9 (104) 2011
15/2
1995 .
,
,
:
( )
:
.
-
77-21121 19 2005 .
.. ,
,
,
.
.. ,
, ,
:
.. ,
,
.. ,
,
.. ,
,
( )
:
.. ,
,
( )
.. ,
,
( )
.. ,
,
( )
. .., .
5
Berberis L.
. .., ..
11
Ficus L. in vitro. . ., ..
15
Crataegus L. (Rosaceae
Juss.) .., . .
20
Sedoideae Berger (Crassulaceae Dc.) - . .., ..
29
Chaenomeles (Thunb.) Lindl . ..,
.. 35
Ribes rubrum L. . .., .. 41
Ribes L. . .. 46
. ..,
.. 50
(Fragaria Ananassa
Duch.) . .., . . 54
(Veronica
austriaca L.). .., .. 60
. .., .. 64
.
..
68
. .. 73
.
..
77
. .., .., ..
81
Iris L. .
.., .., ..
88
. .. 94
. .. 99
Psathyrostachys caespitosa (Sukaczev) Peschkova
. ..
103
.
.. 109
.
.., ..-., .., ..
114
.
.., .., .. 122
_______________________________________________________________
:
.. ,
, (
)
.. ,
, (
)
.. , ,
( )
. , ,
( . .. ,
--)
.. ,
, ( )
.. ,
, ( )
.. ,
, (
)
.. , ,
( )
.. ,
, ( )
.. ,
,
( )
.. ,
,
( )
.. ,
,
( )
.. ,
,
( )
.. ,
, ( )
- ..
E-mail: prisniy@bsu.edu.ru
29.07.2011
6084/8
Georgia, Impact
. . . 33,94
1000 .
236
. .., .. 188
Viola odorata L. .. 191
. .., .., . . 195
-.
.., .. 199
. ..,
.. 202
Amelanchier sp. .., ..,
.., .. 208
. ..,
.., .., ..
215
. ..,
.., .., ..,
..
218
-. .., .., .. 225
,
. .., .., ..,
.., .. 233
Ribes L. ..,
.., .., .., ..,
.., ..
238
, . .., ., .., .., ..
249
. .., ..
257
Crataegus L.
. .., .., ..
265
-.
.., .., .., ..,
.., .., .., ..,
..
270
. .., ..,
.., .. 276
81466
285
-
-
: 308015, . , . , 85
288
, 2011
9 (104) 2011
Issue 15/2
SCIENTIFIC PEER-REVIEWED JOURNAL
Founded in 1995
The Journal is included into the list of the
leading peer-reviewed journals and publications coming out in the Russian Federation
that are recommended for publishing key results of the theses for Doktor and Kandidat
degree-seekers.
Founder:
Federal state autonomous educational
establishment of higher
professional education
Belgorod National Research University
Publisher:
Belgorod National Research University.
Belgorod National Research University
Press
The journal is registered in Federal service
of control over law compliance in the
sphere
of mass media and protection of cultural
heritage
Certificate of registration of mass media
77-21121 May 19, 2005.
EDITORIAL BOARD OF JOURNAL
Editor-in-chief
L.J. Djatchenko,
Rector of Belgorod National Research
University, doctor of sociological sciences,
Professor
Deputy editor-in-chief
V.V. Sokorev,
Vice-rector for scientific research
of Belgorod National Research University,
candidate of pedagogical sciences, associate
professor
Assistant Editors
V.M. Moskovkin,
Doctor of geographical sciences, Professor
of world economy department Belgorod
National Research University
E.N. Krolevetskaya,
Candidate of pedagogical sciences,
associate professor of Pedagogics
department of Belgorod National Research
University
EDITORIAL BOARD OF JOURNAL
SERIES
Chairman of editorial series
L.J. Djatchenko,
Rector of Belgorod National Research
University, doctor of sociological sciences,
Professor
Chief editor:
A.V. Prisnyi,
Doctor of biological sciences, professor
(Belgorod National Research University)
Deputies of chief editor:
O.E. Lebedeva,
Doctor of chemical sciences, professor
(Belgorod National Research University)
A.G. Kornilov,
Doctor of geographical sciences, professor
CONTENTS
BIOLOGY
Phyllophages insects and diseases of the trees and shrubs in the collection of the
Dendrological Garden Northern Research Institute of Forestry. zhov .N.,
Demidova N.A. 5
Productivity of select form of Berberis L. at introduction in the Botanical garden
Belgorod National Research University. Zhilenko V.Yu.,
Sorokopudov V.N. 11
Biological features of reproduction of some representatives of sort ficus L. in culture in
vitro. Mironov J.V., Sorokopudova O.A. 15
Adaptive features of the surface of sheet at kinds of sort Crataegus L. (Rosaceae
Juss.). Rezanova .., Baksutov S. . 20
Estimation of perspectivity species of subfamily Sedoideae Berger (Crassulaceae Dc.)
in the southwest of Russia. Orlova O.N., Sorokopudova O.A. 29
Biological features of some algal genus Chaenomeles (Thunb.) Lindl under
CHTSR. Navalneva I.A., Sorokopudov V.N. 35
Seed efficiency some grades Ribes rubrum L. in the condition of the Belgorod
region. Tohtar L.A., Sorokopudov V.N. 41
On the issue of specific idenfication of genus Ribes L. in Yakutia.
Korobkova T.S. 46
Mitotic polyploidy of asiatic hybrids of lilies. Labunskaya N.A.,
Sorokopudova O.A. 50
Introduction of pink flowering garden strawberry (Fragaria Ananassa duch.)
in Western Siberia. Kuznetsova L.L., Baturin S.O.
54
Study of anatomical structure of herb of Veronica austriaca L.
Bubenchicova V.N., Kondratova Y.A. 60
Evaluation of effectiveness of open pollination of honeysuckle promising variety
samples. Belosohov F.G., Belosohova O.A. 64
Apricot selection on quality of fruits in the conditions of the Belgorod region.
Gavrjushenko E.V. 68
Adaptive ability of grades of the raspberry on Kamchatka. Petrusha E.N. 73
Introduction of the gooseberry for improvement of grades of the Kamchatka edge.
Petrusha E.N.
77
Study of anatomy of herb tragopogon orientalis. Bubenchikova V.N., Trembalya Ya.S., Prokhorova S.A.
81
Rreintroduction of some species of Iris L. in the Botanical garden of Belgorod
State University. Shevchenko I.V., Chernyavskih V.I.,
Sorokopudova O.A. 88
Studying of a gain of corms and other parameters of vegetative efficiency of a gladiolus cultivars. Kuzichev O.B. 94
Symbiotic activity of root nodule bacteria of alfalfa varieties and breeding lines
under conditions of central Yakutia. Atlasova L.G. 99
The bunch formation in Psathyrostachys caespitosa (Sukaczev) Peschkova in the
first years of life under conditions of Central Yakutia. Skobeleva A.A. 103
Studying of key parameters of vegetative growth of gladiolus cultivars.
Kuzichev O. B. 109
Flora of woods of some foothill areas of East Caucasus. Taisumov M.A.,
Astamirova M.A.-M., Israpilova S.A., Gadaeva T.Z. 114
Morphological features of the yakut populations of wild strawberry east.
Vasileva E.P., Belevtcova V.I., Sorokopudov V.N. 122
_______________________________________________________________
(Belgorod National Research University)
Responsible secretary:
Yu.N. Kurkina,
Candidate of agricultural sciences, associate professor (Belgorod National Research University)
Members of editorial board:
L.N. Balyatinskaya, Doctor
of chemical sciences, professor
(Belgorod National Research University)
I.V. Batlutskaya, Doctor
of biological sciences,
professor (Belgorod National Research
University)
A.I. Vezentsev, Doctor
of technical sciences, professor
(Belgorod National Research University)
Rudiger Wittig,Doctor, professor
(I.V. Gete University, Frankfurt-on-Mine)
A.F. Kolchanov, Candidate
of biological sciences, professor
(Belgorod National Research University)
F.N. Lisetskyi, Doctor
of geographical sciences, professor
(Belgorod National Research University)
S.V. Lukin, Doctor of geographical
sciences, professor
(Belgorod National Research University)
A.N. Petin, Doctor of geographical
sciences, professor (Belgorod State
University)
S.V. Sergeev, Doctor of technical sciences,
professor (Belgorod National Research
University)
V.N. Sorokopudov, Doctor
of agricultural sciences, professor
(Belgorod National Research University)
V.K. Tokhtar, Doctor of biological
sciences, senior scientific employee
(Belgorod National Research University)
M.Z. Fiodorova, Doctor
of biological sciences,
professor (Belgorod National Research
University)
Yu.G. Chendev, Doctor
of geographical sciences, professor
(Belgorod National Research University)
V.S. Shilova, Doctor of pedagogical
sciences, professor (Belgorod National
Research University)
______________________________________________________________
632:631.963 (470.11)
.. 1
.. 2
1
,
163000, ,
. , 23.
e-mail: olegezhik@gmail.com
2
, 163062,
, . , 13.
e-mail: forestry@ptl-arh.ru
-
(. ) 2004 2010 . , .
. - .
: , -, , .
- ( )
29 1960 . . (64 0 33/ ..
390 40/ ..). 1969 .
646
595 73 29 . 1100
( 6400 ) (, ,
, , ).
, (Lonicera) 51
, (Rosa) 48 (Crataegus) 37 [1].
. -
.
. , .
-
,
[2, 3, 4, 5].
2004 2010 .
.
_______________________________________________________________
- . , ,
. (Euonymus L.),
(Sambucus L.), (Cornus L.), (Physocarpus Maxim.),
(Sorbaria A. Br.), (Spiraea L.), (Fraxinus L.),
(Lonicera involucrate Sprehg.), (Viburnum lantana L.),
(Picea pungens Engelm.) (P. abies),
(Dasiphora fruticosa (L.) Rydb.), (Corylus avellana L.),
(Elaeagnus argentea Pursh.), (inus pumila (Pall.), (Thuja occidentalis L.), (Padus maackii Kom), (Chaenomeles Lindl.) .
(. 1). ,
( 2004 2010 .). . () () ( ) [6] .
1
-
*
*
*
*
*
*
*
*
*
*
*
*
*
(Microsphaera palczewski Jacz.)
(Puccinia graminis Pers.)
(Cemiostoma scitella L.),
(Aporia crataegi L.), - (Phyllosticta
Michalowskoensis Elenk. et Ohl.)
- (Colopha compressa Koch.),
(Eriophyes brevipunctatus Nal.)
- (Colopha compressa Koch.); -
(Eriosoma ulmi L.),
(Microsphaera alphitoides Griff. et Maubl.)
(Phytagromyza xylostei R. D.), -
(Phopalosiphum lonicerae Sieb.)
(Hyadaphis tataricae Aizend)
- (Daseocheta (Diphtera) alpium Osbek.)
(Podosphaera schlechtendalii Lev., Uncinula salicis D. C.),
(Phyllodecta vitellinae L. Ph.
vulgatissima L)
(Aphis viburnorum F.)
(Rhytisma punctatum (Pers.) Fr.)
(Gloesporium tiliae Oud.),
(Eriophyes tiliae var. Liosoma Nal.)
(Coleophora laricella Hb.), (Melampsora laricis-populina Kleb., Melampsoridium betulae (Schum.) Arth.)
(Phyllotoma vagans Cl.), (Eriophyes laevis Inangulis Nal.),
() (Chionaspis salicis L.)
- (Megachile centuncularis L.), (Blennocampa pusilla Kl.), (Phodites rosarium Gir.)
(Eriophyes goniothorax sorbeus Nal.), (Gumnosporangium orbiculatum (Desm.) Thum.),
(Dentatus sorbi Kait)
______________________________________________________________
*
*
*
*
*
: * ;
** [7].
, . (Berberis): (B. vulgaris f. Atropurpurea Rgl., (B. integerrima Bunge,
(B. circumserrata Schneid.), (B. aristata DC.),
(B. Poiretii C.K. Schneid., (B. dasystachya Maxim.), -
(B. Francisci-ferdinandii Schneid.) (B. laxiflora Schrad.) . , .
(()) [6] (
), ( ). .
(Sorbus discolor (Maxim) Hedl.) (S. aucuparia L.) . (S. sambucifolia (Cham. et Schltd.)
m.Roem.). .
:
(Venturia inaequalis (Cooke) Wint.), -
(Phyllosticta Michalowskoensis Elenk. et Ohl.,
(Rhytisma punctatum (Pers.) Fr.),
(Gloesporium tiliae Oud.).
Agromyzidae (
), Coleophoridae ( ), Gracilariidae (-),
Tenthredinidae (), Eriophyidae ( ), Lyonetiidae ( ), Chrysomelidae (), Megachilidae (-), , ,
.
-
.
(Pines sibiricus Cholodk.), .
, -,
. .
. (.2).
(()) [6] -
_______________________________________________________________
, , .
, .
2
Athelia fibulata M.P. Christ.
Basidioradulum crustosum (Pers.) Zmitr.,
Malysheva et Spirin
Basidioradulum radula** (Fr.) Nobles
Bjerkandera adusta** (Willd.: Fr.) P.
Karst.
Byssomerulius corium(Fr.) Parmasto
Cerrena unicolor** (Bull.) Murrill
Chondrostereum purpureum** (Pers.)
Pouzar
Coniophora arida (Fr.) P. Karst.
Cylindrobasidium leave (Pers.) Chamuris
Daedaleopsis confragosa (Bolton) J.
Schrt.
Exidia glandulosa (Bull.) Fr.
Inonotus radiatus (Sowerby) P. Karst.
Hyphoderma cremeoalbum (Hhn. et
Litsch.) Jlich
Hyphodontia sambuci (Pers.) J. Erikss
Hypochnicium bombycinum (Sommerf.) J.
Erikss.
Nectria cinnabarina (Tode) Fr.
Peniophora incarnata (Pers.) P. Karst.
Merulius tremellosus Schrad.
P. nuda (Fr.) Bres.
Phellinus conchatus (Pers.) Qul.
Ph igniarius (L.) Qul.
Pycnoporellus fulgens (Fr.) Donk
Pseudochaete tabacina (Sowerby) T.
Wagner et M. Fisch.
Radulomyces confluens (Fr.) M.P. Christ
Trametes hirsuta** (Wulfen) Lloyd
Vuilleminia comedens (Nees) Maire
*
*, *
*, *, *
*
.
*, *, *
*
, ,
*
, *
*
*
, *, *
, *
, , *,
,
*
*
*, *, *, *, ,
*
*
*, *
*
*
*
: * ;
** , .
, . ( ) (. 3), . (37,5%) (45,8%) . ,
, , 100%.
59%, 32%,
9%. ( (())) -
______________________________________________________________
. (77,4%),
, [8].
3
- , %
PA
4.8
0
0
0
0
9.5
4.8
9.5
H
0
0
9.5
9.5
MR
0
0
76.2
76.2
4.8
0
95.2
100
. : PA , H , MR , .
: Chondrostereum purpureum, Cylindrobasidium leave, Exidia glandulosa,
Hypochnicium bombycinu, Nectria cinnabarina, Peniophora incarnata, Phellinus conchatus. , .
-
, . : , , . , . ,
. (()) [6] , ,
. , . , , ( ,
); -, (
(())); -, ( ).
, ,
.
: , , ().
.
1. - /.. , .. , .. , .. ; . : , 2006. 20 .
2. .. . ; . 2:
, , , , ,
( ), ( -
10
_______________________________________________________________
), ( ), , ,
, . .: , 1998. 391 .
3. .., - .. . : , 1951. 580 .
4. .. ,
. .: , 1984. 472 .
5. .., .. . I
. Melampsoraceae . Pucciniaceae. : , 1975.
336 .
6. .., .. - // , 2010. 2. .50-54.
7. / .. ,
1990. 86 .
8. .. // . 1997. . 31, . 1. . 1927.
of Ecology Problems in
the North Ural Branch of the
Russian Academy of Sciences,
163000, Arkhangelsk, Severnaya Dvina Embankment, 23.
e-mail: olegezhik@gmail.com
2
e-mail: forestry@ptl-arh.ru
The article lists the main pests and diseases species of introduced trees
and shrubs in the Dendrological Garden Northern Research Institute of Forestry (Arkhangelsk City) for the period of observation from 2004 to
2010.Resistant species, timing of pests and diseases, peculiarities of their prevalent in the collection are identified. That will permit to develop measures
for protection and to create sustainable and highly decorative greenery in cities and settlements of Arkhangelsk Region. The list of wood-destroying fungi
is presented and coeno-environmental analysis is conducted.
