).

. .
( .

5. Gnathochorisis restricta (Rossem, 1980)

. .

6- -

...................................................................................... 1. Eusterinx Foerster

8(5).
9(10).
10(9).
11(12).

. .

3 .

6-7- ............. .

.), ., ;
., ; .

= Gnathochorisis terebrata Strand, 1918

. 17 5?, 12 . , . (

. 3
. ..................................... 1. Eusterinx Foerster

3(2). . 3 . .......... .
.......................................................................................... 3. Trestis Foerster
4(1). .
5(8). .
6(7). , .
3 .

3. Gnathochorisis flavipes Foerster, 1871

. .

aquilonigena), 0,19-0,25

. 6-7-
; 6-11-- , .............................. .

Holomeristus Foerster

. Gnathochorisis xanthocephala (Strol, 1903)

= Blapticus xanthocephalus StroI,

140

1903
141

1.

1(12). .
2(7).

.) ,2 . 0,18-0,20 ............. ..

EUSTERINX Foerster, 1868

EUSTERINX

3(4).

2,8

0,71

. -

. ............................................

4(3).

4.

jugorum

(Strol)

.
.

5(). ,

2,0

0,71

.
, . -

. .

.. .............. .
obscurella Foerster

................................................................................... 5. .
3,0 .
, 0,71 .

............................................................................... 5. . obscurella Foerste

18(17). .
19(20). -
(POL : OOL = 2,5 : 4,5). ( .
0,53-0,57 . 0,18-0,20 .) ............ .
................................................................................... 2. . fabulosa Rossem
20(19). -
(POL : OOL = 2,5 : 5-8,3).
21 (22). -
(POL : OOL = 1,5 : 5,0). .
.l ....... 3. . fleischeri Gregor
22(21). -
(POL: OOL = 1,5 : 3-3,8).
. ,
, .

0,53-0,58 .
, 1-0, 19 .

........................................................................ 7.

(5).

.. ................................................... ..
....................................................................................... 8. . subdola Foerster

. .
. 43 . ; , .

7(2). .
8(9). .
.

1,9 ................................................................. 7.
9(8). (7)- .
10(11). - .

pseudoligomera Gregor

.. .......................................... ..
.............................. .. ..... ........................................... . . oligomera Foerster

11(10).

2. Eusterinx (Eusterinx) fabulosa Rossem, 1990

. .

3. Eusterinx (Eusterinx) fleischeri Gregor, 1941

, -

= Eusterinx pseudoobscurella Gregor

. .

-7- ( 7-

4. Eusterinx (Eusterinx) jugorum Strobl, 1900

). 8- . , .

12(1). .
13(14).

oligomera . ................................ 1.

argutula Foerster

,l-0,18 .

2,5
14(13).

) ........................ .

8.

subdola Foerster

15(1). ,

0,12

0,5 ) .......................... . .
1(15). ,

142

5. Eusterinx (Eusterinx) obscurella Foerster, 1871

0,12

. .
. 14 S?, 2 . . , , ;
.

. ( .

17(18).

. .

( .

pseudoligomera Gregor

1. Eusterinx (Eusterinx) argutula Foerster, 1871

.
. ,

oligomera Foerster

, . ( -

Eusterinx (Eusterinx) oligomera Foerster, 1871

(.

35)

. .
. 13 S?. ; .

143

10. Eusterinx (lschyracis) permiranda Rossem, 1988

. .

4 2, 1

3.

. , .

TRESTIS Foerster

= Catomicrus F oerster, 1871

Tryphon pusillus Zetterstedt, 1838.

11. Eusterinx (Trestis) tartarea Rossem, 1982

. .

12. Eusterinx (Trestis) trifasciata (Aschmead, 1899)

= disparilis Rossem, 1982
. .

3 2, 4

. ; .

13. Eusterinx (Trestis) trichops (Thomson, 1888)

.

35. Eusterinx (Eusterinx) oligomera (: Townes, 1971)

= Tryphon pusillus Zetterstedt, 1838

= Catomicrus trichops Thomson, 1888
. .

7. Eusterinx (Eusterinx) pseudoligomera Gregor, 1941

. .
. 3 2, 5 . , - ;

31 2, 2

. , , . (),

, ; , .

4.

DIVINATRIX Rossem, 1987

Rossem, 1987: 95

8. Eusterinx (Eusterinx) subdola Foerster, 1871

= emiteles pseudominutus Strol,

1900

. .
. 5 2 , 4 . ; , .

14. Eusterinx (Divinatrix) inaequalis Rossem, 1980

. .

2.

ISCHYRACIS Foerster, 1871

1schyracis ispinosa Strol,

9. Eusterinx (Ischyracis) bispinosa

StroI,

1900.
1900

= emiteles ispinosus Strol, 1900

= Catomicrus alpigenus Strol, 1903

144

21 2, 11

. , .,

. (), , ;
.

15. Eusterinx (Divinatrix) inaspicua Rossem, 1988

. .

. .

. 10 2, 1 . , (
, . ), , ; ().

Eusterinx inaequalis Rossem, 1980.

5.

2 2 .

. ..

DALLATORREA Ashmead, 1902

Dallatorrea armata Ashmead , 1902 (. 36).
145

n71
\V

. 37. Eusterinx (Holomeristus) tenuicincta (: Townes, 1971)

. 36. Eusterinx (Dallatorrea) armata (: Townes, 1971)

19. Eusterinx (Holomeristus) refractaria (Rossem, 1982)

1. Eusterinx (Dallatorrea) circaea Rossem, 1982

."

. .
. 4 2, 4 . .

. .
. 2, 2 . ; ; ,

20. Eusterinx (Holomeristus) similis Rossem, 1991

. .
. 1 2 . .

6. HOLOMERISTUS Foerster, 1868

Eusterinx (olomeristus) tenuicincta

21. Eusterinx (Holomeristus) tenuicincta (Foerster, 1871) (. 37)

(Foerster, 1871).
17. Eusterinx (Holomeristus) aquilonigena Rossem, 1982
. .
. 12 2., 18 . , .,
r ., ., .,
, ; .

