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- v. :.

. .

-;
.;
-

.
" ' . A... : .
. .
1991. . 96. .

59

-
(DECAPODA, NATANTIA, PENAEIDAE):
,

. . ,

. .

- (Funchalia
s. lato) ,
. , . ( ,
,

) , ( R a m a d a n , 1938; Paulinose, 1974)


( F u j i n o , 1975).
.
Funchalia s. lato . Solenocerinae (Bouvier, 1908),
F u n c h a l i a e (Balss, 1925), P e n a e i n a e (Balss, 1925; B u r k e n r o a d , 1936) -
. ,
( P e l a g o p e n a e u s Faxon, 1893 Funchalia Johnson, 1867), P e n a e i n a e (Grippa, 1987). , , .
, .
,
.

19741988 . (- , ; - 9
) , (
) ( 8081) . .
( ) 1 .
,
.
.
I . , . . .
II . ,
- - ,

F. villosa,
-
F. woodwardi.
, .
60

III . , . -
F. villosa, - F.
woodwardi.
IV . .
. F. villosa - ,
F. woodwardi -.
V .
,
. F. villosa -
, F.
woodwardi-.
( /'.
villosa).
I .
,

5
.
II . .
.
III . . ,
.
IV . -
;
.

Funchaliinae subfam. nova


, , , .
.
. , - . . . ; , . .
,
,
, . .
1- 2- . 6- . , .
, , . .
,
, , . , , , ,
1- . .
Funchalia Johnson, 1867.

1. . .
.

. . . ,
, .
61

Pelagopenaeus

Burkenroad, 1934. P. balboae


( F a x o n , 1893).
. . , . .
. , .
Funchalia Johnson, 1867.

FUNCHALIA

1. ; 10
2
; 10
.
3
2. 6- ,
; ; , .
F. woodwardi J o n h s o n , 1867.
6- ; ,
; .
F. danae B u r k e n r o a d , 1940.
3.
; .
F. villosa (Bouvier, 1905).

;
.
F. taaningi B u r k e n r o a d , 1940.

Pelagopenaeus
balboae (Faxon, 1893) (. 1, )
: 21.09.82, 23.50, 3025' . ., 28 18' . . ( ), 1 ?; 7.10.82, 22.30, 637 / . ., 2 5 3 / . . (
-), 1 ? ( );
013' . ., ISMS 7 . ., : 1
81 ,
1 2 94 , ; 27.10.81, 5.357.40, 756' . .,
054 / . ., 580 , 580540 , 1 <3
86 .
: 24.07.78, 19.2019.50, 707' . 89 29' . .,
, 1 95 ; 22.07.78, 709 / . ., 8 7 1 . .,
3680 , , 80 , 1
102 .
: . , 2 ?,
; 27.05.81, 1023 / . ., 8224' . ., 4100 , 306 , 1 ? () 116 .
. : , -, ,
3 0 2 . ., 1402 / . .; : ,
. . 1500 (Burkenroad, 1936; R a m a d a n , 1938; Crosnier, Fo62

. 1. F u n c h a l i i n a e :
Pelagope
us balboae (1 , 2 ) ; Funchalia
taaningi,
F. danae,
F. villosa, F. woodwardi
(1 , , 2 , 3 ) . : 1 2, 1 \ 3, 1 3, 1, 2 3
1 , 0,5

rest, 1973). , , (3020 / . . .).

. , , . 63

, . Enoploteuthidae,
( 0,5 ) 4 1 .
Funchalia
taaningi
Burkenroad, 1940 (. 1 , 5 ) ,
: 23.06.78, 0001, 0 0 0 ' . ., 5500' . ., 8030 , , 1 33 .

.
:
(2207 / . ., 4305 / . .),
( 0 6 ' 0 0 / . ., 556205 / . .); :
-, 33 16' . ., 014 / . .
9935' . . 13939 / . .),
(00402 . ., 13939'14228' . .) (Burkenroad, 1940; Crosnier, 1985; Baba et al., 1986; W a s m e r , 1989).

.

. , 19 , , 12
300100 , 85 . 2600440 ( 730440 ) (Wasmer, 1989). . Kikuchi, . Omori (1985), -
240594 ,
483678 .
Beryx splendens,
355
(Baba et al., 1986).
F. taaningi ,
.
Funchalia
danae Burkenroad, 1940 (. 1, )
: 25.02.76, 1044 / . 1305' . ., 1200
1160 , , 12 (); 3.11.76, 328' . ., 547 /
. ., Thunnus
obesus, 1 ? 1 ?; 11.06.78, 1 2 8 ' . .,
0114' . ., 600800 , 2 ? 76 64 ; 11.05.80, 20.4021.10,
800' . ., 0 7 / . ., 490510 , 1 65 ; 22.01.81, 23.3024.00,
0447' . ., 938' . ., 798785 , 1 .; 15.07.87, 0531' . 0 1
. ., 1 .; 27.01.81, 0000.30, 426,8 / . ., 913,8' . ., 807835 ,
2 & 78 74 .