Key words: introduced tree and shrub species, phyllophages insects,
diseases, wood-destroying fungi.
11
______________________________________________________________
631.55:634.746:378.4 (470.325)095
BERBERIS L.
..
..
, . ,
. , 85
e-mail: zhilenko@bsu.edu.ru,
sorokopudov@bsu.edu.ru
Berberis L.,
: B. koreana 1-1, B. vulgaris 4-1, B. heteropoda
6-1, B. vulgaris f atropurpurea 8-8, B. cretica 9-1
B. dielsiana 10-7.
: , , ,
.
: B. koreana: 1-1,
1-2, 1-3, 1-4, 1-5, 1-6, 1-7, 1-8, 1-9; B.vulgaris: 4-1, 4-2, 4-3, 4-4, 4-5, 4-6, 4-7, 4-8, 4-9; B.
heteropoda: 6-1, 6-2; B.vulgaris f atropurpurea: 8-1, 8-2, 8-3, 8-4, 8-5, 8-6,8-7, 8-8 8-9;
B. cretica: 9-1, 9-2, 9-3; B. dielsiana: 10-1, 10-2, 10-3, 10-4, 10-5, 10-6, 10-7, 10-8 109. 3 (2008-2010.). , (1999) [4].
2008-2010 ., .
, (. 1).
1
Berberis L.
1-1
1-2
2008
2009
2010
2008
2009
2010
1,5
1,3
0,9
0,8
0,7
0,6
3,3
2,8
1,9
1,8
1,5
1,3
B. koreana
0,160,02
0,170,02
0,210,03
0,200,20
0,190,02
0,170,01
0,14-0,18
0,12-0,22
0,14-0,27
0,15-0,20
0,12-0,24
0,13-0,19
0,27
0,29
0,28
0,22
0,25
0,26
, V, %
9,3%
17,6%
26,6%
11,1%
1,5%
20,3%
12
_______________________________________________________________
3
1-3
1-4
1-5
1-6
1-7
1-8
1-9
10
4-1
11
4-2
12
4-3
13
4-4
14
4-5
15
4-6
17
4-7
18
4-8
19
4-9
20
6-1
21
6-2
22
8-1
23
8-2
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
0,5
0,6
0,5
0,4
0,5
0,4
0,3
0,3
0,2
0,3
0,2
0,1
0,2
0,2
0,1
0,2
0,1
0,1
0,2
0,1
0,1
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
3,5
3,2
3,1
2,5
2,4
2,2
2,6
2,4
2,3
2,1
2,0
1,9
2,9
2,8
2,7
1,9
1,6
1,5
2,0
1,7
1,6
2,4
2,3
2,2
2008
2009
2010
1,9
1,7
1,6
2008
2009
2010
2008
2009
2010
1,9
2,0
3,1
1,7
1,8
2,9
2008
2009
2010
2008
2009
2010
1,7
1,5
1,6
2,0
1,7
1,6
1,1
1,3
1,1
0,8
1,1
0,8
0,6
0,5
0,4
0,6
0,5
0,2
0,5
0,3
0,2
0,5
0,3
0,2
0,4
0,3
0,2
0,160,02
0,14-0,22
0,170,01
0,15-0,20
0,140,03
0,10-0,20
0,170,02
0,13-0,25
0,170,02
0,13-0,20
0,170,01
0,2-0,18
0,070,03
0,04-0,20
0,190,01
0,15-0,22
0,180,01
0,15-0,21
0,150,01
0,14-0,18
0,160,01
0,12-0,19
0,140,01
0,11-0,16
0,150,01
0,13-0,18
0,180,03
0,08-0,23
0,160,01
0,14-0,17
0,160,01
0,12-0,18
0,20,01
0,17-0,20
0,20,02
0,17-0,23
0,180,05
0,16-0,18
0,220,02
0,18-0,22
0,170,03
0,14-0,22
B.vulgaris
7,7
0,180,05
0,12-0,27
7,1
0,260,03
0,22-0,32
6,8
0,260,04
0,19-0,30
5,5
0,200,03
0,12-0,20
5,3
0,100,02
0,08-0,13
4,8
0,110,01
0,09-0,13
5,7
0,180,05
0,12-0,19
5,3
0,150,01
0,14-0,15
5,1
0,130,01
0,12-0,14
4,6
0,210,03
0,14-0,23
4,4
0,210,02
0,15-0,27
4,2
0,290,03
0,25-0,29
6,4
0,200,05
0,18-0,26
6,2
0,240,03
0,18-0,27
5,9
0,130,01
0,11-0,14
4,2
0,140,01
0,13-0,15
3,5
0,190,01
0,17-0,21
3,3
0,210,01
0,19-0,22
4,4
0,160,01
0,15-0,17
3,7
0,140,01
0,14-0,15
3,5
0,150,01
0,14-0,15
5,3
0,210,01
0,20-0,22
5,1
0,210,01
0,20-0,23
4,8
0,250,02
0,22-0,28
B.vulgaris
4,2
0,150
0,15-0,18
3,7
0,150,01
0,14-0,15
3,5
0,160,01
0,14-0,16
B. heteropoda
4,2
0,040,08
0,04-0,05
4,4
0,120
0,07-0,10
6,8
0,090,01
0,08-0,16
3,7
0,050,01
0,02-0,07
3,9
0,090,01
0,11-0,12
4,6
0,120,02
0,09-0,11
B.vulgaris f atropurpurea
3,7
0,070,01
0,06-0,07
3,3
0,050,01
0,04-0,05
3,5
0,050,01
0,04-0,05
4,4
0,20,03
0,12-0,27
3,7
0,10,01
0,13-0,16
3,5
0,290,03
0,25-0,30
0,24
0,22
0,21
0,26
0,21
0,19
0,21
0,28
0,22
0,23
0,25
0,17
0,31
0,27
0,18
0,24
0,23
0,25
0,19
0,28
0,23
18,8%
10,5%
13,0%
13,3%
10,0%
6,7%
16,0%
14,2%
13,6%
10,6%
12,5%
16,6%
13,5%
33,3%
12,8%
13,3%
16,0%
15,0%
9,2%
16,8%
28,8%
0,32
0,33
0,35
0,27
0,17
0,15
0,27
0,16
0,18
0,27
0,28
0,30
0,27
0,28
0,16
0,17
0,24
0,29
0,18
0,20
0,19
0,23
0,24
0,29
44,0%
19,2%
23,1%
34,0%
26,0%
18,1%
44,4%
4,8%
7,6%
27,5%
23,8%
15,5%
28,0%
22,9%
12,5%
7,1%
10,7%
8,4%
6,3%
4,0%
2,4%
4,7%
7,1%
12%
0,19
0,20
0,18
0%
4,0%
4,0%
0,06
0,19
0,18
0,08
0,13
0,14
8,0%
16,6%
29,1%
15,0%
0%
8,5%
0,08
0,07
0,09
0,28
0,20
0,34
8,6%
15,5%
12,6%
30%
14,3%
15,2%
13
______________________________________________________________
24.
8-3
25
8-4
26
8-5
27
8-6
28
8-7
29
8-8
30
8-9
31
9-1
32
9-2
33
9-3
34
10-1
35
10-2
36
10-3
37
10-4
38
10-5
37
10-6
38
10-7
39
10-8
40
10-9
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
1,5
1,4
1,3
1,7
1,5
1,4
1,4
1,2
1,2
1,1
1,0
0,9
1,3
1,2
1,1
2,2
2,1
2,0
1,4
1,2
1,1
3,3
3,1
2,8
3,6
3,3
3,1
2,9
2,7
2,5
2,4
2,2
1,9
2,8
2,6
2,4
4,8
4,6
4,4
3,1
2,7
2,4
2008
2009
2010
2008
2009
2010
2008
2009
2010
0,5
0,4
0,3
0,2
0,1
0,09
0,1
0,06
0,05
1,1
0,8
0,6
0,4
0,2
0,2
0,2
0,1
0,1
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2008
2009
2010
2,5
2,3
2,1
2,0
1,9
1,8
2,1
2,0
1,9
2,3
2,1
2,0
2,4
2,2
2,1
3,0
2,8
2,7
3,1
3,0
2,9
2,4
2,3
2,1
2,6
2,5
2,3
5,5
5,1
4,6
4,4
4,2
3,9
4,6
4,4
4,2
5,1
4,6
4,4
5,3
4,8
4,6
6,6
6,2
5,9
6,8
6,6
6,4
5,3
5,1
4,6
5,7
5,5
5,1
0,180,02
0,16-0,24
0,190,04
0,12-0,27
0,140,02
0,12-0,18
0,190
0,19-0,24
0,180,05
0,13-0,27
0,130,02
0,10-0,15
0,050,01
0,04-0,05
0,090,03
0,06-0,11
0,080,02
0,06-0,11
0,030,01
0,01-0,05
0,050
0,05-0,06
0,030,01
0,01-0,05
0,080,01
0,07-0,08
0,070,01
0,06-0,08
0,080,01
0,07-0,08
0,160,01
0,13-0,19
0,170,03
0,12-0,27
0,300,03
0,20-0,30
0,080,01
0,06-0,08
0,060,01
0,05-0,06
0,060,06
0,05-0,07
B. cretica
0,120,01
0,09-0,14
0,140,01
0,12-0,17
0,200,02
0,13-0,21
0,120,02
0,03-0,17
0,130,01
0,12-0,14
0,120,01
0,10-0,20
0,120,02
0,09-0,10
0,130,01
0,12-0,13
0,140,02
0,08-0,15
B. dielsiana
0,080,01
0,05-0,13
0,090,02
0,05-0,18
0,140,02
0,08-0,18
0,190,01
0,17-0,21
0,140,03
0,05-0,16
0,120,01
0,10-0,15
0,160,01
0,13-0,18
0,200
0,20-0,21
0,090,02
0,06-0,13
0,160,01
0,14-0,18
0,150,01
0,12-0,18
0,160,01
0,14-0,18
0,130,01
0,11-0,14
0,120,01
0,11-0,14
0,160,01
0,14-0,18
0,090,01
0,08-0,10
0,150,01
0,15-0,16
0,160,01
0,14-0,18
0,200
0,20-0,21
0,170,01
0,16-0,18
0,170,01
0,15-0,18
0,160,01
0,13-0,18
0,150,01
0,12-0,18
0,150,01
0,12-0,18
0,140,01
0,13-0,14
0,120,02
0,09-0,14
0,120,01
0,09-0,14
0,19
0,30
0,19
0,25
0,28
0,20
0,07
0,12
0,13
0,06
0,07
0,08
0,09
0,10
0,09
0,20
0,30
0,31
0,09
0,07
0,09
9,4%
39,4%
0,9%
0%
23,3%
30,0%
12,4%
31,2%
32,9%
36,5%
0%
36,7%
9,3%
7,1%
8,8%
15,9%
35,2%
15,5%
20,1%
12,7%
16,6%
0,16
0,18
0,22
0,18
0,15
0,21
0,14
0,15
0,20
15,8%
13,9%
23,6%
24,5%
7,7%
16,6%
21,6%
4,4%
38,5%
0,15
0,18
0,19
0,23
0,20
0,20
0,19
0,22
0,14
0,18
0,19
0,18
0,15
0,14
0,19
0,11
0,17
0,19
0,22
0,19
0,20
0,19
0,18
0,19
0,17
0,16
0,17
36,6%
56,1%
34,5%
12,1%
18,6%
16,6%
16,0%
0%
40,1%
9,1%
14%
10,0%
11,9%
10,6%
9,1%
11,1%
4,6%
9,1%
0%
5,9%
6,9%
11,9%
0,3%
13,3%
7,0%
21,6%
18,8%
0,1 / (0,3 /)
1-7, 1-8 1-9 (B. koreana) 3,5 / (7,7 /) 4-1 (B.vulgaris).
, 3,5 / 2008-2010 .,
14
_______________________________________________________________
: 1-1 (B. koreana) 1,5 / (3,3 /); 4-1 (B.vulgaris) 3,5 / (7,7 /);
6-1 (B. heteropoda) 3,1 / (6,8 /); 8-8 (B.vulgaris f atropurpurea) 2,2 /
(4,8 /); 9-1 (B. cretica) 0,5 / (1,1 /); 10-7 (B. dielsiana) 3,1 /
(6,8 /). ,
- , , B. heteropoda. B. heteropoda. 6-1. 6-2 2010 , 2008 (. . 1).
2008-2010 . 0,03 (B.vulgaris f atropurpurea, 8-3) 0,26 (B.vulgaris, 4-1)
0,30 (B.vulgaris f atropurpurea, 8-8).
B.vulgaris ( 4-1) B.vulgaris f atropurpurea( 8-8). B.vulgaris ( 4-1) 0,35 ,
B.vulgaris f atropurpurea ( 8-8) 0,31 . : 1-1 (B. koreana)-
0,21 , 0,29 ; B. heteropoda: 6-1, 0,14 , 0,19 ; 9-1 (B. cretica)- 0,20 0,21 ; 10-7
(B. dielsiana) 0,20 0,21 .
Among the populations of Berberis L., are promising form of: B. koreana form 1-1, B. vulgaris form 4-1, B. heteropoda form 6-1, B. vulgaris f atropurpurea form 8-8, B. cretica form 9-1 and B. dielsiana form of 10-7.
form.
Key words: types of barberry, the mass of the fruit, productivity, select
15
______________________________________________________________
635.914.635.918
FICUS L. IN VITRO
..
..
, . ,
. , 85.
e-mail: sorokopudov@bsu.edu.ru
Ficus L.
. .
.
, .
: Ficus L., ,
.
,
in vitro ,
- . ,
,
(F. arica), .
, ..
. , , (, 1997).
. Ficus ; .
. Ficus.
(, 1999).
- , 30 /,
7 /, 0,1 /. , 2-, , , ,
?. 5,7 6,0.
in vitro
Ficus: Ficus elastica Roxb., Ficus Benjaminii L., Ficus lirata Warb., Ficus binnendijkii Mig., Ficus deltoidea Jack.
-,
. , ,
(1 2- ), -
16
_______________________________________________________________
( 5 10 ). (, 1986; ,
, 1983; Jona, 1982).
F. benjamina 2- , F. elastica ,
F. lyrata .
Jona, 1982; Jona, Gribaudo, 1987, 1991.
: 8 , 10-12 . , 12 . .
. 5-7 , , , .
:
F. benjamina ( Daniella, Starlight, Gold Princess, Monique Exotica), F. elastic (
Melany, Rubra, Brazil), F. lyrata ( Bambino).
.
.
F. lyrata
( )
. 3 , . -
(), (1/2 ) - . (.1):
1
F. lyrata
( 1,0 /)
66+1,3
24,8+4,9
5,1+1,8
2,8+0,9
18,0+3,9
13,9+3,0
F. lyrata . :
1. 2- (8,05 /),
(5,4 /), (2,5 /), (5,16 /), 10- 100 , () ().
(.10). , 100- , 10-.
,
.
(. 2).
17
______________________________________________________________
2
, ,
F. Lyrata
1
2
3
4
5
1:100 +
1:100
1:10 +
1:10
1:10 +
7,33+1,10
9,04+1,00
6,33+1,56
6,17+0,91
7,14+1,96
2.
. (. 3).
3
F. Lyrata
, .
1
2
3
4
5
1,5
0,8
0,5
2
12,0+2,0
14,0+4,0
2,0+0,5
3,0+1,0
34,0+7,0
5,0+1,8
3,5+1,1
7,0+1,2
23,0+6,9
8,3+0,8
1,0+0,1
3,0+0,5
1,5+0,1
, F.lyrata
2 ( ), (, 2003; , 2004).
(0,05 /) . . , , ,
F. lyrata 2 (. 3).
, F. lyrata 0,5 /
0,05 / .
F. elastic : Melany, Rubra,
Brazil. I (st), (. 4).
4
F. elastic
( )
Melany
Rubra
Brazil
I (st)
7,2+1,2
5,3+0,7
4,8+0,5
II
III
IV
7,4+1,1
8,5+1,3
10,5+1,5
5,8+0,8
8,1+1,0
8,5+1,1
5,5+0,5
5,8+0,7
6,5+0,9
V
8,3+1,2
6,4+0,9
6,8+1,0
F. elastic F. lyrata ,
F. lyrata (
18
_______________________________________________________________
34,5+7,2). F. elastic Melany ( 10,5+1,5).
. F. lyrata.
( ) .
(. 5).