. .
. 56 2, 40 . , .,
, . (), . , ~ ,
, , ; ();
.

22. Eusterinx (Holomeristus) truculenta Rossem, 1991

18. Eusterinx (Holomeristus) minima (Strol, 1903)

. .

. .

. 3 2, 10 . .,
. , . (), , ., , ;

18.

146

PROELIATOR Rossem, 1982

(.

38)

Proeliator proprius Rossem, 1982.

147

2(1).

. 1-
. .
, ,

1,9 . ...
....... .. .... ............. ...... ........................ .... .. ..... .................... .. 2. Ortholaba Townes

1.

DIACRITUS Foerster, 1868

= Phidias Vollenhoven, 1878

= Stenolais Kriechbaumer, 1894

Foerster, 1868; riechbaumer; Vollenhoven; Momoi, 1966
Diacritus aciculatus (Vollenhoven, 1878).

.

DIACRITUS

1(2).

. 1- 2- -

2(1).

. 1- 2-

.. ........................................ 2. D. incompletus Momoi

. 38. Proeliator captiosus (:

3(4).

. .
, - .......... ..

4(3).

Dasch, 1992)

........ ........... .... .............. .......... .......... .. ... ... 1. D. aciculatus japonicus Momoi

PROELIATOR

1(2). 0.14-0.17 .
. .. ................................................................. 1. . invictus Rossem
2(1 ). 0.23-0.30 ........... ..
....................................................................................... 2. . proprius Rossem

1. Proeliator invictus Rossem, 1987

. . .
2. Proeliator proprius Rossem, 1982
. .

. 24 ~, 11 . , .,
.; , .

4. DIACRITINI Townes, 1965

DIACRIINI

1(2). , 1-
. . ,

0,22-0,57 ............................. 1. Diacritus Foerster

148

, . -,

.
(

D. aciculatus japonicus ) ..
....................................................... .1 . D. aciculatus aciculatus Vollenhoven

1. Diacritus aciculatus (Vollenhoven, 1878)

(.

39)

Phidias aciculatus Vollenhoven, 1878

Stenolabis cingulata Kriechbaumer, 1894
. .
. 23 ~ , 25 . . , ,
-; (), , ,
.

2. Diacritus incompletus Momoi, 1966

. .

2.

ORTHOLABA Townes, 1969

Ortholaba tenuis Townes, 1969.

149

1. Ortholaba tenuis Townes, 1969

(.

40)

. .
. 1 <? . , . ,
, 9.VI.1983 ().

OXYTORINAE Thomson, 1883

Gravenhorst, 1820: 379, 1829: 100, 105, 109; Haliday, 1838; Foerster, 1868:
189, 199, 209; Thomson, 1883: 910; Morley, 1907: 257, 1914: 398;
Schmiedeknecht, 1911: 2515-2517; Kerrich, 1939: 126-128; Momoi, 1965: 32-37;
Townes et al., 1965; Townes, 1971: 185; Rossem, 1980: 88-91, 1990: 316-317;
Dasch, 1992: 290-299.

poO.xytorus,
.

, ;
.
, 2,8 .
. 39. Diacritus aciculatus (: Townes, 1969)

, .

,
;

,
; ,

; 1-

0,6-0,7

, , 4-6-

;
, ,

;
.

. 40. 0l'tholaba tenuis (: Townes, 1969)

150

20 :
. armatus . luridator, . canalis, . corniger, . kamikochianus, . montanus, . nikkoensis,
. norikuranus, . obtusus, . confusus sp. n.,
(, . .)
(Gauld, 1991). ,

.
151

, ;
;
,
.

1.

OXYTORUS Foerster, 1868

Gravenhorst, 1820: 379, 1829: 100, 105, 109; Haliday, 1838; Foerster, 868:
189, 199, 209; Thomson, 1883: 910; Morley, 1907: 257, 1914: 398;
Schmiedeknecht, 1911: 2515-2517; Keich, 1939: 126-128; Momoi, 1965: 32-37;
Townes et al., 1965; Townes, 1971: 185; Rossem, 1980: 88-91; Dasch, 1992:
290-299.
= Delolytus Foerster, 1868
= Callidiotes Foerster, 1868
= Pantoporthus Foerster, 1868
= esatractodes Morley, 1907

.

Oxytorus armatus Thomson, 1883

(.

43, 9).
. , .

1(2).

OXYTORUS

............................................. .

......................................................................... 6.
2(1 ).
3(8).

luridator (Gravenhorst)

(.
.

41, 4).

, (. .

41, 6).

- .

4(7).

,
.

5(6).

, (. .

41, 11).

6(5).

................ 4.

(. .

corniger (Momoi)
41, 10),

, .
.

7(4).

..................................... 3.

8(3).

confusus sp. n.
.....
montanus (Momoi)

(. .

41, 5).

(. .

41, 7).

41. Oxytorus
:

1-

corniger, 2-

canalis;

: 3- . montanus, 4- . obtusus; : 5- . cornige, 6 . norikuranus (: Momoi, 1965); : 7- . Luridator, 8- .

armatus (: Keich, 1939); : 9 - . confusus sp. n., /0 - .

corniger

................ ;............................................................... 7.

- ( .

nikkoensis)
9(12). .
10(11). ................................................ 2. . canalis (Momoi)

11(10).

................ .
................................................................................. 1. . otusus (Momoi)

12(9).
13(14). - ........................... 8. . nikkoensis (Momoi)
14(13). - .
15(16). ,
.
. -

153
152

16(15).

3. Oxytorus confusus Humala, sp. n.

................... .

. armatus Thomson
.

.................................................................................. 1.

, .
2
, . ......................................... 9. . norikuranus (Momoi)
18(17). 2,5
, -

17(18).