.

: - . . (Crosnier, Forest, 1973). , , . , . , (-) , Th. obesus
Alepisaurus ferox ( ), . . ,
- . : 1608 / . ., 97 14' . . (Crosnier,
Forest, 1973).
Funchalia
villosa (Bouvier, 1905) (. 1, )
: 31.01.74, 38 19' . ., . .,
1200 , ?, 4 5; 2.08.81, 21.0022.15, 4033' . ., 1050'
. ., 5200 , 088 , 4 .; 29.0725.09.84,
4358 / 3753' . ., 1418 . ., 1500
3000 , 35110 , 278 ? 2 0 7 ? ; 26.1005.11.84, , 3631 . ., 293702 / . .,
25004100 , 500800 , 29 .

.

- (Burkenroad, 1936),
64

(Grippa, 1987),
--
(Crosnier, Forest, 1973; Grippa, 1987) (2409'
26 18 . . , 0625 7 07 . .: Macpherson, 1984),
(Grippa, 1987); (2952' . . , 3432' . .: Crosnier,
1985; 2002' . . , 9941' . .: W a s m e r , 1989); - . - (Dall, 1957; Kensley et al., 1987) (Wasmer, 1989).
- 1984 . ( ) - F. villosa 46 . .,
29 . . 38 . . , 14 . . , 100%, . . ,
- (4033 / . ., 1050' . .). -,
F. villosa - .
. P. Foxton (1970) , 350700 ,
450 , 50- , . . Omori (1974), F. villosa .
103
1200 . F. villosa .
3550 80% , . 50100 .
,, 100 . ,
-, .

.

3573 . (3570 ) (4473 ).


(57 ). : 66 (. 2, ) .
; (II
). ( 3 0 % ) , , .
. ( I I I I V ) , (. 2, ) .
, F. villosa
- , : - -.
150 (103
, 2 2 ) , ,
, .
, .
, , ,
P a s i p h a e i d a e (, 1986).
. , , () , ,
, ,
}

, . 0

65

. 2. F. villosa:
, , , I , II
III II , IV IIIV , 1 , 2

. , , ( ) . ,
.
. (, ).
(, ), (Carinaria sp.). ,
(- )
Carinaria
(- ) .
,

4050 , 70% . Nyctiphanes
sp. , . (. 1) .
F.

Carinaria sp.

, %

36,9
32,1
30,1
9,7
6,8
6,8
3,9
2,9
2,9

9,2
59,1
14,1

4,5

4,5

villosa

, %

1,9
1,9
1,0
1,0
1,0
1,0
2,9
16,5
103

4,5

4,1
22

Carinaria ( ) ,
60% .
, . , , .
, F. villosa , . F. villosa . Beryx splendens
Ommastrephes
bartrami.
Funchalia
woodwardi
J o h n s o n , 1867 (. 1, )
: 29.0719.09.84, 21.3513.50, 4352'3659'
. ., 2659 / 14 . ., 15003000 , 401050 , 6 74142 , 8 S IIII, 70142 ; 1975 ., 1981 ., 1984 ., 1735 . .,
4002500 ,
, 4749 .; 31.05.69, 0829" . ., 1254 / . ., 280 , 1 <?\ 24.10.85, 08 . 0056' . ., 1 92 ; 1.10.88,
4028' . ., 5302 / . ., 12575
3000 , 1 5 74 .
: 02.01.83, 4028 / . 16347 / . ., 4 , 5 9;
.
3*

67


.

.
(Gordon, Ingle, 1956) -
(Bouvier, 1908) (Relini-Orsi, Costa, 1975; Grippa, 1987). 8 . . (), (20 . .) , - (Kensley, 1977).
- (. : , , 1985)
. - (35 0 36'3539' . ., 15055'15056' . .: Kensley et al., 1987) , -- . (, , 1985) , 1500 (Miller et al., 1983), .

.

400 . , 400500 .
400500 15,4%, 85% .
, 8001300
62,5%, 0,1% (, , 1985). , , 400500 .

.
, , , . .
60 142 ,
130 . , ( 7080 ) . , ( 117 , . 3, 3 ) .