5
F. lyrata
,
/
0,5
1,0
0,5
1,0
0,5
1,0
,
%
52,5+15,1
88,1+11,7
89,2+10,6
95,8+5,4
88,1+11,7
83,3+15,2
68,4+15,9
1,4+0,2
6,9+1,3
7,2+1,5
4,6+0,8
3,8+0,6
2,9+0,5
2,3+0,5
13,4+2,5
30,2+5,1
24,6+3,8
17,6+2,9
11,8+2,1
10,7+1,9
6,7+1,3
. in vitro
(Sphagnum)
( ., 2000).
, 1:1:1. 5 7 . .
, .. .
Ficus .
, .
1. Ficus . ( )
.
2. in vitro
Ficus , (1 2
); F. lyrata
0,5 / 0,05 / ;
0,5 /, 1,0 /;
.
3.
, .
19
______________________________________________________________
1. .. . .: . , 1986. 179 .
2. .. . . ., 1997. 20 .
3. .., .. . ., 1983. 97 .
4. .., .., .. - // , . . 179. ., 2000. . 136-143.
5. Jona R. In vitro propagation of Fig through Shoot Tip culture // Hort Science 17 (1):
86-87. 1982.
6. Jona R., Gribaudo J. Adventitious bud formation from leaf explants of Ficus lyrata //
Hort Science. 1987. - Vol 22, 4. - P. 651-653.
7. Jona R., Gribaudo J. Ficus spp. // Biotechnology in agriculture and foresty. Vol. 16, 1991.
P. 79-93.
20
_______________________________________________________________
581.821
CRATAEGUS L. (ROSACEAE JUSS.)
..
. .
, . ,
. , 85.
e-mail: Rezanova@bsu.edu.ru,
baks922@rambler.ru,
sorokopudov@bsu.edu.ru
- 20
Crataegus L. . , , ,
. Crataegus L. .
.
: Crataegus, , ,
, .
, , . , , , [12, 14]. . , ,
[1].
. 30 60
. ., , .
,
[4, 2]. , , .
, [13].
, .
. .
Crataegus L.,
, 2001 : Crataegus flabellata
(Bosc ex Spach) K.Koch, C. douglsii LINDL., C. punctata Jacq., C. macracantha Lodd. ex
Loudon, C. holmesiana Ashe., C. ellwangeriana Sarg., C. maximowiczii C.K.Schneid.,
C. prunus Spiendens, C. sanguinea Pall., C. chlorosarca Maxim., C. monogyna Jacq.,
C. submollis Sarg., C. turkestanica Pojark., C. pentagyna Waldst. & Kit. ex Willd., C. arnoldiana Arnold Hawthorn, C. Crus-Galli L., C. rotundifolia Moench., C. pringlej, C. canadensis Sarg, C. canadensis Sarg.
C. monogyna Jacq., - , , , .
21
______________________________________________________________
.
, . (7- ) , . : , , (1:1:1) [11]. [10].
13, -5, 10 Quanta 200 3D , -.
,
50- .
, , () [7], . ,
.
. . (1984), Microsoft Office.
, , ,
[6]. , ,
. (, ), , , 2, [8].
.
. , [12].
. .
, . ,
, .
[8]. [3] , , , , , , , (. 1). , , , , , , , ,
, , .
[3]. , , , , , , , , , .
(-0,532).
4,25-9,64%, ,
. :
, , . -.
, .
22
_______________________________________________________________
, , .
1
Crataegus
C. flabellata
.
C. douglsii
.
C. punctata
.
C. macracantha
.
C. holmesiana
.
C. canadensis
.
C. ellwangeriana
.
. densiflorum
.
C. maximowiczii
.
C. prunus
.
C. sanguinea
.
C. chlorosarca
.
C. monogyna
.
C. submollis
C. turkestanica
.
C. pentagyna
.
C. pringlej
.
C. arnoldiana
.
C. Crus-Galli
C. rotundifolia
.
885,1122,50**
1,250,016*
69,704,96
8,070,511
154,337,54
857,8913,19**
1,360,014
80,733,06
9,530,393**
125,595,12**
687,4023,85
1,280,028
56,563,43
8,330,486
125,396,33**
867,1124,55**
1,240,024
62,134,35
7,030,371
194,975,86**
664,6526,87
1,330,039
67,0220,05
7,120,429
176,9910,54
765,8026,90*
1,330,020
59,274,32
7,410,431
133,663,48**
617,6822,28
1,290,03
43,602,89
7,070,415
165,356,60
1,160,019
71,485,12**
7,660,510
153,699,38
1,180,024
54,905,83**
4,250,407**
22,522,26**
715,3625,74
1,380,025
52,694,72
7,310,512
210,1310,04**
852,3128,80**
1,300,024
60,143,72
7,110,398
233,189,81**
754,5526,09*
1,260,020
51,744,22
6,830,469
210,278,89**
649,4228,78
1,390,034
45,043,47
6,950,418
146,757,25
665,1321,03
1,330,029
47,804,70
7,170,576
161,477,43
679,2119,67
1,340,024
48,583,21
7,210,480
147,526,64
869,5431,44**
1,260,026
73,795,44
8,490,558
190,156,51*
873,7233,40**
1,280,026
56,953,57
6,590,390
185,337,23
646,0617,03
1,290,025
47,003,47
7,300,521
217,718,49**
754,1229,94*
1,240,026
56,083,74
7,480,440
164,959,12
668,0836,79
1,250,023
48,623,93
7,520,597
183,266,88
1026,7134,34**
1,440,325
97,545,27
9,640,567
156,166,88
.
945,1539,05**
1297,8261,51**
23
______________________________________________________________
, , , , , , , , ,
(. 2).
2
Crataegus
C. flabellata
C. douglsii
C. punctata
C. macracantha
C. holmesiana
C. canadensis
C. ellwangeriana
. densiflorum
C. maximowiczii
C. prunus
C. sanguinea
C. chlorosarca
C. monogyna
C. submollis
C. turkestanica
C. pentagyna
C. pringlej
C. arnoldiana
C. Crus-Galli
C. rotundifolia
S . .
1120,3865,22**
12,700,363**
878,06 57,42*
14,900,550
966,6049,27**
14,360,379**
722,1638,40
17,400,626
818,1147,70
15,660,540
327,7416,17
22,760,622**
819,8544,24
14,310,447**
489,2230,03**
21,950,237**
645,6139,35
17,020,618
550,6929,30**
20,900,714**
422,0016,52**
19,050,388**
704,8726,67
16,220,353
710,9028,54
18,340,384**
869,5540,82**
16,100,429
620,9550,38
17,400,665
524,4124,27**
18,860,457**
553,7827,89**
19,130,455**
766,6626,00
16,830,339
1150,0754,91**
13,240,317**
S . .
1132,6996,93*
12,760,647
1296,4544,51**
11,160,227**
998,1034,88
12,320,193**
1228,6643,06**
11,480,240**
641,8023,12**
15,810,332**
1035,0244,63
12,220,277**
540,1624,69*
17,900,450**
881,5943,10**
13,010,094
831,6728,15
13,460,677
776,8625,95
14,400,288
783,8423,61
15,370,343
501,6428,82**
18,220,470**
865,1446,66
14,100,445
858,3437,49
14,150,317
1129,0248,65**
11,820,281**
605,9018,28**
15,730,251**
615,4018,78**
15,410,188*
1157,3538,62**
12,750,230*
746,1922,26
14,850,174
996,1552,27
12,900,368
24
_______________________________________________________________
,
.
. C. arnoldiana
. . densiflorum
. C. rotundifolia
. C. chlorosarca
. C. canadensis
. C. macracantha
. C. submollis
. C. pringlej
. C. turkestanica
. 1. Crataegus
,
, , , , , , .
. 20 Crataegus : (. 2), (. 2), (. 2).
, , 2 .
, , ,
[15].
25
______________________________________________________________
. ,
.
. C. canadensis
. C. canadensis
. C. ellwangeriana
. 2. Crataegus
,
(. 3). ,
(.3).
. 3. C. pringlej
, , ,
(. 3).
.
, , .
.
, , , , ,
,
.
. . : , ; ; . , [5].
26
_______________________________________________________________
3
,
Crataegus
C. flabellata
C. douglsii
C. punctata
C. macracantha
C. holmesiana
C. canadensis
C. ellwangeriana
. densiflorum
C. maximowiczii
Crataegus prunus
C. sanguinea
C. chlorosarca
C. monogyna
C. submollis
C. turkestanica
C. pentagyna
C. pringlej
C. arnoldiana
C. Crus-Galli
C. rotundifolia
70,056,37
62,6710,45
119,8210,02
82,792,01
23,514,28
50,955,70
119,736,59
125,848,20
87,7412,82
86,615,47
87,635,62
53,893,39
102,417,94
39,535,19
63,183,78
106,996,90
87,206,44
65,214,90
46,703,80
56,044,16
30,573,89
52,993,85
66,236,0
79,993,57
92,106,08
70,314,30
S
265,1915,14**
208,9111,32
225,3911,21
368,3426,31**
280,3219,66**
228,9914,20
281,0613,92**
325,7714,99**
232,4514,25
231,5811,29*
223,3211,04
185,5313,89
375,4541,07**
462,0733,59**
422,4641,88**
329,5635,38**
408,4630,72**
334,3718,72**
463,5636,91**
.
1,270,028
1,370,047
1,340,028
1,340,041
1,320,035
1,360,035
1,350,028
1,330,038
1,440,047
1,310,026
1,380,041
1,310,058
1,290,031
1,290,033
1,380,072
1,300,035
1,360,029
1,330,032
1,320,042
* 0,95; **
0,99 ( C. monogyna).
(. 4). . . (. 3). . .
. . Densiflorum
. C. canadensis
. C. submollis
. C. turkestanica
. C. Crus-Galli
. C. ellwangeriana
27
______________________________________________________________
. C. sanguinea
. C. prunus
. C. macracantha
. 4. Crataegus
, , , ,
, , , , .
(-0,392).
rataegus L. .
,
.
: ,
, (
). , , , ,
, ,
.
,
, .
20
: , , .
, , 2 .
, ,
.
1. .. . .: , 1974. 224 .
2. . . (Crataegus L., Rosaceae) //
3 / 2004. . 18.
3. .. /
.. / . .. . .: . , 1988. 208 .
4. . ., . . ( ). .: , 2003. 224 .
5. .. / .. . .: , 1953. 152 .
6. .. - / .. // , , -
28
_______________________________________________________________
: XII . : , 2009. . 44-48.
7. , ..
/ .. , .. // : . .-. ., 19-22
2006. : - , 2006. . 344.
8. .. /
.. , .. : , 1991. 307 .
9. ..
/ .. , .. , .. //
. 2010.
. 42, 1. . 3-21.
10. .., .. - Ribs americanum Mill. //
: , , ,
: . .-. . : , 2008. .133-135.
11. .. - Ribs
americanum Mill. : . . .
. , 2010. 20 .
12. .. - /
.. , .. // , , : XII . . . : , 2009. . 103-107.
13. , .. / .. , .. . .: , 1986. 72 .
14. .. . : , 1971. 388 .
15. , . / . . .: , 1969. 572 .
29
______________________________________________________________
582.715:581.522.4
SEDOIDEAE BERGER (CRASSULACEAE DC.) - 1
..
..
, . ,
. , 85.
e-mail:
orlova-belgorod@yandex.ru,
Sorokopudova@bsu.edu.ru
25
- .
-
. S. rupestre, S. sediforme,
S. sarmentosum, H. spectabile.
: , , , , .
30
_______________________________________________________________
H. Ohba, H. telephium `Matrona`, H. cauticola (Praeger) H. Ohba, Ph. kamschaticus
(Fisch) t Hart, Ph. kamschaticus f. variegatum, Ph. kamschaticus f. album, Ph. spurius
(M. Bieb.) t Hart ( Ph. spurius1, Ph. spurius2, Ph. spurius3), Ph. spurius var.
variegatum, Ph. spurius Album, Ph. hybridus (L.) t Hart, Ph. aizoon L., S. sediforme (Jacq.)
Pau, S. acre L., S. acre `Elegans`, S. album L., S. album f. murale, S. sexangulare L., S. hispanicum L., S. rupestre L., S. rupestre `Aureum`, S. sarmentosum Bunge).
GRIN Taxonomy for Plants [4].
,
. - , [5].
,
. : ,
, , .
. 3-
4- , 0 (,
), 1 3 .
3 , 2 , (30-60 %),
1 ( , ), 0 , ( ) (. 1).
Sedum Phedimus ( ,
, ).
H. spectabile, H. telephium subsp. maximum, H. telephium `Matrona`, Ph. aizoon
, , ( )
. 3
, ;
2 , (
), 1 .
2008-2011 .
. , , .
. 3 ,
, 2 ,
( 5-10 1 ), 1 - ,
50%, 0 , 50%
(, , ).
,
. 3 ,
, 2 , -
31
______________________________________________________________
, 1 ,
.
, , . , , 1 2, , , , , (. 2). 3- .
5- ,
.
1 2 1
1-5 , 2 6-10 , 3 11-16
1 2.
1
2-7 , 2 - 8-13 , 3 14 .
1 ,
13-16 , 2 17-20 , 3 21-24 ,
4 25-28 , 5 29 .
, .
1 , , 2 , (
, -, , - ), 3 , .
1 -- , , 2 -
, ,
, 3 -
, ,
. ,
Phedimus Hylotelephium H. spectabile H. telephium `Matrona` -
.
()
: (1), (2), (3), (4), (5), (6). 3 ,
, 2 3-4 , 1 , 1-2 .
. 3
14-17 ; 2 11-13 ; 1 -
, 8-10.
.
:
(5-8 ), (9-12 ) (13-15 ).
________________________________________________________________
Sedoideae, , ,
Ph. kamtschaticus
Ph. kamtschaticus f. variegatum
Ph. kamtschaticus f. album
Ph. spurius
Ph. spurius var. variegatum
Ph. spurius 1
Ph. spurius 2
Ph. spurius 3
Ph. spurius Album
Ph. hybridus
Ph. aizoon
3
2
2
3
2
2
1
2
3
3
3
2
2
3
3
2
3
3
3
3
2
2
S. sexangulare
S. acre
S. acre `Elegans`
S. sediforme
S. hispanicum
S. sarmentosum
S. rupestre
S. rupestre `Aureum`
S. album
S. album f. muralis
3
3
2
0
3
0
0
0
0
0
3
3
3
3
3
2
3
3
3
3
H. spectabile
H. cauticola
H. telephium subsp. maximum
H. telephium `Matrona`
0
1
3
0
3
2
2
3
family Aphididae
Phedimus Raf.
2
3
2
0
2
1
3
0
2
1
2
1
1
3
2
3
2
3
2
1
1
1
Sedum L.
2
3
2
3
2
3
2
0
2
3
2
3
2
0
3
2
3
3
3
3
Hylotelephium H.Ohba
2
0
2
3
1
0
3
0
3
3
3
3
3
3
3
3
3
3
3
13
9
11
12
10
11
11
13
14
11
10
3
3
3
3
1
1
2
3
3
3
14
14
13
8
12
8
7
11
12
12
3
3
3
3
8
11
9
9
* , ,
32
32
33
2
Sedoideae, , ,
S. sexangulare
S. acre
S. acre `Elegans`
S. sediforme
S. hispanicum
S. sarmentosum
S. rupestre
S. rupestre `Aureum`
S. album
S. album f. muralis
H. spectabile
H. cauticola
H. telephium subsp. maximum
H. telephium `Matrona`
2
2
2
2
2
2
2
2
2
2
2
11
13
12
14
13
12
9
13
13
12
8
3
3
3
3
3
3
3
3
3
3
13
13
12
12
13
11
11
14
17
16
2
2
2
2
13
11
10
14
Ph. kamtschaticus
Ph. kamtschaticus f. variegatum
Ph. kamtschaticus f. album
Ph. spurius
Ph. spurius var. variegatum
Ph. spurius 1
Ph. spurius 2
Ph. spurius 3
Ph. spurius Album
Ph. hybridus
Ph. aizoon
1 2
Phedimus Raf.
2
1
4
1
1
2
1
4
3
1
2
1
4
1
2
2
1
5
2
2
1
1
4
3
2
1
1
4
2
2
1
1
1
2
2
1
1
5
2
2
2
1
4
2
2
2
1
4
1
2
1
1
2
1
1
Sedum L.