................................. .
....................................................................... 5. . kamikochianus (Momoi)

1. Oxytorus armatus Thomson, 1883

Gravenhorst, 1820: 379, 1829: 100, 105, 109; Haliday, 1838; Foerster, 1868:
189, 199, 209; Thomson, 1883: 910; Morley, 1907: 257, 1914: 398;
Schmiedeknecht, 1911: 25215-2517; Keich, 1939: 126-128; Townes et al. 1965;
Townes, 1971: 185; Rossem, 1980: 88-91
. .

. 4 9, 153 . ; , , -

(): 8,5 .;
. 1,04 ;
1,56 0,5 ,
, ;
0.67 ,
.
, ; .
, , ,
,
; 2,85 ;
.

6,3

, , -

(OOL)

1,26

; ,
;

3,1

, -

37

2,6

; .

; ;
; ;

, .

2. Oxytorus canalis (Momoi, 1965) (. 42)

. .

, (. .

41, 10).

.
(. .

41, 4).

.
, , ;

4,35

0,42

3,1 ;
, 0,84
. .
. 2-ro 1,25
. ,
.
2- 3- . .
, , , ,
.

. , 3-, 4- ,
2- .
, -.

. cornige

. 42. Oxytorus canalis () (: Momoi, 1965)

154

Momoi

, .

155

. .
. : 2. , 30

17.VII.l981

().

. 59 2 , 105 . , .,
., ., .,
; , , , ,
, , .

4. Oxytorus corniger (Momoi, 1965)

7. Oxytorus montanus (Momoi, 1965)

Momoi, 1965: 32-37.

. .
. 9 2 , 4 . , .

5. Oxytorus kamikochianus (Momoi, 1965)

. .

8. Oxytorus nikkoensis (Momoi, 1965)

. .

Momoi, 1965: 32-37.

. , .
. 1 2 , 3 . ,

9. Oxytorus norikuranus (Momoi, 1965)

. .

Oxytorus luridator (Gravenhorst, 1820)

(.

1. Oxytorus ohtusus (Momoi, 1965)

42, 43)

=!chneumon luidator Gravenhorst, 1820

= Atractodes properator Haliday, 1838

= Mesoleptus ventat01- Gravenhorst, 1829
= Mesoleptus coxator Gravenhorst, 1829
= Callidiotes l-idato (Gravenhorst) forma nigicoxa Kiss von Zilah, 1924

156

43. Oxytorus luridator (: Townes, 1971)

Momoi, 1965: 32-37

. .
. 5 2 , 3 . .

 ..

(Hymenoptera):

, //
- i . .. .
. i. .,

1905. . 1-23.
(Diptera: Bolitophilidae, Ditomyiidae,
Keroplatidae, Diadocidiidae, Mycetophilidae) . :
, 2000. 84 .
. (Diptera, Mycetophilidae)
// . . 1977. . 56, . 1.
. 71-78.
.. . ., 1978. 447 .

..

..
/1 .
.: . 1984. . 3-20, 179-221.
. .., ..
Ichneumoninae (Hymenoptera, Ichneumonidae)
// . . 1996. . 75; . 8. . 1182-1194.

.., . . ~
(Hymenoptera, Ichneumonidae, Oxytorinae) ,

. .. , ,

"" //
"". : ,
.

..

(Hymenoptera, Apocrita)

1991.

31-44.

"" //

. : ,

11 Ekologija. 2000. . 3.

. 15-21.
. . . .

3.

, . 3. Ichneumonidae. .: ,
1981. 688 . ; . 129).
..

1996.

75,

..

4.

(Hymenoptera) //
756-789.

. .

1997.

50-72.

. ..

Cylloceria Schisz>dte Allomacrus Foerster

(Hymenoptera, Ichneumonidae) // . .
2002. . 81, . 2. . 370-385.

..

//

. : . .

1980.

45-60.

(Hymenoptera,
1998. . 77,

.. ,

.., . ..

.., .., . . . ,

Ichneumonidae)
. 1. . 216-223.

. // .

Oxytorinae (Hymenoptera, Ichneumonidae)

// . 1995. . 74, . 2. . 416-419.

.., .., . .
(Hymenoptera, Ichneumonidae, Tryphonoidae)
11 . . 1975. . 69. . 65-71.
: . . 4-
. .,

2000.

.. .

. .
; . 15).

3.

.,

1934. 271

Ichneumonidae

. (

.. . Ichneumonidae
. . 4. ., 1936. 535 . (
; .

16).

.. .

Ichneumonidae

. . . 5. ., 1936. 340 . (
; . 21).
.. . Ichneumonidae
. . . ., 1936 . 356 .(
; . 21).

158

. : ,

1994. 127

// -

.
, 1994. . 128-145.
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159

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absconditus, Proclitus 110

Acaenitinae 42*
Acanthostroia 141
accusator, Cylloceria 30
aciculatus, Diacritus aciculatus 149
aciculatus, Diacritus 8, 35 , 4, 149, 150*
Aclastoneua 10
Acroapticus 139

Acrodacty1a 31, 32, 45

Adelognathina 32
Adelognathinae 32
Adelognathus 31, 32
adversarius, Aperi1eptus 112
aemulus, Megasty1us, Myriarthrus 120
agitator, P1ectiscidea 54, 4, 127, 129
aino, Cylloceria 49, 54, , 88, 89*, 90, 91
alacris, Fetialis 5, 102, 124
alidipes , Proclitus 5
a1ipalpus, Aperileptus 5, 25*, 30, 50, 51,
4 , 110,111
Allornacrus 4, 15, 18, 19, 22, 24, 31, 32,
40,41,42*,0, 8,89 * ,94