70 (. 3, ) .
: 64
( ) 89 . 6275 , 110135 .
, - ,

, , , .
11 V ( 72122 )
: 300032000 . , .
,
, - , ,
(0,5 ) ,
,
Ocythoe
tuberculata
(
.
. . ). ( 9 1 % )
68

. 3. F. woodwardi
(3)

(, ): 1984 . (503 .), 1986 . (359 .),
1982 . (84 .), - 1981 . (144 .), 1984 .
(2459 .), 1984 . (144 .),
1975 . (1090 .), 3 1975 .
(587 .), 1985 . (110 .),
(31 .)

Terebellidae
( . . )
(. 2 ) . 99,6% .
Brachiellas,
Udonellidae
( . . ) . 1060 ( 50% ). , , ,
F. woodwardi .
, , . ( 3 4 , 4 % ) , (25,3%) ( 2 5 % ) . , , ( ), ,
300500- .
F. woodwardi

, Todarodes
angolensis,
Merluccius
sis ( ) Thunnus maccoyii ( S h a n n o n , 1986).
69


F.

, %

91.0
16,2
12,6
6,3
1,8
2,7
0,9
3,6
- -

1,8
0,9
2,7
9,9
5,4

111

woodwardi

13,0


, %

99,6

6,0

32,6
6,7
42,7
7,5
18,8
42,7
0,4
3,3
1,7
1,2

0,4

34,3

1,0
0,7
25,0
25,0
1,3
0,3

0,4
239

40

34

F u n c h a l i i n a e , . F. woodwardi,
F. villosa , Pelagopenaeus
balboae, F. danae F. taaningi.
,
F. taaningi, . F. danae . F. woodwardi
F. villosa

, P. balboae
. . , .
, F. danae,
.

P. balboae ( ,
,
. .), ,

F. woodwardi,
. F. villosa
50-, a F. taaningi
100-
.
,
, F. woodwardi .

, , . , P e n a e i n a e
( P. balboae
Penaeus spp.), ,
70

S e r g e s t i d a e Benthesicyminae. , , , , . F. woodwardi (, ) ,
,
. ,
, .
,

- , - ( O p l o p h o r i d a e ) .


P e n a e i n a e , (, 1968).

. . Pasiphaea
semispinosa/l.
.
1986. 7. . ., . .
, / / . . 1985. . 2. . ., . .