3
1
2
1
3
2
1
3
1
3
2
1
1
2
3
2
1
1
2
3
3
1
1
2
3
2
1
1
1
3
1
1
1
2
3
1
1
3
3
3
3
1
5
2
3
2
1
5
2
3
Hylotelephium H.Ohba
2
2
5
1
1
1
1
5
1
1
1
2
3
1
1
1
3
5
2
1
______________________________________________________________
33
34
_______________________________________________________________
. (13-15 ) Ph. kamtschaticus, Ph. spurius, Ph. spurius3, Ph. spurius Album, S. sexangulare, S. acre, S. acre `Elegans`; (9-12 ) - Ph. kamtschaticus f. variegatum, Ph. kamtschaticus f. album,
Ph. spurius var. variegatum, Ph. spurius1, Ph. spurius2, Ph. hybridus, Ph. aizoon,
S. rupestre `Aureum`, S. hispanicum, S. album, S. album f. muralis, H. cauticola, H. telephium subsp. maximum, H. telephium `Matrona`. S. rupestre, S. sarmentosum, S. sediforme H. spectabile, - (7-8). H. spectabile, S.
rupestre, S. sediforme , S. sarmentosum
. , , (11-13 ).
, - Ph. kamtschaticus, Ph. kamtschaticus f. variegatum, Ph. kamtschaticus f. album, Ph. spurius, Ph. spurius var. variegatum,
Ph. spurius1, Ph. spurius2, Ph. spurius3, Ph. aizoon, S. album, S. album f. muralis, S.
rupestre `Aureum`, S. sexangulare, S. acre, S. acre `Elegans`. Ph. kamtschaticus
Ph. aizoon .
- , , S. rupestre, S. sediforme, S. sarmentosum, S. hispanicum, H. cauticola, H. spectabile, H. telephium subsp. maximum, H. telephium `Matrona`, ( , , ).
1. .., .. . .: , 2006. 95 .
2. .. . : . .: +, 2011. 432 .
3. .., .., .. -
// , .
. . . . --
; . , 2003. . 46-49.
4. Genera of Crassulaceae subfam. Sedoideae / United States Department of Agricul-ture.
Germplasm Resources Information Network (GRIN). 2007. : http://www.arsgrin.gov/cgi-bin/npgs/html/gnlist.pl?1764.
5. .., .. // . . . . 1978. . 107.
. 80-86.
35
______________________________________________________________
582.715:581.522.4
CHAENOMELES (THUNB.) LINDL 1
..
..
, . ,
. , 85.
e-mail: irina.navalneva@mail.ru,
sorokopudov@bsu.edu.ru
Chaenomeles L. (Rosaceae)
(Maloideae). Chaenomeles .
, , . , , . ,
,
. ,
, .
.
, [2].
: . . , 2,5 3,5 [4].
, .
( ) - -. .
; . ; [3].
,
. , , -, , .
, .
[4].
1
-
2009-2013 , 267 - Chaenomeles .
36
_______________________________________________________________
( 6) , , ,
[3].
Chaenomeles.
Chaenomeles.
.. [1]. : Ch. japonica Ch. maulei.
,
. .
, ( ) ( , ).
(, ).
, .
Chaenomeles 2-3 .
. 15-20 .
37 .
50 7080 2030
. 1000 .
: 4080 , 100120 ,
34 .
, (, ),
, . . .
2
: , . , , .
( )
. : , . Chaenomeles .
.
18 , 50,0 %,
23. : ,
, . ,
, , , (. 1).
37
______________________________________________________________
. 1.
, . , , .
, .
10
1-2 . . , 12 .
.
, . , .. , , , . , - . , . - .
()
(. 2). , 1, 2 6
, .
, .
3, 4, 5 7 , .
. , 3 5.
38
_______________________________________________________________
100,00
90,00
80,00
70,00
60,00
%
50,00
40,00
30,00
20,00
10,00
0,00
1
. 2. ()
,
(. 3).
. 3.
(),
. . , . , . .
.
. , .
, , .
, , , , , , , , .
39
______________________________________________________________
, . - . . .
.
, , .
(), .
. , , . , , . , , , ,
. .
, . . , , . , . , , .
, . , , ,
, . , , , , , , . ,
, , .
. , , , .
, .
, , ,
20 .
,
. , 4,35 13,27 , 4,56 16,40 . , 2,50 11,83 . : 12,66*8,45-16,80*13,25 2 (
). , , 11,550,13 , 1,000,00 .
, 3,300,15 ,
4,820,08 . , 1,920,06 ,
1,060,08 (. 1).
1
,
4,35
0,10
13,27 0,25
4,56
0,06
16,40 0,63
2,50
0,05
11,83 0,08
12,66 0,18
16,80 0,05
8,45
0,14
13,25 0,06
11,55
0,13
1,00
0,00
3,30
0,15
4,82
0,08
1,92
0,06
1,06
0,08
40
_______________________________________________________________
, Chaenomeles
: , , ,
, ,
,
.
1. .. Pentaphylloides Fruticosa
(L.) O. Schwarz, // . 2003. . 39. . 4.
. 68-76.
2. . 6- . . 5 (2). . .: , 1981. 511 .
3. .. . .: - ; : , 2004. 62 .
4. , .. , ..,
- // : . . .-. : , , , 24-27 2008. , 2008. . 402-406.
41
______________________________________________________________
634.722:582.717
RIBES RUBRUM L.
1
..
..
, . ,
. 85
E-mail: sorokopudov@bsu.edu.ru
Ribes rubrum L. .
: , . .
: Ribes rubrum L., ,
,
, , , ,
[1].
,
, , [2].
,
.
. , ,
,
, , 1000
. 27 2
[3].
: ( 5 ), ( 5 8 ),
( 8 ) [4].
2007 ,
4,6 9,3
(. 1). 2007 (4,5), (4,6), (4,7), (4,8), 14%
(. 2). 2007 7% (9,3) (8,2). ,
83% , .
1
2007
5,5
8,0
6,7
,
2008 2009
4,0
5,2
5,3
2,8
5,3
4,2
2010
4,7
5,0
5,8
.
4,860,66
5,282,14
5,531,05
14
41
19
- 2009-2013 ., 508 14.05.2010 .
1
42
_______________________________________________________________
1-63-22
56-2
()
5,8
9,3
5,3
7,0
6,7
6,5
4,7
5,7
5,3
6,6
8,2
5,0
6,3
4,6
6,8
8,0
6,0
5,3
4,8
4,5
6,5
5,5
6,7
5,5
7,2
5,0
7,0
8,8
3,7
5,8
4,8
11,7
5,00
4,4
4,6
8,0
6,8
5,2
9,3
3,9
2,8
9,5
4,6
6,5
5,6
6,8
5,6
3,8
4,4
4,4
6,0
4,6
6,0
7,0
4,3
4,1
3,0
7,7
3,2
4,8
3,3
7,5
4,3
4,7
8,0
3,5
2,4
8,0
6,2
4,9
4,3
3,3
6,5
4,9
3,1
3,4
8,6
3,0
7,1
7,4
4,7
6,8
4,3
9,0
5,4
5,0
4,3
8,4
5,2
5,4
6,4
4,3
7,8
7,3
6,6
5,0
5,4
2,8
5,4
4,4
4,6
4,8
7,7
3,7
6,480,68
8,121,12
4,520,65
5,921,32
4,71,54
8,722,22
4,570,97
4,950,56
4,360,85
7,630,78
6,131,73
5,060,30
7,501,43
4,060,47
4,981,76
8,210,91
5,840,87
5,420,74
5,020,60
4,351,78
6,010,57
4,640,74
4,71,48
5,540,86
7,111,31
4,090,89
10
14
14
22
33
25
21
11
20
10
28
6
19
15
55
11
15
14
12
41
10
16
32
19
17
22
2008
2,8 () 11,7 ( ).
, 41%. 2007 ,
45%. , , , 2008
. ,
, , , , , , , , , 1-63-22.
,
2008 14% .
, , , .
2
( 5 )
( 5 8 )
( 8 )
2007
14
79
7
, %
2008 2009 2010 2007-2010
41
66
34
42
45
34
59
48
14
7
10
2009 , 8
(), .
2,8 8
. (66%) 5 .
, 2008 .
34% .
2010 (59%)
5 8 ().
35%, 7%.
43
______________________________________________________________
2,4 11,7 . 20072010 42%. ,
, , , , , , , , , , , . (48%) - ,
, , , , , ,
, , , , , 1-62-22, 56-2. ,
, 10 %.
, ( 0,55),
(0,41), (0,41), (0,32). (0,06), (0,10),
(0,10), (0,11), (0,11), (0,12), (0,12), (0,13), (0,14), (0,14).
1000 4,8 7,5 . (. 3).
3
1000
1-63-22
56-2
()
1000 ,
2007 2008 2009 2010
6,00
7,33
5,32
5,50
6,45
7,42
5,20
5,72
6,00
5,04
5,79
6,34
6,12
7,14
4,81
5,05
6,12
8,49
4,80
5,33
6,12
8,64
6,40
6,97
5,83
8,00
4,80 6,40
5,03
7,34
5,93
5,20
6,00
7,14
5,0
5,37
6,35
9,33
8,3
5,90
5,95
8,58
6,50
5,97
7,05
8,04
7,7
6,80
6,04
8,0
5,3
5,7
6,88
8,63
6,00 5,80
5,56
7,54
4,6
5,34
5,75
7,03
4,30
6,25
8,07
8,90
6,20 6,00
6,64
8,90
7,10
5,50
6,76
7,89
5,40
5,86
6,03
7,04
5,40 6,90
5,00
6,25
5,90
4,50
5,90
7,14
5,33
5,90
5,80
7,82
4,55
5,71
5,04
6,02
3,85
4,23
6,02
7,0
5,90
6,4
7,75
8,09
6,50
6,78
6,01
6,68
5,00 5,80
5,32
6,25
4,2
5,00
5,95
7,12
6,3
6,07
.
6,060,91
6,200,96
5,790,55
5,781,07
6,191,63
7,031,13
6,261,34
5,881,05
5,880,94
7,471,62
6,751,25
7,390,57
6,261,20
6,831,29
5,762,56
5,831,15
7,291,42
7,041,41
6,481,10
6,340,77
5,410,80
6,070,76
5,971,36
4,790,96
6,330,49
7,280,76
5,870,69
5,190,85
6,360,53
15
16
10
19
26
16
21
18
16
22
18
8
19
19
22
20
19
20
17
12
15
13
23
20
8
10
12
16
8
(41%) 1000 6
7 . 38% 1000 6 .
(4,8 ), 56-2 (5,2), (5,4), (5,5 ), (5,8), (5,8 ), (5,8), (5,9 ), (5,9 ), 1-63-22 (5,9 ).
44
_______________________________________________________________
(21%) 1000 7 . (7,5 ), (7,4 ), (7,3 ),
(7,3 ), (7,0), (7,0).
0,08 0,26.
(0,08), (0,08), (0,08), (0,10). 1000 (0,26), (0,237), (0,22), (0,22), (0,21).
, . ,
.
4,5 8,3% (. 4). 5% (4,5%),
(4,5%), (4,5%), (4,7%),
(4,9%). (76%) 5 7,5%. (8,2%),
(8,3%), (7,6%), (7,9%).
(0,23), (0,22), (0,21). (0,04),
(0,07), (0,07).
1000
(r= - 0,36),
(r=0,16), 1000 (r =0,31).
4
1-63-22
56-2
()
2007
6,31
6,00
4,50
7,80
6,00
7,00
5,35
4,65
7,75
7,04
6,00
5,34
6,20
6,40
6,05
4,85
5,51
5,16
5,00
7,14
6,00
4,85
4,92
4,06
5,18
5,16
5,12
4,87
3,88
, %
2008 2009
7,00
7,20
4,85
5,70
5,30
3,90
9,02
8,2
7,05
6,5
7,85
6,1
8,2
4,9
6,03
4,1
8,34
7,9
8,70
8,00
8,10
6,48
6,80
7,40
7,88
6,86
8,84
6,90
8,70
6,14
7,70
5,62
7,88
7,00
7,20
4,8
5,23
4,70
8,68
7,60
7,08
6,5
6,60
4,70
6,76
5,00
4,73
3,49
7,04
6,46
8,12
6,10
6,05
4,70
6,00
6,60
5,30
4,40
2010
6,2
4,61
4,40
7,70
5,54
5,35
4,80
5,2
9,0
6,80
5,80
6,10
6,48
6,50
7,20
4,90
6,60
7,10
4,90
8,08
7,50
5,80
5,71
5,80
7,43
6,89
4,70
5,80
5,16
.
6,680,50
5,290,67
4,530,58
8,180,60
6,270,65
6,581,09
5,811,61
4,500,82
8,250,56
7,640,88
6,601,04
6,410,89
6,860,73
7,161,14
7,021,23
5,771,34
6,750,98
6,071,26
4,960,22
7,860,66
6,770,66
5,490,89
5,600,85
4,520,99
6,530,98
6,571,25
5,140,64
5,820,72
4,690,67
.
, %
7
13
13
7
10
17
28
16
7
12
16
14
12
16
18
23
15
21
4
8
10
16
15
22
15
19
12
12
14
45
______________________________________________________________
1. (41%) 1000 6
7 . 1000 6 38% : (4,8 ), 56-2 (5,2), (5,4), (5,5 ), (5,8),
(5,8 ), (5,8), (5,9 ), (5,9 ), 1-63-22
(5,9 ). 1000 7 (21%): (7,5 ), (7,4 ), (7,3 ), (7,3 ), (7,0),
(7,0).
8 26. (8), (8), (8), (10).
(26), (23), (22), (22), (21).
2.
4,5 8,3% : (4,5%), (4,5%), (4,5%), (4,7%), (4,9%).
(8,2%), (8,3%),
(7,6%), (7,9%).
(23), (22), (21). : (4), (7), (7).
3. 1000
(r= - 0,36),
(r=0,16), 1000 (r =0,31).
1. , .. / .. , .. . : , . -, 2003.
296 .
2. .. / .. , .. ,
.. : , 2008. 62 .
3. .. : /
.. , .. , .. , .. , .. , .. ,
.. , .. , .. , .., .. , .. ,
.. : - , 2008. 36 .
4. , .. - . Grossulariaceae Dumort. / .. , .. , .. ,
.. , .. , .. , .. // . 2009. 1. . 111 117
Seed efficiency of grades of red currant Ribes rubrum L is studied. In the conditions of the Belgorod region. Grades are divided on
groups. The share of seeds from weight of berries as a perspective direction in selection on it is not enough seeds is defined.
Key words: Ribes rubrum L., seeds, a share of seeds from weight
of berries, weight of seeds.
46
_______________________________________________________________
582.717.7
RIBES L.
..
, 677980
. , . , 41
e-mail: korobkova_t@list.ru
, .
8 : R. fragrans Pall., R. procumbens Pall.,
R. dikuscha Fisch, R. pauciflorum Turcz ,R. nigrum L. spp sibiricum
E. Wolf, R. triste Pall, R. palczewskii Pojark, R. glabellum Trautov.
et Mey.
: , , ,
, .
, , , ..
- . , . , . Ribes . ,
, ,
, , . -, , .
, R. vulgare (R.rubrum) ,
, (R. triste).
Ribes .. [11]
R. rubrum sense Jancz.
Rubrae A.Pojark., , R. spicatum. , . R.spicatum,
( ) -R. pubescens, - R. hispidulum,
, - - R. cidum,
, - R. palzewskii [3].
Ribesia
. : R. glabellum (Trautov. et Meyer) Hedl, R. rubrum var glabellum,
R. acidum, R. spicatum Robson, R. hispidulum (Jancz) Pojark [1];
R. acidum Turcz. ex Pojark (R. rubrum Pojark) . , [13] ;
R. rubrum L. s. 1 - R.acidum Turcz. ex Pojark, R. glabellum . ,
[17]; R. rubrum L. s. 1 - R. acidum Turcz. ex Pojark, R. hispidulum (Jancz) Pojark, R. palczewskii (Jancz) . Pojark, R. spicatum Robson s.str. . , [6].
Eucoreosma Jancz. 16 5 [2, 11,
13]. : (R.dikuscha Fisch.),
(R. ussuriense Jancz.), (R. fontaneum Boczk.), (R. pauciflorum Turcz.), (R. procumbens Pall.),
R. nigrum L. [16].
47
______________________________________________________________
R. nigrum L. . . [13] 4 11 , 4 10 . Ribes L. . ,
. , R. rubrum L. R. vulgare Lam. , R. pauciflorum Jurcz, R. nigrum L. spp europaeum Jancz, R. nigrum L. spp sibiricum E. Wolf. - R. nigrum L [6,19].