alpicola, Syrnpleci~ 137

alpigena, Cylloceria 3, 88, 91
alpigenus, Eusterinx, Catomicrus 144
ambulator, Plectiscus 132, 134
arnicalis, Plectiscidea , 4, 12, 130
arnoenus, Megastylus 51
Aniseres 4, 17, 18, 19, 24, 31, 34, 40, 42,
42*,43,57,0, 72,100,102
antennatus, Oxytorus 4
Aperileptus 5, 14, 18, 19, 21, 24, 25, 2* ,
27, 32, 34,42*,43,51,55,57, ,2,
7, 70, 72,100,110
Apoclirna 4, 14, 1*, 17*, 18, 19, 24, 40,
42,0,2, 8,99,99*

aquilonia, Plectiscidea , 4, 124, 130

aquilonigena, Eusterinx 8, 4, , 141, 14
aquisgranensis, Microleptes 3, 4,
arcticus, Allomacrus 4, 4, 89*, 94, 95*
1

ardentis, Proclitus 4, 51, 4, 1, 107

argutula, Eusterinx 7, 4, 142, 143
arrnata, Eusterinx 51, 145
arrnatus, Oxytorus 9, 21, 33, 4, 151, 152,
153*, 154
Asph1agis 87
Atabulus 7, 15, 19, 24, 42*, 43, 44, ,
101, 13
Ateleute 31, 32
Atractodes 34, 4
attentus, Proclitus 4, 51, 4, 107
autumnalis, Proclitus 109
Banchinae 8*, 87
basalis, Syrnplecis 137
beaurnontor, Syrnplecis 7, 137
belokobylskii, Microleptes 3, 7
cainatus, Proclitus 109
icingulata, Syrnp1ecis 7, 28*, 30, 4, 9,
137, 138*
cingulatus, Syrnplecis, Mesoleptus 137
nodulus, P1ectiscidea, Plectiscus 130
ispinosa, Eusterinx 8, 51, 4, 144
istriata, Plectiscidea , 54, 4, 124, 130
landita, Plectiscidea , 128, 130
Blapticus 31, 32, 139
borea1is, Cylloceria 3, 1*, 17, 4, 88, 91
borealis, Helictes , 4, 122
brachycera, Cylloceria 3, 5, 89*, 90, 91
brachyurus, Plectiscidea, Plectiscus 134
breviuscula, Syrnp1ecis 7, 51, 58, 4, 137,
138
caligata, Cylloceria, Phytodietus 30, 87
Callidiotes 33, 34, 152
Campopleginae 8*
Campoplegoidae 31
cana1icu1ata, Plectiscidea , 52, 4, 12,
130
cana1is, Oxytorus 9, 151, 152, 153*, 154,
154*
carinata, He1ictes , 1*, 4 , 122, 123

n ,

; .

168

caseyi, Megasty1us 51
Catastenus 7, 14, 21, 31, 42*, 43, , 2,
100, 102, 13
Catomicus 141, 145
caudatus, Aniseres 4, 24, 57, 4, 102, 103
caudiger, Proclitus 108
cautus, Proclitus 110
Chalinoceras 87
Cidaphus 2*, 27
cinctula, Plectiscidea , 52, 4, 12, 130
cingulata, Diacritus, Stenolas 149
cingulator, Megasty1us, Myiathidea 120
cingulator, Megastylus, Myriarthus 120
circaea, Eusterinx 8, 4, 145
clypearis, Proc1itus 109
collaris, Plectiscidea , 30, 52, 53, 54, 4,
, 124, 127, 129, 130
cornes, Proc1itus 4, 24, 25*, 30, 51, 4,
10, 108
cornrnunis, P1ectiscidea, Plectiscus 7, 24,
4, 124, 134, 135
confomis, Aperileptus 111
confomis, Megastylus 120
confusus, Oxytorus 9, 151 , 152, 153*,
155
conjuncta, Plectiscidea , 4, 127, 131
connexa, Plectiscidea , 128, 131
conspicua, Helictes , 17, 4, 122, 123
contemptilis, Proclitus 110
contrubator, Proclitus 109
comiger, Oxytorus 9, 151 r 152, 153*, 155,
15

coxalis, Idioxenus 122

coxator, Mesoleptus 15
coxator, Plectiscus 133
crassicomis, Hyperacrnus 3, 30, 3, 84,
85,85*,93
crassicomis, Plectiscidea , 4, 127, 131
crassipes, Dialipsis, Plectiscus 135
crassula, Gnathochorisis 7, 4, 139
Cressonia 37, 41, 59, ,
Criptopimpla 94
cruentator, Megasty1us 5, 30, 51, 3, 9,
11, 117*, 118,119, 119*
cruentatus,
Megastylus,
Cryptus
(Helictes) 120
Cryptinae 32, 34, 4, 47, 8* ,
Cyptus (Clepticus) 30, 1
Ctenopelmatinae 27, 45, 4, 47, 8* ,
cupidus, Proclitus 109
cuiosus, Proclitus 109
cuticauda, Plectiscus 130
Cushrnania 3, 14, 15, 17, 18, 22, 34, 37,
38, 39,49, 72,83
custoditor, Aperileptus 111

Cylloceria 3, 13, 15, 17, 18, 19, 21, 23, 24,

2*,27, 28, 29, 30, 31,32, 3, 38, 39,
40,41,42*,49,54,55,57,58,0,2,
7, 70, 8,87, 89*
Cylloceriinae 3, 37, 40

Cylloceriini 3, 18, 19, 37, 39, 40, 41, 43,

49,55,59,2, 72,8
Dallatoea

delis,

8, 14, 21, 141, 145

Gnathochorisis, Aco!apticus

139
definitus, Proclitus 110
degener, Acrodacty1a 32
Delolytus 152
denticulatus, Proclitus 11
dentifer, Gnathochorisis 7, 4, , 139,
140*
deterior, Plectiscidea , 128, 131
determinatus , Plectiscus 130
Diacritinae 11, 37
Diacritini 8, 18, 33, 37, 39, 41, 59, 2, 72,
148
Diacritus 15, 18, 2*, 31, 33, 35, 41, ,
148, 149
Dia1ipsis 14, 24, 32, 42*, 43, 44, , 101,
135
Dico1us 23, 32, 11,117, 118,120
dimidiatus, Proclitus 109
Diplazon 2*
Diplazontinae 27, 29, 42*, 5
dispar, Pantisarthrus 4, 4, 104
disparilis, Eusterinx 145
displicitus, Proc1itus 11
distinctus, Plectiscus 130
Divinatrix 8, 14, 29, 141, 145
edwardsi, Proclitus 4, 51, 5, , 1, 109
egegia, Microleptes, Gnathoniella 80
electus, Aperi1eptus 112
e1egans, Megasty1us 5, 117, 121
Entelechia 31, 32, 97
Entelechius 97
Entyporna 4, 15, 19, 21, 22, 24, 31, 32, 40,
42,42*,57,2,7,8,97