/ / . . . . 1985. . 90,
. 4. . . ., 1968. a b -, H a y s h i . I., T o r i y a m . D e c a p o d c r u s t a c e a n s f r o m c o n t i n e n t a l shelf a n d slope
a r o u n d J a p a n / / J a p . Fish. Res. Cons. Assoc. Tokyo, 1986. B a l s s H. M a c r u r a der
Deutschen Tiefsee-Expedition. 2. N a t a n t i a , Teil A . / / E r g e b n . Valdivia Exped. 1925.
Vol. 20. B o u v i e r E. L. C r u s t a c e s D e c a p o d e s ( P e n e i d e s ) p r o v e n a n t des c a m p a g n e s de
r H i r o n d e l l e et de la P r i n c e s s e Alice ( 1 8 8 6 1 9 0 7 ) / / R e s u l t . . Sci. M o n a c o .
1908. Vol. 33. B u r k e n r o a d M. D. The Aristeinae, S o l e n o c e r i n a e a n d p e l a g i c Pen a e i n a e of the B i n g h a m O c e a n o g r a p h i c C o l l e c t i o n / / B u l l . B i n g h a m o c e a n o g r . Coll. 1936.
Vol. 5 (2). C r o s n i e r A. 1985. C r e v e t t e s P e n e i d e s d ' e a u p r o f o n d e recoltee d a n s l'ocean
I n d i e n lors des c a m p a g n e s Benthedi, S a f a r i I et II, M D 3 2 / R e u n i o n / / B u l l . M u s . n a t n .
Hist. nat. P a r i s , 1985.
4 ser., sect. A. N 4. C r o s n i e r A., F o r e s t J. Les crevettes
p r o f o n d e s de 1'Atlantique oriental t r o p i c a l / / F a u n e tropicale. XIX. O R S T O M . P a r i s , 1973.
D a 1 1 W. A revision on the A u s t r a l i a n species of P e n a e i n a e ( C r u s t a c e a , D e c a p o d a : Pen a e i d a e ) / / A u s t . J. M a r . F r e s h w . Res. 1957. Vol. 8 ( 2 ) . F o x t o n P. The vertical distrib u t i o n of p e l a g i c d e c a p o d s ( C r u s t a c e a , N a t a n t i a ) collected on the S o n d cruise 1965.
II. P e n a e i d e a a n d g e n e r a l d i s c u s s i o n / / J . m a r i n e biol. ass. U. K. 1970. Vol. 50. F u j i n o T. F u n c h a l i a s a g a m i e n s i s sp. nov. f r o m C e n t r a l J a p a n , w i t h discusion of the generic c h a r a c t e r s ( D e c a p o d a , N a t a n t i a , P e n a e i d a e ) / / C r u s t a c e a n a . 1975. Vol. 2 8 ( 2 ) . G o r d o n I., I n g l e R. W. O n a p e l a g i c penaeid p r a w n s , Funchalia
woodwardi
Johnson,
new to the British f a u n a / / J . m a r . biol. ass. U. K. 1956. Vol. 35. G r i p p a G. A revision
of gen. Pelagopenaeus
B u r k e n r o a d , 1934 a n d Funchalia
J o h n s o n , 1867 ( C r u s t a c e a , Decapoda, P e n a e i d e a ) / / I n v . Pesq. 1987. Vol. 51 ( S u p l . 1). K e n s l e y B. The S o u t h African M u s e u m ' s M e i r i n g N a u d e cruises. P a r t 5. C r u s t a c e a , D e c a p o d a , R e p t a n t i a a n d Nat a n t i a / / A n n . of the S o u t h Afric. M u s e u m . 1977. Vol. 7 4 ( 2 ) . K e n s l e y .,
Trant e r H. A., G r i f f i n D. J. G. D e e p - w a t e r D e c a p o d C r u s t a c e a f r o m E a s t e r n A u s t r a l i a
( P e n a e i d e a a n d C a r i d e a ) / / R e c o r d s A u s t r . M u s e u m . 1987. Vol. 39. K i k u c h i ., r i M. Vertical d i s t r i b u t i o n a n d m i g r a t i o n of oceanic s h r i m p s at t w o l o c a t i o n s off the
Pacific coast of J a p a n / / D e e p - S e a Res. 1985. Vol. 32. M a c p h e r s o n E.
Crustaceos
Decapodos del B a n c o Valdivia (Atlantica s u d o r i e n t a l ) / / R e s . Exp. Cient. 1984. Vol. 12.
M i l l e r D. G. M., A u g u s t y n L. J., H a m p t o n J. Am u n u s u a l record of the p r a w n
Funchalia
woodwardi
( C r u s t a c e a : D e c a p o d a ) / / S . Afric. J. m a r . Sci. 1983. Vol. 1. P a ul i n o s e V. T. Decapod C r u s t a c e a f r o m the I n t e r n a t i o n a l I n d i a n O c e a n E x p e d i t i o n :
the species of Funchalia
(Penaeidae)
and
their p o s t - l a r v a e / / J . N a t u r . H i s t . 1974.
Vol. 8 ( 4 ) . R a m a d a n M. M. C r u s t a c e a : P e n a e i d a e / / J o h n M u r r a y Exped. Sci. Rep.
1938. Vol. 5 ( 3 ) . R e l i n i - O r s i L., C o s t a M. R. S e g n a l a z i o n e di Funchalia
woodwardi J o h n s o n in M a r L i g u r e ( C r u s t a c e a D e c a p o d a R e n a e i d a e ) / / B o l l . M u s . e I n s t . biol.
Univ. G e n o v a . 1975. Vol. 43. S h a n n o n L. V. The t u n a s of the B e n g u e l a r e g i o n off
S o u t h e r n Africa A s y n t h e s i s / / S C R S (86)31. IKKAT, 1986. W a s m e r R. A. Supple71

i n e n t a r y d e s c r i p t i o n of Funchalia
taaningi
B u r k e n r o a d , 1940 ( C r u s t a c e a , D e c a p o d a . P e n a e i d a e ) f r o m the C e n t r a l P a c i f i c O c e a n a n d a n e w r e c o r d of F. villosa
( B o u v i e r , 1905)
f r o m the E a s t e r n I n d i a n O c e a n / / J . N a t . H i s t . 1989. Vol. 23.
,


25.10.89

FUNCHALIA SHRIMPS (DECAPODA, NATANTIA, P E N A E I D A E ) :


TAXONOMY, D I S T R I B U T I O N A N D S O M E P R O B L E M S OF BIOLOGY
R. N. Burukovsky,

L. L.

Roinensla/

Summary
A c o m p i l a t i o n on the t a x o n o m y , g e o g r a p h i c a l d i s t r i b u t i o n , v e r t i c a l c i b p o s i ' i o n . biology and f e e d i n g in a s m a l l g r o u p of p e l a g i c P e n a e i d a e s h r i m p s , b e i n g v i e w e d a s
F i m e b a l i i m i c s u b f a m i l y , is p r e s e n t e d . T h e s e s h r i m p s a r e s h o w n l l u n c p e c u l i a r
nurt u r e of m a n d i b l e s , o v a r y , f r o n t pai'is of the s t o m a c h a n d body c o v e r i n g .