6 8 .
, Ribes acidum
Turcz. ex. Pojark [8]. .. , Ribes glabellum Hedl
Ribes acidum Turcz. ex. Pojark. [17]. , ..
[18] .. , .. .
[4] Ribes glabellum Hedl. Ribes acidum Turcz. ex. Pojark Ribes
glabellum Hedl. R. spicatum Robson.
: R. fragrans
all., R. procumbens Pall., R. dikuscha Fisch, R. pauciflorum Turcz, R. triste Pall., R. acidum (R. rubrum L., R. glabellum Trautov. et Mey), R. alczewskii Pojark [8].
R. fragrans Pall., R. procumbens Pall., R. dikuscha Fisch., R. pauciflorum Turcz. ex
Pojark (R. kolymense (Trautov) Kom. ex Pojark, R. cyanthiforme Pojark), R. triste Pall.,
R. glabellum (R.rubrum var glabellum Trautov et Meyer, R. acidum Turcz. x Pojark,
R. rubrum auct non L.), R. spicatum subsp. alczewskii (Jancz.) (R. rubrum
var.palczewskii (Jancz.), R. palczewskii (Jancz.) Pojark) [18].
R. fragrans Pall., R. procumbens Pall., R. dikuscha Fisch., R. nigrum var. pauciflorum Jancz, R. nigrum var. kolymense Trautov, R. triste Pall., R. acidum Turcz. x Pojark,
R. spicatum subsp. Palczewskii (Jancz.)Malyschev [17].
R. fragrans Pall., R. procumbens Pall., R. dikuscha Fisch, R. nigrum (R. kolymense
Kom. ex Pojark, R. pauciflorum Turcz ex Pojark), R. triste Pall., R. rubrum L. (R..acidum
Turcz. x Pojark, R. hispidulum(Jancz.) Pojark, R. palczewskii (Jancz.), R. spicatum Robson) [6].
R. fragrans Pall., R. procumbens Pall., R. dikuscha Fisch, R. nigrum L., R. glabellum Trautov et ey, R. triste Pall., R. spicatum Robson; R. palczewskii (Jancz.) [4].
, ,
, , , .
.
70-
, . Ribes L. ,
,
.
[7, 12].
. ,
. . ,
(.). ( 8-12, 15-20,
[5]. ..
[10], .. [14] ,
.
48
_______________________________________________________________
,
,
R. nigrum var. kolymense [Trautv] Kom. ex
Pojark ( -,
. , .
)
( 1),
R. pauciflorum Turcz
(
1980 -,
70 . ,
,
.
.
,
,
, ,
. , , 3 3-6 .
- , -
.
.
( 3 - 6 )
,
.
, ,
. Ribes rubrum L.
Ribes vulgare Lam., - , , Ribes rubrum L.
Ribes vulgare Lam., .
R. nigrum spp sibiricum E. Wolf. R. pauciflorum Jurcz.
. R. spicatum
R. palczewskii .
. Ribes alczewskii
, , , , , .
. . - , , . -. (9-15) , , 1 3.
R. spicatum , , , , , , , . -, , .
, , R. pauciflorum Turcz , R. fragrans Pall, R. procumbens
Pall, R. dikuscha Fisch, R. nigrum L. spp sibiricum E. Wolf., Ribes glabellum Hedl,
R. palczewskii (Jancz.), R. triste Pall.
1. . .: , 1971. .6. .228-230
2. . . . , 1973.
184 .
49
______________________________________________________________
4. .. . : - ,
2005. 328 .
5. .. : ...., 2001. 15 .
6. .., .. .
: - , 2002. 707 .
7. .. . ., 1972.
284 .
8. . : , 1974. 542 .
9. . . . .: , 1930.
196 .
10. . . . .; .: , 1955. 277 .
11. .. Ribes L. // . .; ., 1939. .9.
. 226-267
12. , .
: - , 1999. 608 .
13. . . . , , . .: ,
1987. 216 .
14. // , , . . : , 1987. .3. .12-20.
15. .. ( )
: . . . . . -,
1994. 22 .
16. .., .., ..
. : , 2005. 120 .
17. .. . , 52 .
18. : Berberidaceae Grossulariaceae. : , 1994. .7.
312 .
19. .. . .:
, 1995. 992 .
50
_______________________________________________________________
582.572.226:575.224.46.044
..
.. 2
1
, 308033,
. , . 11
e-mail: Labunskaya@bsu.edu.ru
2
, 308015,
. , . 85
e-mail: Sorokopudova@bsu.edu.ru
, ,
22 .
, . ,
: Connecticut King
( 4n = 48),
( 4n = 48),
( 4n = 48).
: , , , , .
2n=24,
L. lancifolium Thunb., L. rhodopaeum Delip., L. szovitsianum Fisch. et Ave-Lall. (2n=36) [1,2].
. , , , , , ,
, , . , [3-5], (
70%, 86%, 65%)
;
, [6-8].
2006-2010 .
[9].
22 : (Longiflorum/Asiatic) (Algarve, Royal Delight, Royal Drim, Royal Fantasy, Royal Justice, Royal Sunset, Showroom, Spirit,
84-592), (, , , , , , , , , , Connecticut King, Rosello), , Lilium henryi Baker.
, (
) ,
[9].
. 10-
51
______________________________________________________________
0,2 % 6 , 3
, -.
5- , L. henryi,
, - .
- ,
, ,
, ,
(. 1, 2).
. 1.
-
. 2.
1,5 - .
,
, (, Royal Delight, ) 22
. .
, , .
() , 48,5%.
Royal Delight
. . , - ; , , . , , .
, ,
, .
, , . ,
(3 ) 30%, 31%, 48% , 37%, Connecticut King (2 ) 60% 72%, (2 )
31% 49%, (2 ) 62% 60%, 57%.
52
_______________________________________________________________
,
.
, , 14
( 25% ) 4: (4n = 48) (. 3), (4n = 48) (. 3),
Connecticut King (4n = 48) ( , . 3 ,).
. 3.
( , , , Connecticut King)
, 288 22 ,
0,2% 6 ,
195 , 6 (2% ), .
37-72%.
: Connecticut King
66% , - 6%,
- 4%, , , -
53
______________________________________________________________
, ,
.
1. , .. / .. . .: ,
1980. 300 .
2. / .: .. , .. ,
.. , .. . .: . . -, 1969. 927 .
3. , .. / .. . .: . . .
. . . . . , 1982. . 176-178.
4. . : - , 1992. 32 .
5. , . / . . 2008. :
http://www.molo.ru/articles,special,1,14.htm.
6. , .. : . /
.. ; . .. , .. . 3- ., . : .
. -, 2006. 479 .
7. , .. / .. . 1998. : http://svitlo.by.ru/bibloteka/ot_sozd_mira/ot_sozd_mira11.html.
8. Polyploidy. 2003. : http://www.biologie.uni-hamburg.de/bonline/e37/37d.htm.
9. , .. / .. . : - , 2005. 244 .
e-mail: Labunskaya@bsu.edu.ru
Belgorod National Research
University,
Pobeda str., 85, Belgorod,
308015, Russia
2
e-mail: Sorokopudova@bsu.edu.ru
54
_______________________________________________________________
581.6 : 631.527
(FRAGARIA ANANASSA DUCH.)
.. 1
. . 2
1
, 630039,
, . , 160
e-mail: childofsiberia@mail.ru
2
, 630090, ,
- . , 10.
e-mail: SO_baturin@mail.ru
1996
- , . (Fragaria x
ananassa Duch.), 5 ,
30 , .
.
: , Fragaria x
ananassa, Comarum palustre, , ,
.
55
______________________________________________________________
.
.
-
Pink anda () 61-1 61-2 F1 C 141
( ABZ Seeds Gourmet Starwberries [6]. F1
( - 97/3-76-6, 97/2-1-4 97/3-75-6,
97/2-1-6), F. x ananassa
Pink anda.
Pink anda. F2 :
00/9-30-5 [97/3-75-6 97/2-1-4]; 05/11-51-2 [96/10-78-4 (58 x Pink Panda)].
, . . ,
.
.
. , . .
3- +5+6 ,
5-7 .
RHS [9].
(. )
:
1)
, ;
2) ,
, . 1996 Pink Panda.
, , ,
, . 60%
[4]. Pink Panda ,
. .
. , -
Pink anda . () . Pink anda Fragaria
x ananassa , . 707 , .
48,7%. 91,3%.
56
_______________________________________________________________
Pink anda. .
- -.
. 24 (41,4%) 58
. - , .. , Pink anda.
05/11-51-2 00/9-30-5. , , .
F1 C 141, - .
, . , , [10]. ( 7-9 61-2 7-28 61-2),
80-90% . 20
: 90%
01 , 0 - , 5 .
: 1) ; 2) - 3) . ,
. , , 19
, 3 , 17 , 6 .
, 4,5 , ,
. . .
, ,
, .
- ,
(. 1),
. , , 90-97 % (08/15-12-5 08/15-4-2;
08/15-12-5 08/47-27 .). 92% (. 2). , (2n=8x=56)
, .. ,
. , ABZ Seeds
Gourmet Starwberries (),
(Pikan F1, Florian F1, Merlan F1, Tarpan F1, Roman F1, F1 C141) (Sarian F1, Grandian F1, Loran F1,
Chesan F1), [11].
(Sarian F1,
57
______________________________________________________________
Grandian F1, Loran F1, Chesan F1, Florian F1) , Chesan F1, Loran F1 Sarian F1 ,
Florian F1 Grandian F1 ( ,
, .) [12].
.1. -
. 2.
.
-
58
_______________________________________________________________
,
. .
- , .. .
. , , , [12].
, , .
.
, 30 . . . 500 . , .
1. Mangelsdorf A.J., East E.M., Studies on the genetics of Fragaria // Genetics. 1927. 12.
. 307-339.
2. .. . .: , 1975. - 184 .
3. Ellis J.R. Fragaria-Potentilla intergeneric hybridization and evolution in Fragaria // Proc.
Linnean Society of London. 1962. V.173. . 99-106.
4. Baturin S.O., Ambros E.V. The use of intergeneric crosses in strawberry breeding // International Meeting of Young Scientists in Horticulture. Materials of the 7th Intern. Conf. Lednice,
Czech Repablic. 1999. P.13-17.
5. Khanizadeh, S. New Hardy Day-Neutral Red Flowering Strawberry Cultivars // Acta Hort.
(ISHS). 2000. 538. . 779-780.
6. Bentvelsen G.C.M., Bouw E. Breeding Ornamental Strawberries // Acta Hort. (ISHS).
2006. 708. P. 455-457.
7. .., ..
(Fragaria x ananassa Duch.) // .
2010. . 14. 1. . 165-171.
8. Khanizadeh, S., J. Cousineau, M. Deschnes, A. Levasseur, O. Carisse. Roseberry and Rosalyne: two new hardy, day-neutral red flowering strawberry cultivars // Acta Hort. (ISHS). 2002.
567. P. 173-174.
9. Tucker A.O., Maciarello M.J., Tucker S.S. A survey of color charts for biological descriptions // Taxon. 40. 1991. P. 201-214.
10. .., .. (Fragaria L.) // i , i i: .
. . . 2007. . 2. . 16-20.
11. Bentvelsen G.C.M., Bouw E., Veldhuyzen van Zanten J.E. Breeding strawberry (Fragaria x
ananassa Duch.) from seed // Acta Hort. (ISHS). 1997. Vol. 1. 439. P. 149- 153.
12. .., .., ..
// . 2010. 1. . 40-45.
59
______________________________________________________________
INTRODUCTION OF PINK FLOWERING GARDEN STRAWBERRY (FRAGARIA
ANANASSA DUCH.) IN WESTERN SIBERIA
L.L. Kuznetsova1
S.O. Baturin2
1 Novosibirsk
State Agrarian
University, Novosibirsk, Russia
e-mail: childofsiberia@mail.ru
2 Institute
e-mail: SO_baturin@mail.ru
At 1996 in IC&G SB RAS (Novosibirsk, Russia) a program on creation pink flowering forms of garden strawberry with double purpose (ornamental and berrylike) as well as forms adapted for growing in Siberian region has been started. The gene pull of pink flowering strawberry
included 5 varieties of European selection, 30 winterhardy and highyielding hybrids with ornamental qualities of flowers was formed. All selected hybrids are day-neutral strawberry varieties during summer and
autumn period of vegetation.
Key words: pink flowering strawberry, Fragaria x ananassa, Comarum palustre, selection, day-neutral strawberry, seed germination.
60
_______________________________________________________________
: 582.951.64:581.8
(VERONICA AUSTRIACA L.)
..
..
,
305041, . , . .,3
e-mail: salvia_julia@mail.ru
. , .
: (Veronica austriaca L.),
,
61
______________________________________________________________
. 1. (. 140)
. 3.
(. 400)
. 5.
(. 200)
. 2.
(. 120)
. 4.
(. 300)
. 6.
(. 200)
62
_______________________________________________________________
. 7.
(. 200)
. 8.
(. 200)
.
. 5-6 .
. , .
, . 1-3
, .
.
, .
. ,
. , .
.
, (. 3, 4).
( ). .
, . .
. , - -,
, -
, (. 5). - .
,
.
, .
. , (. 6). . . , --, , ,
63
______________________________________________________________
, . .
(. 8). , , (. 7).
,
.
1. . 11- . .: , 1987. . 1. 277 .
2. .., .., .., .. . . 3: (: ). : - , - . 2004.
.105.
3. .., .. :
, 2005.- 80 .
4. : , , ; Carifoliaceae Plantaginaceae / .. ; . .. . .: , 1990 . 173-182 .
5. . 22 .: , 1955. 329-487 .
V.N. Bubenchicova
Y.A. Kondratova
Kursk State Medical University,
Kursk K. Marks St., 3, 305041
e-mail: salvia_julia@mail.ru
Study of microscopic diagnostic characters of herb of Veronica austriaca L. for identity control. Air-dried herb of Veronica austriaca L. The
histological slides were studied with microscope Biolam C-11 supplied
with a digital camera. Microscopic diagnostic characters of herb of Veronica austriaca L. for identity control were determined. Microdiagnostical sings of the plant material of Veronica austriaca L.have been determined. Then they will be used to the standarting documents for this
plant material.
Key words: Veronica austriaca L., herb, microscopic diagnostic
characters
64
_______________________________________________________________
635.939.73
.. 1
.. 2
1
,
393760, , . , . , 101
e-mail: mgau@mich.ru
2
, 393760, , . - ,
. , 274
. ,
( 75%) 20%
, 50%
.
: , , ,
.
e-mail: mgpi_lab@mich.ru
. , , 7-10
; , , ,
, , . , , ,
. , ,
- .
[1]. .
, .
.
1.
.
2.
.
: , , , , , , , , , , , ,
, , 1-10-49, 1-11-45, 1-17-48, 12-2, 1-39-28,
14-9, 1-67-47, 1-8-57, 1-9-58, 1-9-59, 2-20-14, 2-47-45, 2-56-47, 2-56-48, 2-59-41, 2-61-33,
2-61-35, 2-62-43, 2-65-51.
65
______________________________________________________________
1989 - 2009 . [2, 3],
. .. . , , 100
.
( ) . 10-12 ,
.
, 10-15 ,
.
[4].
Microsoft EXCEL 2000-2007,
STATISTICA 6.1. StatSoft.
.
1
1989 - 2009 .
1-10-49
1-11-45
1-17-48
12-2
1-39-28
14-9
1-67-47
1-8-57
1-9-58
1-9-59
2-20-14
2-47-45
2-56-47
2-56-48
2-59-41
2-61-33
2-61-35
2-62-43
2-65-51
(),%
59,67
58,38
76,64
78,09
90,24
84,69
55,77
70,72
62,54
51,65
84,72
68,69
97,33
75,09
80,50
66,48
57,59
66,81
50,35
96,22
97,00
97,40
79,16
99,00
86,00
71,96
81,06
78,28
61,00
70,21
83,61
79,46
83,21
(Sx), %
0,87
10,67
9,41
17,40
5,29
0,43
14,70
17,60
7,84
16,52
2,84
7,52
1,19
7,67
6,16
3,95
14,39
3,14
5,02
3,07
2,60
9,18
0,47
1,33
12,99
14,48
16,39
5,42
12,08
0,91
0,82
0,54
0,66
, %
59,74
69,05
67,28
60,69
84,95
85,12
41,07
84,11
51,81
58,32
87,56
61,17
98,52
80,66
74,35
70,43
58,60
60,53
55,37
96,79
77,23
94,79
68,53
88,00
80,54
72,43
83,01
65,29
78,00
84,69
67,22
89,00
95,29
0,36
0,40
0,52
0,47
0,77
0,72
0,23
0,59
0,32
0,30
0,74
0,42
0,96
0,61
0,60
0,47
0,34
0,40
0,28
0,93
0,75
0,92
0,54
0,87
0,69
0,52
0,67
0,51
0,48
0,59
0,56
0,71
0,79
,
, , , , , , , ,
66
_______________________________________________________________
, , , 2-56-47, 1-39-28, 2-20-14, 14-9,
2-59-41, 12-2, 1-17-48, 2-56-48.