Entypomus 97
Epitropus 47, , 100
erythropyga, Plectiscidea 53, 4, 12, 131
erythrostorna, Helictes 30, 4, 121, 123
Euryproctini 45
eurystigrna, Plectiscidea , 12 , 129, 131
euyzonus, Aperi1eptus 111
Eusterinx 7, 13, 14, 15, l, 17, 18, 19, 21,
22, 24, 25, 2*, 27, 28, 29, 31 , 34, 38,
39,40,41 , 42*,43, 55, 57,0, 2, 7 ,
101, 141, 142
evacuator, Proc1itus 109
eversorius, Plectiscus 132

169

excuitor,

Megastylus 5, 64, 117*, 119,

121
exilis, Dia1ipsis 7, 15*, 52, 64, 135, 135*
?exilis, Proclitus 11
Exochides 33
Exochini 33
Exochoidae 3 1
Exochus 30
extirpator, Aperi1eptus 111
fabularis, Helictes 6, 64, 122, 123
fabulosa, Eusterinx 7, 143
?facialis, Megastylus, Myriarthrus 120
facialis, Symplecis 137
faustus, Atabulus 7, 64, 136
femoralis, Catastenus 7, 21, 64, 66, 136,
136*
femoratum, Dialipsis, Paentypoma 135
ferale, Entypoma 4, 66, 97
festata, Pantomima 5, 114
Fetialis 6, 18, 43, 44, 60, 102, 124
? flavicoxis, Plectiscus 131
flavipes, Gnathochorisis 7, 51, 58, 64, 69,
139, 140
flavipes, Hemiphanes 5, 17*, 64, 114,
115, 115*
flavizonus, Plectiscus 132, 133
flavopictus, Megastylus, Myriarthrus 5,
30, 64, 116, 118, 120
flavus, Aperileptus 5, 65, 111, 112
fleischeri, Eusterinx 8, 143
foersteri, Plectiscidea 6, 127, 131
fossulatus, PJ-oclitus 108
fratema, P1ectiscidea 6, 128, 131
fontalis, Aperileptus 111
frontosum, Entypoma 4, 65, 97
Fugatrix 7, 124, 134
fulvicomis, Cryptus (Helictes) 123
fulvicomis, Proclitus 5, 52, 64, 106, 109
fulvipectus, Proclitus 5, 65, 107, 109
fulvocingulatus, Proclitus 109
fulvus, Plectiscus 132
fungicola, Aperileptus 111
fuscicomis, Megastylus 120
fuscifemur, Plectiscidea 6, 64, 124, 131
fusciventris, Cylloceria 3, 64, 66, 90, 92,
93
gilvus, Plectiscus 134
glabriventris, Microleptes, Miomeris 82
Gnathochorisis 7, 15, 18, 19, 21, 22, 24,
27,28,29, 31, 32,42*,43,45, 57, 60,
62, 102
gracilentus, Proclitus 108
gracilis, Pantisarthrus 4, 64, 69, 104
grandis, Microleptes 3, 77
gandis, Proclitus 109

170

gravator, Hemiphanes 5, 64, 114, 116

grossepunctata, P1ectiscidea 6, 66
grossepunctatus, Pantisarthrus 105
halis, Plectiscus 134
haese1barthi, Apoclima 4, 64, 99
Helictes 6, 13, 16, 18, 21, 25, 27, 30, 32,
34, 40, 42*, 43, 60, 62, 67, 101, 121,
122*
Helictinae 37, 41,42
Helictini 4, 18, 21, 22, 23, 37, 39, 40, 41,
44,45,55,58,59,62, 72,100
helleni, Plectiscidea 6, 64, 131
helvola, Plectiscidea 6, 64, 102, 126, 132
Hemiphanes 5, 14, 18, 22, 23, 25, 28, 29,
31,42*,43,45,60,62, 71,100,114
emiteloidae 3 1
hetaocerus, Plectiscus 109
?hirticomis, Megastylus, Dicolus 121
Holomeristus 8, 15*, 18, 31, 44, 141,146
hortense, Hemiphanes 5, 114, 116
hostilis, Plectiscus 132
humera1is, Plectiscidea 6, 65, 129, 132
humilis, Proclitus 110
Hyperacmus 3, 13, 14, 18, 21, 23, 24, 30,
31, 33, 34, 37, 38, 39, 57, 60, 62, 72,
73,84
hyperborea, Plectiscidea 64
Ichneumon 30,
lchneumoninae 68*
Ichneumonoidae 31
ldioxenus 32, 116, 121
ignota, Cushmania 83
immundus, Aperi1eptus 112
impactus, Aperileptus 111
imperspicua, Cylloceria 3, 90, 92
impressor, Megastylus 5, 30, 51, 64, 66,
117, 117*, 119, 120, 121
impurator, Plectiscus 36
impurus, Aperileptus 5, 65, 111, 112
inaequalis, Eusterinx 8, 64, 145
inaequalis, ldioxenus 123
inaequalis, Pantisarthrus 104
inaestimalis, Proclitus 110
inamoenus, Aperileptus 112
inaspicua, Eusterinx 8, 66, 145
incompletus, Diacritus 8, 66, 149
incongruens, Helictes 6, 64, 121, 123
indomita, Plectiscidea 6, 65, 125, 132
infavoralis, Symplecis 138
inferior, Proclitus 110
infirmus, Plectiscus 134
infuscatus, Aperileptus 5, 65, 69, 111, 112
inquietus, Proclitus 109
inquilinus, ldioxenus 123
insectator, Megastylus, Dicolus 121

insolitus, Epitropus 65
instigator, Proclitus 109
intricator, ldioxenus 123
inusitatum, Hemiphanes 5, 114, 116
invalidus, ldioxenus 122
invicta, Cylloceria 3, 64, 66, 88, 89*, 92
invictus, Proe1iator 8, 148
invisitata, Symplecis 7, 64, 137, 138
Ischyracis 8, 14, 21, 31, 141,144
jakuticus, Allomacrus 4, 65, 89*,
94,95
japonicus, Diacritus aciculatus 149
jugorum, Eusterinx 8, 64, 142, 143
kamikochianus, Oxytorus 9, 151, 154, 156
karelica, Helictes 6, 16*, 17*, 64, 122, 123
Kentrotryphon 4, 15, 18, 24, 40, 42*, 43,
60,86,99
koponeni, Plectiscidea 6, 64, 124, 132
Laepseus 139
Lamponota 3 1, 87
lapponicus, Aniseres 103
leptoderus, 120
leptosomus, Proclitus 108
leucostomus, Blapticus 137
lineator, Megastylus 120
Lissonotides 32,
Lissonotini 32, 87
Lissonotoidae 31
litigiosus, Proclitus 1 10
longecaudatus, Kentrotryphon 4, 65, 99
longitasis Plectiscus Proclitus 109
longithorax, Rossemia 4, 64, 96, 96*
lubricus, Anisees Pantisarthrus 4, 50, 64,
69, 104
1uridator, Oxytorus 9, 22, 30, 33, 46, 64,
71, 151, 152, 153*, 156, 156*
luridus, Pantisarthrus 4, 64, 104, 105,
105*
macurus, Proclitus 108
maderensis, Megasty1us, Mesoleptus 120
maginatus, Proclitus 110
matilei, Symplecis 51
mediato,
Helictes, ldioxenus sensu
Foerster 123
mediato, Megasty1us 119
Megalostylus 116
Megastylus (Helictes) 121
Megastylus 5, 13, 16, 18, 21, 22, 23, 24, 25,
27, 29, 30, 31, 32, 40, 42*,43, 45, 51,
55,56,57,60,62,67, 71,100,116,118
melancholica, Cylloceria 3, 25*, 28*, 30,
63, 88,89*,90,91,92,92*
melanocephalus, Poclitus 108
melanocera, Plectiscidea 6, 54, 64, 126,
132

me1anopsis, Aperileptus 5, 65, 111, 112

mendica, Plectiscidea 6, 128, 132
meridionator, Gnathochorisis 7, 140
maitus, Aperileptus 112
Mesatactodes 46, 152
Mesochorinae 27
Mesochorus 26*, 27
Mesoleptina 33, 45
Mesoleptini 33, 45
Mesoleptoidae 33
esoleptoidea 45
Mesoleptus 30, 46
mesoxantha, Plectiscidea 6, 128, 132
mesoxanthus, Plectiscus 129
mesoxanthus, Poclitus 107
Microleptes 3, 14, 15, 17, 18, 21 , 23, 26*,
27, 30, 31, 33 , 34, 36, 38, 39, 40, 46,
49,54,57,60,62,67, 71 , 72,73
Microleptinae 3, 10, 11, 13, 14*, 20*, 22*,
27,30,33,36,37,38,39,40,41,44,45,
46,47,48,49,59,62,67,68*,69, 71
Microleptini 3, 13, 14, 15, 16, 17, 18, 19,
21, 22, 23, 25, 28, 37, 38, 39, 41, 45,
49,58,62, 71,72
microspi1us, Aperileptus 5, 65, 111, 112
minima, Eusterinx 8, 64, 146
minor, Microleptes 3, 78
Miomais 31, 33
Miomaoides 116
Mischoxoides 106
moerens, Plectiscidea 6, 65, 125, 132
montanum, Hemiphanes 5, 65, 114
montanus, Oxytorus 9, 151, 152, 153*,
157
monticola, Plectiscidea 6, 54, 64, 126, 133
Myriarthidea 116
Myiarthrus 32, 121
nava, Plectiscidea, Plectiscus 6, 54, 65,
127, 133
nemorensis, Plectiscidea 6, 65, 124, 133
Neoproclitus 59, 60
Neoxorides 33
nigicapus, Aperileptus 111
nigricoxus, Helictes 123
nigitus, Plectiscus 134
nigriventis, Megastylus 119
nikkoensis, Oxytorus 9, 151, 153, 157
norikuranus, Oxytorus 9, 151, 153*, 154,
157
nuptialis, Plectiscus 133
obenbergeri Gregor, Microleptes 3, 49,
54, 74, 75*, 79
oliquus, Aperileptus, Plectiscus 113
obscura, Plectiscidea 6, 65, 125, 126, 133
obscurella, Eusterinx 8, 64, 142, 143