75% 90% . , , 1-8-57, 2-47-45, 2-62-43, 2-61-33
66% 72% .
, 1-9-58, 1-10-49, 1-11-45, 2-61-35, 1-67-47, 1-9-59,
2-65-51. 50% 62%.
, , 8 33 ( 25%) .
- ,
, .
- 5-6 ,
, , - (, ..).
.
- (.). ,
, , , , , 1-8-57, 12-2, 1-9-59, 1-67-47, 2-61-35, 1-11-45 10,67% 17,60%. ( 5,02% 9,41%)
, , 1-17-48, 1-9-58, 2-56-48, 2-47-45, 2-59-41, 1-39-28, 2-65-51.
, , , , , , , , 2-61-33, 2-62-43, 2-20-14, 2-56-47, 14-9, 1-10-49.
(r = -0,498). (D = 0,248)
25%-
.
.
(.1). , , ,
, , , , , , , 2-56-47, 220-14, 14-9, 1-39-28, 1-8-57, 2-56-48 77% 99%
. , , , , 2-5941, 2-61-33, 1-11-45, 1-17-48 ( 65% 74%). 2-47-45, 12-2, 2-62-43, 1-10-49, 2-61-35, 1-9-59, 2-65-51, 1-9-58, 1-67-47. 41% 61%.
(.1).
. -
67
______________________________________________________________
, , ,
, , 2-56-47, 1-39-28.
0,75 0,96.
, , , , 2-20-14, 14-9,
2-56-48, 2-59-41, 1-8-57 0,59 0,74.
, , ,
, , ,
0,23 0,56. ,
, ,
( 75%)
20% , 50% .
1.
( 75%) 20%
.
2. 50%
23% 56% .
3.
, , , , , 2-56-47, 1-39-28.
1. , .. /.. , .. // . . . . -., 2. - ,
2006. . 82-87.
2. , .
, 1973. 495 .
3. , .
, 1980. 454 .
4. , .. . / .. . ., 1979. 416 .
e-mail: mgau@mich.ru
2 FGOU
e-mail: mgpi_lab@mich.ru
The indicators of open pollination of honeysuckle promising variety samples is reviewed. It was established that the reserves of productivity in open pollination realize a sufficient degree (75%), only about 20%
of modern variety samples. More than 50% of them in these conditions
are able to realize only one-quarter to slightly more than half the capacity of this component of productivity.
Key words: honeysuckle, open pollination, ovary, productivity.
68
_______________________________________________________________
631.527:634.2
1
..
, . ,
. 85
e-mail: sorokopudov@bsu.edu.ru
.
, . .
: , , , , , Armeniaca vulgaris Armeniaca sibirica, Armeniaca mandshurica.
, , , , , .
XIX XX . .. .
, , . - .
.
, , . ,
.
, [ 1].
, , . (,
). .
. 6-9 , , , , -
. , ; ,
. : ; , ; ,
, , [1]. 6-14
, , , .
( -25...-27 )
.
. .
- 2009-2013 ., 508 14.05.2010 .
1
69
______________________________________________________________
.
, ,
, , - . , .
,
: , , .
, ,
, ,
. , .
,
, .
. , , , , , .
,
.
,
. , , . ,
.
, , .
, : Armeniaca vulgaris Lamarck., Armeniaca mandshurica (Maximowicz) B. Skvortzov., Armeniaca sibirica (L.) Lamarck.
, [2].
. . , 2008 13,2 27,5 , 2009 9,5 42,9 . 11, 13, 18, 22, 26, 28 29,7 42,9 , 33,77
26,9 .
17, 24, 32, 37, 42, 45, 54, 55.
11,1 14,5 , 11,7 14,9 .
. -10, -3, -6
-7
. 8- 5.
, 5,2 10,2 ,
11,3 (. 1).
70
_______________________________________________________________
Armeniaca L.
11
13
14
16
16
17
18
22
23
24
26
28
31
26
31
85
93
2009
2009
2009
2009
2009
2009
2009
2009
2009
2009
2009
2009
2009
2009
2009
2008
2009
2009
2009
2009
2008
2009
2008
2009
2009
2009
2009
2008
2009
2009
2009
2009
2008
2009
2009
2009
2009
2009
2009
2009
2008
2009
2009
2008
2009
2009
2009
-10
2009
-4
-5
-8
-9
-12
2009
2009
2009
2009
2009
32
37
42
44
45
46
47
48
51
52
54
55
63
64
65
71
73
74
75
81
82
83
84
,
,
, %
Armeniaca vulgaris
42,90,8
46,90
2,90,2
6,80,2
32,81,1
37,04
2,60,5
7,90,8
28,72,4
36,36
2,81,1
9,91,8
24,32,6
33,04
4,51,0
18,50,5
21,42,0
25,10
2,2,08
10,30,4
14,50,8
14,90
2,01,2
13,41,4
31,71,3
34,22
2,80,6
8,81,2
29,71,8
35,74
2,50,1
8,51.0
23,22,6
33,79
2,50,6
11,01,5
12,91,0
14,70
2,50,7
20,23,0
30,61,3
35,55
2,00,2
6,50,5
30,61,2
33,77
2,50,3
8,20,4
17,70,7
19,50
3,00,2
16,90,5
16,81,0
19,74
2,40,3
14,40,5
20,52,0
27,04
2,30,8
11,50,3
18,61,2
22,30
2,10,3
10,60,3
10,30,7
11,75
2,00,4
19,10,2
11,50,4
12,17
1,80,1
15,60,2
12,88
13,82
2,00,2
15,80,3
17,60,5
18,40
2,50,2
14,20,3
14,01,0
18,20
1,50,4
10,80,3
10,00,5
10,81
1,40,2
14,30,4
20,51,4
27,10
2,20,2
10,20,2
16,31,2
19,58
2,20,1
14,11,1
9,51,1
11,20
1,70,5
18,30,4
20,31,1
23,88
1,90,5
9,51,5
21,40,7
22,52
2,30,3
10,70,2
27,51,3
31,80
3,20,1
10,90,5
17,01,5
19,67
2,80,6
16,80,4
13,80,3
14,50
2,10,4
14,91,5
11,10,5
11,75
1,60,1
14,20,6
15,10,4
16,02
2,10,7
13,61,4
18,91,1
21,20
1,90,4
10,60,4
14,20,9
16,40
2,40,4
17,10,4
15,20,8
17,13
2,20,3
14,50,5
22,01,0
23,99
2,10,4
9,60,3
11,30,6
12,24
1,60,4
14,30,5
11,30,6
12,93
1,80,2
16,30,2
17,00,6
18,30
1,90,3
11,00.1
18,90,6
21,49
2,40,3
12,70,4
22,21,4
26,50
2,80,5
12,50,5
22,11,0
24,93
2,90,4
12,90,6
13,80,5
14,11
2,00,3
14,20,3
13,21,5
18,30
1,20,6
8,90,5
11,10.5
11,92
1,60,6
14,41,0
20,31,1
23,36
2,20,3
11,10,4
13,90,7
15,88
2,50,3
17,90,6
Armeniaca mandshurica
6,60,3
7,04
1,50,2
23,30,2
Armeniaca sibirica
10,20,5
11,32
2,00,3
19,30,3
6,71,5
8,83
1,60,8
22,02,2
6,91,2
9,24
2,00,6
28,30,7
5,20,6
5,88
1,20,2
24,01,0
6,70,6
7,77
1,10,1
17,21,0
71
______________________________________________________________
2
Armeniaca L.
26
28
-31
26
31
32
37
42
44
45
46
47
48
-11
-13
-14
-16
-16
-17
-18
-22
-23
-24
51
52
54
55
63
64
65
71
73
74
75
81
82
83
84
85
93
-4
-5
-8
-9
-12
-10
Armeniaca vulgaris
4,7
5,0
4,8
4,7
4,9
4,3
5,0
4,8
4,0
4,8
4,3
5,0
4,8
4,1
4,7
5,0
3,6
4,7
4,2
4,2
4,9
4,0
4,8
4,9
3,2
4,7
4,4
4,5
4,6
4,2
4,7
4,9
4,2
4,6
4,3
4,7
4,8
4,2
4,4
4,5
-
-
4,6
5,0
4,7
4,4
4,8
4,6
5,0
3,6
4,2
4,8
4,5
4,8
4,2
4,8
4,6
4,7
4,9
4,4
4,6
4,7
4,6
4,6
4,8
4,7
5,0
5,0
4,6
4,6
4,8
5,0
5,0
3,7
4,9
3,9
4,0
4,7
4,9
4,4
4,1
4,4
4,4
4,6
4,6
4,9
4,2
4,5
4,7
4,5
4,5
4,7
4,9
4,5
4,8
4,6
4,7
4,7
4,8
4,8
5,0
5,0
5,0
4,6
4,6
3,8
3,5
4,7
4,7
4,6
4,2
4,6
4,7
4,7
4,8
4,7
4,5
4,6
4,7
4,1
4,2
4,3
5,0
4,0
4,1
4,3
5,0
4,9
4,8
5,0
4,7
5,0
4,6
4,8
4,8
3,8
4,9
4,0
4,8
4,4
5,0
4,5
4,0
4,2
4,1
4,5
4,5
4,7
4,7
4,9
4,6
4,7
4,7
4,6
4,4
4,6
4,6
4,7
4,9
4,6
4,7
4,6
4,9
5,0
4,9
4,2
4,8
4,4
4,8
4,5
4,5
3,9
4,1
4,3
Armeniaca sibirica
4,0
3,0
3,0
3,5
4,0
Armeniaca mandshurica
3,6
3,5
3,8
4,0
4,5
4,0
4,5
72
_______________________________________________________________
. 3 (. sibirica, .mandshurica,
.vulgaris) (.2),
. , 26 .
.
, , .
(
, , ) ,
,
. 4, .
:
, , , .
- .
1. , .. / .. , .. .
.: , 2007. 188 .
2. , .. / ..
, ...
.. . / , / : - , 1999. .300-350.
3. , .. /.. // /
2002. 5. .32-35.
73
______________________________________________________________
634.71:631.527
..
, . ,
. , 4
e-mail: sorokopudov@bsu.edu.ru
-
,
- .
: , , , ,
- ,
. -
, .
.
[1 - 5].
-
.
,
.
2001-2006 .
22
, (, 1999 .).
.. , . .. , . : , .
,
1 . ,
, ( 5.6-6.0). 2,8 1 .
.
: , , ,
, .
:
, , ,
.
, .
2004-2005, 2005-2006 .
. III 2004
(-7), 2005 (-10,9) I 2004 (-10,0), 2005
(-18,2) (9-11 )
(2-15 ),
.
74
_______________________________________________________________
2005 2006 4,6-5,0
, , , , ,
.
(. 1).
1
,
2003
2004
20032004
20042005
20052006
20042005
20052006
1,0
1,2
1,2
1,1
1,4
1,6
1,0
1,3
1,0
1,0
3,0
1,0
3,0
2,7
2,0
3,6
3,0
2,8
5,0
3,8
1,5
3,0
3,0
3,5
5,0
4,6
1,5
2,5
3,0
2,4
4,3
3,7
1,0
1,0
3,5
1,5
3,0
3,6
1,6
3,0
3,4
2,8
5,0
3,2
2,0
3,0
3,0
4,0
5,0
5,0
1,5
2,3
3,3
2,8
4,3
3,9
2,0
2,4
3,0
2,5
1,0
1,6
4,0
2,2
1,8
1,0
1,3
2,7
1,0
1,2
3,2
2,0
1,2
4,0
2,5
5,0
2,8
2,6
1,8
3,2
2,5
5,0
2,5
5,0
2,8
3,0
1,2
2,6
1,8
3,3
2,1
4,2
2,2
2,3
2,1
2,6
1,6
2,0
1,0
2,3
1,3
1,0
3,0
1,6
1,4
3,4
2,5
5,0
3,0
2,8
2,4
3,8
3,0
5,0
3,3
5,0
3,4
3,2
2,0
3,5
2,0
3,5
3,3
4,1
2,6
2,3
2,5
3,0
2,8
5,0
5,0
4,3
3,2
4,4
5,0
4,2
1,4
3,0
1,0
1,2
2,6
2,6
5,0
3,5
5,0
2,5
3,0
2,9
3,0
2,3
1,2
1,6
3,0
4,8
5,0
3,4
5,0
3,2
3,0
2,7
.
:
(1,1-1,8 ) , , ; (2,1-2,4 )
, , , , ;
(2,5-3,3 ) , , , ,
, , ; (3,7-4,3 ) , ,
, .
20 31 4
61,3-188,0. . II
III (12-23 ), 14,5-16,8.
,
607,0-811,5.
:
15 18
1094-1125 , ; 19 22
1146-1160 , , ,
, , , , , , ,
, , , ; 25
1186 .
75
______________________________________________________________
.
.
19 . ,
28 . , , , .
125-198 .
(1,7 ) (2,0 ),
(1,9 ). , , 1,6
1,7 (. 2).
4,2 ,
4,0 , 3,8 , 3,5 , 3,5 , 3,4 .
2
( 2004-2006 .)
N (0.95)
,
/
,
%
,
%
, %
1,7
2,9
5,0
12,40
6,24
35,5
2,0
0,6
0,8
0,4
0,6
3,8
3,0
2,4
2,8
3,3
4,5
4,0
4,5
4,0
4,5
11,45
12,07
11,39
11,18
11,36
5,27
4,76
5,22
5,0
5,66
37,84
31,28
35,93
37,39
24,17
0,9
4,0
5,0
10,32
7,33
37,10
1,9
0,9
0,7
0,5
0,3
1,7
1,6
3,4
3,0
2,7
3,2
3,1
3,5
4,2
5,0
4,0
4,0
4,0
4,0
5,0
4,5
11,39
11,21
11,90
11,90
6,07
11,06
10,16
5,41
4,36
6,43
5,71
7,57
4,51
4,83
42,83
45,82
43,56
35,20
33,40
40,19
23,47
0,5
2,4
4,5
10,78
4,11
37,84
0,9
1,0
3,5
4,5
11,21
3,68
36,08
(4,5-5,0 )
, , , ,
, , , , , .
(6,07-12,40%) (3,68-7,57%).
945,82 %),
(43,56 %), (37,1 %), (40,19 %) (. 2).
, - - .
1. .. -
// . . . , 1982. 195 .
76
_______________________________________________________________
2. .., ..
. // -
. , 2003. .110-113
3. .., ..
// . . . ., 1977. . 10. - . 175-179.
4. .., .., ... -
// . :
. . . : , 2004. . 293-299.
5. .. . . //
. .-, 2005. . 132-143.
In the Southeast zone of the Kamchatka edge Illusion and the Imagination most adapted for local prirodno-environmental conditions
have allowed to recommend the conducted researches for introduction in
amateur gardening of a grade of a raspberry with a complex of economic
valuable signs.
Key words: a raspberry, grades, winter hardiness, productivity,
weight of a berry
77
______________________________________________________________
634.71:631.526
..
, . , . , 4
e-mail: sorokopudov@bsu.edu.ru
- ,
, , , , , , . .
: , , , , .
, , , . -
.
-
.
: , , , , , .
, , , , ,
.
,
.
, .
.
. , , . , . 80-90 . - , ,
(780-1119 )
(60-80%). , , +3,7. 3,7, - 26.
30-50 .
0 20-22 , 5 20-23 .
- . 120-160 . 1300-1600.
.
. .
- (70%). .
78
_______________________________________________________________
. 36 5 2001
2002 . , -
. - , , , . .. , .
.. , .
, , .
, (.
1999 ., . 1973 .). : ,
, , ,
.
(2004-2007 .) 13 23 . 28-35
14-23 .
3-11 .