171

obtusus, Oxytorus 9, 151, 153, 153*, 157

occupator, Cylloceria 3, 88, 90, 93
ochropus, Pantisarthrus 104, 105
oligomera, Eusterinx 8, 64, 66, 142, 143,
144*
Ophioninae 33, 34
oritator, Megastylus 5, 30, 51, 64, 69,
117,118,120
orientalis, Cylloceria 3, 21, 88, 89*, 90,93
orientalis, Microleptes 3, 17*, 79
Orthocentrinae 18, 26*, 27, 32, 3, 37,
42*,45,68*
Ortholaba 8, 15, 18, 33, 41, 60, 149
Orthopelma 26*, 44, 45,
Orthopelmatinae 27, 44
Oxytorides 33, 45
Oxytorina 33, 45
Oxytorinae 9, 10, 11, 13, 22, 23, 27, 34,
3,37,45,47,59, 71,151
Oxytorus 9, 11, 21, 22, 23, 2*, 27, 33, 34,
35, 3,45,4,47, 59, ,, 151,152,
153*
paganus, Proclitus 5, 30, 51, 52, 4, ,
9, 10, 107, 109
pallens, Proclitus 108
paiiipes, Aniseres 4, 50, 57, 4, 103, 103*
Pantisarthrus 4, 15, 17, 18, 21, 24, 25, 32,
34,40,42,42*,43,44, 57, , 2, 7,
70, 72, 100, 104
Pantomima 5, 19, 21, 24, 43, 45, , 72,
100, 114
Pantoporthus 152
paradoxa, Symplecis 7, 5, 137, 138
paradoxus, Aniseres 4, 5, , 102, 103
Paentypoma 135
parvu1a, P1ectiscidea , 64, 125, 133
pauxillus, Megasty1us 120
pectoralis, Megastylus 5, 51, 64, , 11,
117*, 119,121
penetans, Aperileptus 111
perditorius, Proclitus 108
1j, Plectiscidea 133
perfoimidatum, Hemiphanes 5, 4, 115, 11
peiculosus, Proc1itus 109
permiranda, Eusterinx 8, 66, 145
petiolatus, Plectiscus 132
Phaeogenoidae 31,
Phidias 31, 32, 33, 149
Phosphoriana 7, 18, 42*, 43, , 102, 139
Phosphorus 139
Phytodietus 30
pilicomis, Helictes 122
pimplaius, Allomacrus 94
Pimplinae 19, 27, 29, 32, 33, 37, 42*, 44,
45,8*

172

placidus, Aperileptus 111

plagiatus, Aperi1eptus 5, 5, 111, 112
Plectiscidea , 10, 15, 17, 19, 21, 24, 25,
27, 34, 3,42*,43,44, 53, 54, 55,57,
, 2, 67, 9, 70, 101, 102, 124, 129,
129*
Plectiscides 31, 33
Plectiscina 31, 32
Plectiscinae 32
Plectiscini 33
Plectiscini auct. 37
Plectiscoidae 30, 31
Plectiscus auct. 30, 31, 32, 3, 10, 110,
124, 135
Plectiscus 3
?pleualis, Megastylus 120
Poemenia 33
Poemenini 33
Polyaulon 31, 32
Polysphincta 32
Polysphinctini 19, 32, 45
posticata, Plectiscidea , 4, 127, 133
paepositus, Plectiscus 134
praetor, Proclitus 5, 30, 51, 52, 4, 9,
10, 109
proceulus, Proclitus 108
Proclitus 4, 15, 18, 19, 21, 24, 25, 27, 30,
32, 34,42*,43, 51,5,57, , 2,7,
70, 101, 10
Proeliator 8, 15, 19, 42*, 43, 44, 60, 102,
147
prognathor, Plectiscidea 7, 65, 126, 133
prominens, Entypoma 4, 5, 97, 98
tr, Atractodes 156
popinquus, ldioxenus 122
proprius, Proeliator 8, 4, 147, 148
povidus, Proc1itus 109
poximus, Plectiscus 132
psarevi, Cushmania 3, 5, 84, 84*
pseudochropus, Pantisarthrus 105
pseudoligomera, Eusterinx 8, 4, 142,
143, 144
pseudominutus, Eusteinx, Hemiteles 144
pseudoobscuella, Eusteinx 143
pumilio, Megasty1us 120
punctatus, Proc1itus 110
pungens, Plectiscus 133
pusillus, Eusterinx, Typhon, 30, 145
quaesitorius, Proclitus 107
rectangulus, Microleptes 3, 3, 73, 73*,
74, 75*,80
refractaria, Eusterinx 8, 4, 147
remotus, Bassus 98
restricta, Gnathochorisis 7, 140
retroligatus, Megastylus 120

obustator, Entypoma 4, 5, 66, 97, 98

robustum, Entypoma 4, 50, 64, 97, 98, 98*
rossemi, Aperileptus 5, 24, 25*, 65, 111,
112
Rossemia 4, 14, 15, 18, 19, 24, 40, 41, ,
86,9

rudepunctatus, Pantisarthus 4, 65, 104,

105
rudis, Proclitus 5, 5, 107, 110
?uficaudator, Proc1itus 109
ufipleuis, Megasty1us, Myriarthus 118,
120
rufus, Aperileptus 112
rugosissima, Phosphoriana, Phosphorus 7,
5, 139
ugosissimum, Phosphoriana, Entypoma
139
salisbugensis, Micoleptes 3, 4, 74, 75*, 81
Scutellato 1
secetus, Apeileptus 112
sharkeyi, Sweaterella 9
Siiakoffia 94
signaticome, Apoclima 4, 1*, 4, 99
similis, Eusterinx 8, 5, 147
sinceus, Proclitus 108
sodalis, Plectiscus 130
sodidus, Proc1itus 110
spasskii, Microleptes 3, 81
spectalis, Proclitus 110
Sphingozona 21, 42*, 43, 60, 62
sp1endidu1us, Micoleptes 3, 1*, 30, 4,
74, 75*,82
spoliat01, Aperileptus 112
spuria, Plectiscidea 7, 53, 64, 125, 133
stenofaster, Proclitus 108
Stenolas 149
Stoia 141
subalpinus, Pantisarthrus, Aniseres 105
subangu1ata, Plectiscidea 7, 65, 129, 134
subangulatus, Plectiscus 129
subcuvatus, Plectiscus 130
subdo1a, Eusterinx 8, 4,
subsignatus, Aperileptus 113
subsimilis, Plectiscidea, Ephalmato 133
subsimilis, Plectiscidea, Plectiscus 132
substantiva, Plectiscidea 7, 5, 127, 134
substiatus, Proclitus 108
subsulcatus, Poclitus 5, 51, 4, 9, 107,
110
subteres, Plectiscidea 7, 4, 125, 134
subtilis, Allomacrus 4, 4, 89*, 94, 95
subtilis, Plectiscus 130
subtiliventris, Megasty1us, Dicolus 121
suecicus, Megastylus 5, 118, 120
suerinensis, Cyiioceria, Lampronota 3, 93