:
( 29 4 ) , , , ; ( 5 8 )
, , , , , ;
( 9 13 ) , , , ; ( 14 20 ) , , .
, . - . , , , , . (20042007 .) . , , , , , , , , , , , , , , , ().
( 2004-2007 .)
,
/
, %
, %
, %
2001
(st)
(st)
N (0,95)
2,4
6,2
10,5
11,18
7,68
38,38
0,8
0
0,6
0,2
0,3
3,6
3,2
2,7
2,6
2,5
4,7
6,3
4,4
7,5
3,6
4,8
4,7
8,4
3,6
4,9
2002
12,02
14,27
10,95
12,33
11,34
6,56
9,05
7,73
8,32
7,73
51,04
53,31
51,04
51,50
56,32
0,9
6,0
9,8
12,21
8,29
48,67
0,3
0,4
0,3
3,2
1,6
1,5
1,0
2,7
4,6
6,0
3,7
8,9
8,2
14,49
10,42
11,10
8,61
6,70
6,02
44,88
36,52
59,39
79
______________________________________________________________
. . 2005, 2006 .
.
, , , , , , , ,
. (70-50%)
, . , . ,
15 20.
, , , , ,
, , , , , , , .
, .
2,2 0,6 3,7 (. 1). 0,6 1,3
: , , , , ,
, . 34-62-42. (1,5-2,7 ) , , , , .. 12-1-90, 25-1-90 .
(3,0-3,7 ) , , , , , .
2,0 6,2 . , , , , ,
, , , ,
12-1-90 (7,5-10,5 ). (2,0 ),
(2,4 ), (2,5 ), (2,7 ), (2,8 ).
(5 ) , , , , , , , 34-62-42.
10,22% () 14,49% (), 5,69% ()
11,08% ().
(57,84 %), (51,50 %), (56,32 %), (53,31 %), (51,04 %), (59,39 %),
(55,44 %) (. 1).
-
,
, , , , , , , .
1. . - .- , 1963. .49-50.
2. .., .. , 1999. 89 .
3. .., .. // .
., 1990. . 101-102.
80
_______________________________________________________________
4. .. ,
. // . /
. . . .-, 2007. . 153-161.
5. .. . // . .-, 2003. 20 .
81
______________________________________________________________
635. 168: 581.8
..
..
..
,
305041, . , . ., 3
e-mail:
kaf.farmakognoz@kurskmed.com
, ,
(Tragopogon orientalis L.), -11 Adobe Photoshop
10.0. .
: , , , , .
82
_______________________________________________________________
4-6 2-3 . , . 5-6 .
, ,
. .
, . , (. 1).
. ,
(. 2, 3).
(. 5).
(. 2, 3, 5).
1
2
3
4
7
5
6
8
2
1
9
10
11
. 2.
(. 600)
1
2
. 1.
(. 80)
1
2
3 ; 7
4 ;
8
5
9
6
10
11
1
1
2
. 3. (. 200)
1
. 4.
(. 600)
1
2
83
______________________________________________________________
.
, ,
(. 4, 6). (10 ) ,
(. 6).
, , ,
,
(. 4). , (. 7).
2
1
1
2
. 6. (. 300)
1
2
. 5. (. 300)
1
2
2
1
. 7.
(. 200)
1
. 8. (. 300)
1 ; 2
.
, ,
(. 8, 11). (. 8).
, (. 9, 10, 15). , (. 9).
. ,
.
84
_______________________________________________________________
1
1
2
. 10.
(. 300)
1
. 9.
(. 600)
1
2
:
(. 11, 12, 13);
-, -, (. 11, 13);
, 2-4 ,
(. 11, 14);
( 10 ) , , (. 15);
, , , (. 11, 12, 13).
4
5
. 11.
(. 300)
1
2
3
4
5
. 12.
(. 300)
1
2
3
85
______________________________________________________________
3
1
1
2
. 13. -
(. 300)
1
2
3
. 14.
(. 300)
1
. - .
, .
, .
(. 12). (. 16).
1
1
. 15.
(. 200)
1
. 16.
(. 300)
1
. . , .
(. 17). , (. 18) . , -
86
_______________________________________________________________
(. 19). (. 19).
, . (. 20).
1
2
. 17.
(. 600)
1
2
. 18. (. 600)
1
1
1
. 19. (. 300)
1
2
. 20. (. 600)
1
. , , .
1. . 11 . .: , 1987. . 1.
277 .
2. / . .. , .. . ., 2001. .130.
3. , .. / .. . , - , 1978. 116 .
4. . /
. . , . . ., , 2007. 768 .
87
______________________________________________________________
STUDY OF ANATOMY OF HERB TRAGOPOGON ORIENTALIS
V.N. Bubenchikova
Ya.S. Trembalya
S.A. Prokhorova
Kursk State Medical University,
K. Marks str., 3, Kursk,
305041, Russia.
e-mail: kaf.farmakognoz@kurskmed.com
88
_______________________________________________________________
582.579.2:581.522.4(234.81)
IRIS L.
..
..
..
, . ,
. , 85
e-mail: ostapenko_I@bsu.edu.ru,
sorokopudova@bsu.edu.ru
-
. . ,
,
, , . ,
,
[2]. 1400 12000 ,
, 80 , 500
[3].
Iris L.
. 25 Iris
,
, [1].
I. aphilla L. (. ), I. pumila L. (. ), I. pineticola Klok. (. ), I. pseudacorus L. (. ) [2]
, 2004 ., I. pumila , I. aphilla 2008 .
[4, 5].
, , , , ( , ).
. ,
, .
.
,
-- 1992 ., -
89
______________________________________________________________
[6].
.
I. aphilla I. pumila.
().
I. aphilla I. pumila 2003 .
I. pumila , I. aphilla . I. aphilla L. 2 (
), I. pumila - 3 ( ) [5]. 2008-2009 . , Iris, Iris .. [7-8].
I. aphilla I. pumila
( 20 . ), . 2 : 2007 .
2008 . , 40
.
- . 9-12 ,
25-28 .
0 225-237 , 148 153 .
467 , -
[9]. 2008-2009 . ( 2008 . 40 %,
2009 . 64 %), .
[10-11].
I. aphilla , ,
, .
I. aphilla ,
(, ; ; ), (
), ( , ) [5].
, ;
. , , . I. aphilla ,
. 17 24 .
I. pumila - . I. pumila
, , (; ), (,
) [5]. ( 1 3) , ().
6 9 . I. pumila ,
. I. aphilla I. pumila .
I. aphilla I. pumila , . - - ( 1), Bromopsis inermis (Leyss.) Holub, Poa pratensis L., Coronilla varia L., Fragia vesca L., Thalictrum minus L., Salvia verticillata L., Agrimonia eupatoria L., Chamaecytisus austriacus (L.) Link, Achillea millefo-
90
_______________________________________________________________
lium L., Leucanthemum vulgare Lam., Anemone sylvestris L., Clematis integrifolia L.,
Stachys annua L., Euphorbia waldsteinii Waldst. et Kit., Galium verum L., Hieracium pilosella L. Asteraceae Dumort., Ranunculaceae Juss., Fabaceae Lindl., Lamiaceae Lindl., Poaceae Barnchart. Rosaceae Juss.
-
( 2), Bromopsis
inermis (Leyss.) Holub, Agrimonia eupatoria L., Polygala comosa Schkuhr, Plantago lanseolata L., Saponaria officinalis L., Centaurea jacea L., Salvia nemorosa L. , Salvia pratensis L., Salvia verticillata L., Fragaria vesca L., Galium verum L., Stachys annua L., Euphorbia waldsteinii Waldst. et Kit., Lathyrus pratensis L., Coronilla varia L., Chamaecytisus austriacus (L.) Link. Lamiaceae Lindl., Fabaceae Lindl. Rosaceae Juss.
- ( 3), Calamagrostis epigeios (L.)
Roth, Genista tinctoria L., Fragaria vesca L., Polygala comosa Schkuhr, Agrimonia eupatoria
L., Centaurea jacea L., Plantago media L., Plantago lanseolata L., Veronica teucrium L. Plantaginaceae Juss. Rosaceae Juss.
2008-2009 2011 .
I. aphilla I. pumila, . 2008 .
, 2007 .
; 2009 ., 50 % . 2011 .
I. pumila I. aphilla
. , I. aphilla,
, 10 %. I. aphilla
. 100 %. I. pumila , ,
2009 . ,
(. 1).
1
I. aphilla I. pumila
(2008-2011 .)
1
2
3
2007 .
2008 .
2007 .
2008 .
2007 .
2008 .
2007 .
2008 .
%
I. aphilla
I. pumila
2008
2009
2011
2008
2009
2011
100
90
30
100
90
90
80
50
0
80
50
40
100
100
90
100
90
90
90
30
10
80
50
50
100
100
100
90
90
90
90
20
0
80
60
40
100,0
96,7
73,3
96,7
90,0
90,0
86,7
33,3
3,3
80,0
53,3
43,3
I. aphilla I. pumila .
2009 .
1,46, 3
(. 2). .
91
______________________________________________________________
2
( 2009 .)
-
-
2007 .
I. aphilla
I. pumila
1,10 0,07
1,04 0,04
2008 .
1,08 0,15
1,20 0,08
2007 .
1,20 0,09
1,25 0,09
2008 .
1,04 0,04
1,04 0,04
2007 .
1,32 0,08
1,46 0,11
2008 .
1,17 0,07
1,30 0,10
1,15 0,08
1,22 0,08
2011 .
I. aphilla I. pumila . , 2007 ., , 2008 ., ( ), , ().
2 (
) -
I. aphilla, I. pumila.
. I. aphilla
.
- . I. pumila
, (. .)
, , (
) .
, , . 2-3 .
, , . I. pumila . . , -
- . I. aphilla
. 1-2 , . I. aphilla . .
I. aphilla I. pumila
, , 2
, , 2 .
.
.
, .
92
_______________________________________________________________
1. .. Iris L. (Iridaceae) // Turczaninowia. 2008. . 11, 2.
. . 5-68.
2. .., .., ..
( ). .: , 2004. 120 .
3. . , , . / . .. . , 2004. 532 .
4. () / . - . .. ; . . -; . . . - . .
-.: . . . . .. . .: , 1988. 590 .
5. ( ) / ; ;
; ; . .. ; . .: .. .; . ..
. .: , 2008. 855 .
6. . .: , , 1994.
62 .
7. .., .., .. . / . . .. . .: , 1981. 156 .
8. .. . .: , . -, 1988. 159 .
9. . .:, 1972. 92 .
10. . . 2. / . . .. , .. . .; .:
- , 1960. 499 .
11. .. . 2- . .: - , 1970. 367 .
93
93
_______________________________________________________________
. I. aphilla I. pumila :
, , .
:
1, ,
- 2, ,
- 2, ,
2, ,
- 3, ,
- 3, ,
- .
94
_______________________________________________________________
635.9:582.579.2
..
, 393760,
,
. ,
. , . 101
e-mail - info@mgau.ru
,
.
( ) -
.
: , , , ,
.
, . , . , 10000,
, - , .
( ) (
) (). .
,
.
.
2007 .
. . . , . 7 : , , , , , ,
2 ( ) . . . .
10 : I ( 3,2 ), II (2,5-3,19 ) III (1,4-2,49 ) .
, . . . 1972 .
, (
+ (m1)), ()
(m2), m2/m1 ( ), , (d.) (d.) , ( ), (h) (d/h), (d) (d/d., ).
95
______________________________________________________________
. 1 2. 1, , , ( II III )
( III ). ( ) .
1
I
II
III
05
I
II
III
05
I
II
III
05
I
II
III
05
I
II
III
05
I
II
III
05
I
II
III
05
I
05
(
II
-)
III
-
d- d.,
%
0,92
32,09
0,7
35,67
0,43
33,25
30,38
1,27
57,78
0,62
32,75
0,43
36,32
86,2
0,83
29,83
0,5
22,69
0,6
49,91
41,81
1,17
46,57
0,97
41,43
0,57
34,8
30,58
0,73
14,14
0,48
21,58
0,4
21,04
13,79
1,03
40,24
1,03
62,33
0,5
30
45,02
0,77
30,97
1
64,77
0,3
16,36
59,2
h,
d/h
d,
d/
d, %
1,8
1,67
1,23
2,03
1,73
1,47
2,1
1,77
1,4
1,7
1,5
1,37
2,0
1,72
1,37
2,37
1,83
1,47
1,83
1,67
1,33
-
2,1
1,61
1,46
0,31
1,72
1,58
1,32
0,2
1,74
1,65
1,40
0,16
2,01
1,88
1,54
0,34
1,74
1,62
1,71
0,29
1,52
1,49
1,55
0,2
1,8
1,64
1,6
0,23
1,23
0,8
0,48
0,97
0,73
0,43
1,02
0,7
0,46
0,65
0,6
0,5
0,83
0,65
0,68
0,77
0,73
0,65
0,63
0,6
0,57
-
32,88
30,16
26,96
8,31
27,63
26,97
22,36
6,71
28,0
25,86
24,03
7,61
18,99
20.28
23,2
6,69
26,08
23,65
29,29
7,12
21,48
26,92
28,67
3,86
19,23
22,04
26,99
7,61
22,29
0,19
0,22
0,12
6,45
33,3
0,18
0,22
0,22
6,34
43,32
0,14
0.16
0,21
7,24
m1
m2
m2/m1,%
d.
23,33
20,67
17,67
15,67
15
11
2,39
16,33
10,0
8,0
3,89
14,67
8,33
7
3,34
15,0
11,33
7,67
5,64
11,0
9,33
7
5,53
14,33
14,33
8,67
3,03
15
10,67
7
1,2
67,37
72,78
62,2
5,25
73,05
60,12
58,09
20,88
73,38
60,98
53,91
11,45
73,54
70,34
49,99
32,71
75,69
55,79
51,69
32,46
66,2
78,56
68,65
15,16
74,97
62,85
67,57
11,82
2,87
1,97
1,37
2,23
2,12
1,5
2,83
2,2
1,37
2,3
1,9
1,53
2,93
2,28
1,93
2,57
1,7
1,77
2,53
1,73
1,83
-
3,78
2,67
1,8
3,5
2,73
1,93
3,67
2,7
1,97
3,43
2,9
2,03
3,5
2,77
2,33
3,6
2,73
2,27
3,3
2,73
2,13
-
4,82
4,09
20,97
0,61
5,47
4,25
23,21
0,34
4,88
2,01
21,3
0,21
22,33
16,67
14,67
20
14
13
20,7
17,33
16,0
23,3
16,67
14
21,67
18,33
12,67
20
17
10,33
.
( 78,56% II ), . ,
,
.
I
(3,78 ). II III
(2,67 1,8 ).
. II
(64,77%) (62,33%), I (57,78%)
(46,57%).
( 14,14-21,58%). ,
. , ,
96
_______________________________________________________________
- ,
. , . , , .
I
2,37,
(. .). (d/h)
,
(2,1) (1,52) ( I ). () :
1. ( d/h 1,52
).
2. (d/h = 1,72-1,8 ,
, , )
3. (d/h = 2,01-2,1
).
2
1
2
3
I
II
13,67
9,33
3,33
12,39
12,67
7,33
4,67
6,6
8
4,33
3
2,93
7,67
4
4
4,04
5,3
5
3
4,51
6,67
4,67
4
3,67
11,67
7,33
4,67
4,0
7,75
3,41
0,6
0,63
0,53
0,63
0,62
0,67
0,77
0,65
0,53
0,77
0,63
0,5
0,7
0,77
0,67
0,9
0,7
0,6
0,73
0,67
0,57
0,15
0,17
III
2,69
0,13
05
1
2
3
05
1
2
3
05
1
2
3
05
1
2
3
05
1
2
3
05
05
05
( , )
1
2
3
%
9
64,44
5,33
56,93
1,33
41,11
17,84
8,67
67,18
3,33
49,05
2
36,08
20,61
5,33
66,67
2,33
53,33
1,33
44,44
28,09
4,33
56,11
2
64,29
2,33
57,94
61,66
3,33
71,85
2,67
50,16
1,33
47,22
55,18
5,33
77,74
2,67
60
2,67
74,44
86,62
6
50,77
4,33
58,99
2,33
50
22,3
18,27
25,19
-
43,01
: , 2, , 05 .
97
______________________________________________________________
II 1 1,49, 2 1,58-1,65, 3
1,88. III : 1 1,32-1,46,
2 1,54-1,6, 3 1,71. ,
, , .
. d/h , . ,
,
.
,
, , -
( ).