suspiciosum, Entypoma 4, 4, 97, 98

40, 9
sylvestris, Cylloceria 3, 24, 30, 64, 90, 91, 93
Symphylus 31, 32
Symplecis 7, 14, 19, 21, 22, 25, 27, 28, 31,
41 , 42*, 43, 57 , , 2, 102, 137
tartarea, Eusteinx 8, 65 , 145
tenellus, Megasty1us 5, 118, 121
tener, P1ectiscidea 7, 54, 125, 134
tenuicincta, Eusterinx 8, 17, 51, 4, 14,
147, 147*
tenuicomis, Cylloceia 4, 4, 6, 89*, 90,
91,93
tenuicomis, Plectiscidea 7, 65, , 125,
134
tenuis, Ortholaba 8, 6, 149, 150*, 151
terebata, Gnathochoisis 140
tt, Plectiscidea 7, 53 , 64, 128, 134
tetraglyptus, ldioxenus 123
tiialis, Microleptes 3, 82
tipulivora, Cylloceria 49, 54
townesi, Hemiphanes 5, 63, 114, 115, 11
Trestis 8, 13, 14, 21, 31, 141, 145
trichops, Eusterinx 8, 30, 4, , 145
tricinctus, Aperi1eptus 5, 65, 111, 113
trifasciata, Eusterinx 8, 64, , 145
truculenta, Eusterinx 8, 65, 147
Tryphon 22
Tryphonidae 31
Tryphoninae 33, 8*
Tryphonini 5
tutorius, Aperi1eptus 111
unicinctus, Proclitus 110
ussuriensis, Cyl1oceria 4, 88, 89*, 90, 93
vacuus, Aperileptus 111
vagator, Plectiscidea 7, 4, 12, 134
vagato, Plectiscus 132
validus, Proclitus 108
vanus, Aperileptus 19, 50, 57, 4, , 69,
110, 113, 113*
variator, ldioxenus 122
ventosa, P1ectiscidea 7, 128, 134
ventrator, Mesoleptus 156
viduatus, Aperileptus 5, 5, 111, 113
vilis, Aperileptus 112
visitato, Proclitus 11
volcanus, Allomacrus 4, 14, 89*, 94, 9
xanthocephala, Gnathochorisis 7, 4, 139,
140
xanthoneuris, Plectiscus 132
xanthostoma, Symplecis 137
Xoridides 33
zelator, Proclitus 11
zonaria, Symplecis 137
zonatus, Proclitus 5, 30, 64, l, 107, 110

Sweateella

 p,-oclitus ... ................................................................................

Aperileptus ..................................................................... .. ........
Pantomima ...............................................................................
. emiphanes .... ..... ... ... ...... ..... ... ... .. .... .. .... ...... .... .. .... ... ... .. ... ... ...

3.
4.
5.

...........................

....................................................................................

...................................

3
10
13

Microleptinae Oxytorinae
-

............................. .. ..... .......... .......... .. .. ........... .. ... ......

13

7. Megastylus ...............................................................................
8. Helictes ................................................................................... .
9. Fetialis .....................................................................................
10. Plectiscidea ........................................................... .................
11. Dialipsis ......................................... ............. ...........................
12. Atabulus ............... ..................................................................
13. Catastenus ..............................................................................
14. Symplecis ...............................................................................
15. Phosphoriana ....................................................... ............ ......
16. Gnathochorisis ............................................................... ........
17. Eusteinx .. ... ... ... ...... ... .... ............ ...... ....... .. ... ...... ..... .... ...... .. .. .
18. Proeliator ...............................................................................
4. DIACRITINI ..................................................................... .........
1. D iacritus .. .............. ...... ...... ... ............ ............ ....... .. ............ ......
2. Ortholaba .................................................................................

..................................................

30

MICROLEPTINAE .......................

OXYTORINAE ..............................................................
1. Oxytol'US ...................................................................................

36

48
48

....

... .. ......... .... ... .

...........................................................................................................

59

................................

.................................................................................

59

.
..........................................................................................

63

,
Microleptinae ( ) .......................... .... .

67

.......................................................
MICROLEPTINAE .......................................................

1. MICROLEPTINI ........................................................................
1. Microleptes ....................................................................... .......
2. Cushmania ... .. ..... .. ........ ......... ... .. .. .. .......... .. .... .. ... ....... .. ... ... .....
3. Hyperacmus ....... ..... ................... .. .... ..... .. ........ .. ......... ... ... ... .... .
2. CYLLOCERIINI .. ..... .. .............. ............... ......... ........... ... ... ........
1. Cylloceria ..... ... .......... ......... ..... ......... ..... ...... .. .. .... ... ...... ...... .... .
2. Allomacrus ............ ......... .... ........ ........... ...... ...... ....... .. ...... ...... ..

Rossemia ..................................................................................
Entypoma ..... .... ........... ..................... ...... ... ... .. .. .. ..... .. .......... ... ..
Kentotryphon ..........................................................................
. Apoclima ............................................................ ......................
HELICTINI ................................................................................
1. Aniseres .... ........ ... ... ... ....................... .... ....... .. .... ... .. ... ...... ... .....
2. Pantisarthrus ...........................................................................

3.
4.
5.

3.

174

71
71
72
73
83
84
86
87
94
96
97
99
99
100
102
104

106
110
114
114
116
121
124
124
135
136
136
137
139
139
141
147
148
149
149
151
152

........................................................................................ 158
168

MICROLEPINAE

OXYTORINAE
(HYMENOPTERA: ICHNEUMONIDAE)


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E-mail: secet@naukaran.ru
Internet: www.naukaran.ru
- " "

199034,

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12

90