(32,88% I - . . 1). , ( I
1,23 ), , , .
()
(d/h = 2,1 ). , , , , I (
3,2 4,5 ), ,
. ,
.
(. .).
, ( ),
(, , , ) ( ). -
98
_______________________________________________________________
, , .
( 2). (
) I
13,67. I .
( 0,9 ).
( 70 % I III ).
1. ,
, : , ,
.
2. (d/h)
() : ,
. ,
, ,
.
3. I , . , , 0,9 ,
0,5 I 77,74%.
1. . . . .: , 1981. 192 .
2. . ., . ., . . . .: +, 2002.
3. . . . .:
1972. 36 .
Gain of corms - the important parameter which reflects opportunities of a grade in the shortest terms to receive a large and qualitative
planting material. The relation of corm bottom diameter by the diameter
of corms (in percentage) - the main indicator of ageing of a planting material has been calculated. On various biometric parameters mathematical models of corms various grades of a gladiolus are made.
Key words: gladiolus, cultivar, gain, corm, diameteranatomy.
99
______________________________________________________________
579.81.:633.31(571.56 191.2)
..
, 677980,
. , . 41
e-mail: mila_atlasova@mail.ru
. .
: , , ,
, , ,
.
, (, 1989).
, ,
(,
, 1981).
.
. , , .
. , ,
.
, ,
.
.
- () .
- , .
.
:
1. - () .
2.
3.
.
. -
100
_______________________________________________________________
10 15 .
, 50 60 .
.
,
. ,
. , - 3-5 [2].
,
. . .. .
,
,
(. .). , 202030 0,25 .
2,0 2,5 .
0 20
,
/
N60P80K80
/
N60P80K80
/
N60P80K80
/
N60P80K80
- /.
137
100
,
,
142
125
,
109
49
69
34
181
164
76
68
-
/
8.22
6.00
4,94
3,57
,
/
960
3240
8.52
7.50
5,12
4,46
660
2910
6.54
4.14
10.86
4.56
2.94
2.04
9.84
4.08
3,9
2,49
6,53
2,74
1,75
1,21
5,86
2,43
1092
1548
2400
1020
. . . . ,
.
,
, ,
[1].
, , ( )
. , ,
.
. . .
. 1
3 [1].
, 0 10 . ,
, . , -
101
______________________________________________________________
. ( )
10.86 .
/, 9.84 . / (.).
.
( )
25 % .
N60P80K80 .
, .
, . . .
.. ( ), 1 ,
. () ,
()
. () , , 1 [1].
,
102
_______________________________________________________________
., .
. (6,53 /) .
, -, , .
.
, N60P80K80
.
1. .., .. .
.: , 1983. 256 .
2. .. . -: , 1995. 224 .
3. .., . .
//-. , 1981. . 16. 3. . 349 358.
4. .. / .
, 1989. 109 .
103
______________________________________________________________
633.28:581.4
.
.
: , , , , .
(, 1971; , 1989).
Psathyrostachys caespitosa . (Psathyrostachys caespitosa (Sukaczev)
Peschkova) .
,
[1]. (- , ) [2].
[3]. , [4, 5].
- , ,
, [6, 2]. ,
.
, .
,
.
, , ..
.
,
, ,
[2].
2 . Psathyrostachys caespitosa , , .
, .
104
_______________________________________________________________
2003 .
3030
. - Psathyrostachys caespitosa, : ( ) (,
).
- [7,10,11].
: , , , .
, [8,9]. .
, , ,
.
1-2 . , , , ,
, .
3030 ( ).
. , 2003-2004 .
.
P. Caespitosa . 1 . 1.
10-12 . 0,2-1,7 1-3 .
1- 2,0-6,5 0,2 . 18-20
2- . .
. , 2 ( ).
3-
II1 II2.
4 (2 , 1-2 , 1 ).
8-9, ().
II1 1-
, 4-
5- (.. 4-5- ) .
II1 2- () .
1 (II1) 2 (. 1).
5- II2. 3-5.
3-4 (1-2 , 1 1 ). 10-11. III
II1. II1 2 .
105
______________________________________________________________
5
2
4
1
6
5
7
5
2
1
1
4
11
8
II
1
2
10
9
8
64
97
5
3
II
III
105
. 1. Psathyrostachys caespitosa :
- 3- ; - 5- ; - 6- ; - 7- ; - 9- ; - 12-
( ); I - ; 1-12 - ; - II- ; - III-
106
106
_______________________________________________________________
IV2-1-1
3-4
5
IV1-1-2
3
3
1
1
2
2
IV1-1-1
4
5
10
III4-2
4
5
III4-1
5
5
III3-2
6
6
III3-1
6
7-8
III2-3
40-50
50-60
III2-2
11
12
11
12
13
14
15
16
17
18
19
20
21
22
III 2-1
1
2
III1-3
1
1-2
3
III1-2
1
2
4
1
1-2
2
2
3
III1-1
1
3
3
4
II6
1
2
3
4
5
6
II5
1
2
3
4
5
5
6
II4
1
1
1
2
3
4
6
10-12
15
15-20
II3
1
2
3
4
5
6
7
8
9
10
II2
II1
1
1
1-2
1
1
2
1
1-2
3
2
2-3
2
3
2
3
2
4
3
1
1
1
1
3
1
2
1-2
1
4
4
3
3
2
2
1
2
2
2
1
2
3
5
8
12
17-18
30-35
35-40
55-60
1
2
80-100
100-120
107
______________________________________________________________
6- 3-4 ( 2-3 II
) 5-6 . 4 .
II1 3 , II2 1-2 (. 1).
7- (7- ) II1 4 , .
III III1-1, 1-
II1 1- . 7- 4- , II (II2, II3 ..).
II 2-4 ,
III 1 . III (III1-1 III1-2)
1-2 . 5-13 ,
II
6-7 . 13
. , 7-
3- II , 1-2 III , 3-11 .
IV III1-1.
II
. II2, 3- ,
2- 5- ,
II3, II4, II5 6, 7 8- . II :
2-3-
II (. 1).
2-3, 3-4- 4-5- II ,
1, 2 3 .
III II
,
II-III . , ,
4-5- .
9- II
7-8, III 5. , II ,
3-4- , . III
IV 3-4 .
17-18. II 4-5-, III 3- (. 1).
Psathyrostachys caespitosa 9-10
. 2- 12- (. 1). ,
9-10 . 15-20,
.
, Psathyrostachys caespitosa
:
1. II1
1- , 4- 5- . ,
.
2. ,
15-20. 1- 9-10 . , , 4-5- .
108
_______________________________________________________________
, ( II
) 4-
5- , ( III )
7- .
1. .., .., .. .
. : , 2005. 320 .
2. .. . /
. . -. . . , 2006. 240 .
3. ..
// . -:
, 1991. 185 .
4. .. () . -: ,
1975. 71 .
5. .. . :
. -, 1977. 110 .
6. .. . :
. , 1988. 172 .
7. .. . -: , 1995. - 225 .
8. .. . .:
, 1971. 357 .
9. .. . ., 1973. 225 .10. .. // . . . 3. . .; .: - , 1950. . 6.
. 7-204.
11. ( ) / .., .., .. . .: , 1976. 217 .
There are set out results of investigations of the tillering and bunch
formation of Psathyrostachys caespitosa have been presented in ontogenesis. The law-governed nature of simple and complex bunch has been
established depending on the growth and development of leaves of plant
main shoot.
Key words: morphogenesis, tillering, organogenesis stages, morphostrukture, ontogenesis.
109
______________________________________________________________
635.9:582.579.2
..
, 393760,
,
. ,
. , . 101
e-mail - info@mgau.ru
8 , . k, (r)
. .
: , , , , , ,
. ,
.
. -, .
. . ,
(, , , , 1994). 4 ( , ,
) ,
:
S = L W 0,62,
L , ,
W - , .
0,62 (k)
, ,
, , , k=0,62
. , k
.
. - , , , . , , .
- . 5-9, 10-12 , . 5-6
, . 2-3, 4 .
, - ( ).
,
.
110
_______________________________________________________________
2008 8
, . , 7 ( ()
- ). (
1 ). , ,
k :
k = (S. / S..) 100%
S. , 2;
S.. ( ) , 2.
, , .
, I (. 1), 3-
( , , 4- ,
,
2-). II ,
: 5 6 2- , 4-. , III
( ) 2-
, ( ) 1-.
, (I ) 5 ( , ), 6 ( , ). II : 4
, , 5
: . , III , (
4 : , , ).
344,53 2 ( I ). II III S
(137,03 74,47 2 ).
(r)
,
. 1 , ( 0,92 0,999).
( II - 0,864) III
(0,766), .
( , , , ) 66,4 I 4- (. 2).
( I ). ,
3- . II 5 7 3- ( (l = 48,3 ) ).
, 2- . III 2- ( (42,9
)).
111
_______________________________________________________________
1
(S), 2
3,67
(S), 2
(r)
13,87
73,83
94,7
81,97
60,8
16,3
344,53
0,972
0,972
3,63
2,77
1,97
I
II
III
47,47
20,07
24,07
59,43
43,97
26,9
53,8
38,83
8,0
37,37
18,07
-
20,73
-
213,97
120,93
58,97
0,998
0,999
0,944
0,974
2,7
II
25,8
36,63
35,0
24,87
16,9
127,93
0,998
0,998
3,33
2,83
2,07
3,53
2,83
2,7
2,17
3,6
3,3
2,8
2,4
I
II
III
I
II
II
III
I
I
II
III
20,23
22,65
20,4
34,37
26,47
38,4
29,33
37,4
27,27
10,57
20,07
34,27
43,2
25,37
58,43
43,87
38,03
27,73
43,27
37,73
29,07
27,4
53,03
35,2
7,0
69,2
17,07
35,0
17,4
50,77
44,8
40,37
5,67
37,17
23,2
63,87
10,73
25,6
51,57
37,17
24,0
-
9,6
26,13
44,33
18,97
8,43
-
35,8
22,17
-
144,27
112,65
50,43
264,17
94,67
137,03
74,47
249,5
166,07
112,43
53,13
0,947
0,864
0,911
0,969
0,953
0,999
0,766
0,98
0,934
0,948
0,999
0,92
0,943
0,989
0,98
0,989
(d),
111
_______________________________________________________________
112
112
(d),
3,67
17,57
45,73
63,1
66,4
60,6
30,03
1,2
2,4
2,23
1,83
1,47
0,77
0,645
0,67
0,672
0,674
0,692
0,708
3,63
2,77
1,97
I
II
III
35,4
24,5
27,33
46,77
41,8
42,93
48,13
48,3
23,03
41,2
35,47
-
23,97
-
2,07
1,17
1,5
1,93
1,57
0,9
1,7
1,2
0,47
1,23
0,77
-
0,97
-
0,647
0,62
0,586
0,659
0,666
0,694
0,670
0,679
0,751
0,715
0,681
-
0,741
-
2,7
II
26,37
37,7
44,57
41,23
31,8
1,33
1,43
1,2
0,87
0,75
0,736
0,682
0,655
0,687
3,33
2,83
2,07
3,53
2,83
2,7
2,17
3,6
3,3
2,8
2,4
I
II
III
I
II
II
III
I
I
II
III
22,33
32,6
27,2
25,0
28,6
32,23
34,33
32,47
26,93
26,93
25,27
33,53
44,0
38,23
39,1
37,23
38,2
39,63
37,8
33,93
39,87
38,1
48,0
45,8
22,15
46,03
19,37
38,13
39,13
46,0
43,7
47,03
18,07
49,57
45,3
45,53
13,37
34,17
44,63
45,6
28,03
-
30,2
27,83
43,9
36,33
12,83
-
31,7
29,87
1,63
1,1
0,98
2,17
1,5
2,1
1,37
1,77
1,57
0,63
1,23
1,67
1,475
0,97
2,43
1,87
1,8
1,13
1,77
1,77
1,17
1,1
1,6
1,2
0,4
2,4
1,37
1,33
0,7
1,67
1,6
1,3
0,43
1,13
0,8
2,37
1,13
1,03
1,73
1,2
1,27
-
0,5
1,32
1,43
0,73
0,9
-
1,9
0,93
-
0,557
0,655
0,767
0,613
0,62
0,565
0,632
0,653
0,653
0,631
0,646
0,627
0,672
0,688
0,628
0,629
0,552
0,617
0,650
0,633
0,625
0,657
0,694
0,641
0,625
0,637
0,693
0,641
0,663
0,646
0,662
0,770
0,654
0,65
0,581
0,716
0,737
0,671
0,682
0,676
-
0,635
0,63
0,71
0,716
0,698
-
0,797
-
113
______________________________________________________________
I II
2- . ( I II )
( I ), 1- ,
( II ) 3-
. III
1- .
k . 2. , .
, , , k , , 2- . , , , k, . ,
(, 4-6-), , (1- 2- ).
, ,
S = L W k.
1. , I ,
5-6 , II 4-5, III 3.
2.
(
(r) 0,766 0,999).
3. ,
S = L x W x k.
.
, .
1. . . . .: , 1981. 192 .
2. . ., . ., . . . .: +, 2002.
3. . ., . ., . .
. // . . . .- : ,
2005. . 503-509.
4. . ., . ., . .
. . .: , 1997. 144 .
5. . . . .:
1972. 36 .
6. G. Barbieri, S. De Pascale, R. dAndria. M. Mori. Estimating Gladiolus leaf area by linear
measurements // Advance in Horticultural science, 8, 1994, . 85-89.
114
_______________________________________________________________
635.9:582.579.2
..
..-.
.
..
, 87 .
7 , 19 , 37 , , 24 , 11
locus classicus .
: , , .
, . ,
, .
, .
(
, .,)
, , , , .
: - [1; 2; 3; 4; 5; 6 .].
.
[7].
. ,
. . , , . [8].
.
[8; 9; 10; 11
], . ,
, , . ( , , , ..) , , ,
. . .
115
______________________________________________________________
. .
.
, , , ,
, , , , .
[12]. -
( ) . (-, , )
.
.
,
.
, .
. .
, . ,
, . ,
. , , . , . ,
. - - .
, , ,
.
.
- ,
.
, ,
, . ,
, , , , , ( ), , .
.
Stipa, Festuca, Koeleria, Teucrium, Scutellaria, Botriochloa, .
( 1500 )
( 1000-1500 )
- . - .
-
116
_______________________________________________________________
. . .
,
, ,
. ( 2500 )
- .
- ( ) ,
[13;14;15;16]
. - .
,
, . , ,
- .
Colutea orientalis ( , ); Spiraea
hypericifolia; Tragacantha aurea, Tragacantha denudata, Paliurus spina-christi, Rhamnus pallasii, Berberis vulgaris. - Cotinus coggygria Celtis glabrata. . , , ,
, Ephedra procera, Cerasus incana, Cotoneaster meyeri, . integerrima, Crataegus monogyna, Juniperus oblonga, Amelanchier ovalis, Clematis
integrifolia, Rosa valentinae, Cerasus incana, Rhamnus pallasii .).
, Festuca brunescens, Elytrigia
gracillima, Carex tomentosa, C. pallescens, Allium fuscoviolaceum, A. paniculatum, A. albidum, Herniaria besseri, Dianthus capitatus, D. ruprechtii, D. fragrans, Alyssum hirsutum, Erysimum repandum, Potentilla pimpinelloides, Potentilla recta, Ononis pusila, Coronilla coronata, Hypericum hirsutum, Helianthemum ciscaucasicum, Vinca herbaceae, Dracocephalum austriacum, Phlomis tuberosa, Salvia tesquicola, Galium aureum, Campanula
hohenackeri, Aster amelloides, Achillea millefolium, Reichardia glauca, Salvia
daghestanica, Astragalus haesitabundus, Thymus dagestanicus, Xanthobrychis bobrovii
- , Medicago daghestanica ( ), Convolvulus lineatus, Linum
orientale, Salvia kuznetzovii, Vincetoxicum funebre, Teucrium chamaedrys, Onosma armeniacum, Dianthus cretaceus, Muscari pallens, Veronica propinqua. Teucrium polium, Galium ruthenicum, Botriochloa ischaemum , Stipa capillata, Koeleria gracilis, Festuca valesiaca, Medicago falcata, Chondrilla juncea, Herniaria
incana, Cleome daghestanica, Scleranthus annuus, Euphorbia szovitsii.
Parietaria judaica, Scrophularia rupestris, Galium brachyphyllunu
Onosma caucasica, Campanula sarmatica, Rumex hastifolius . Rosa .
. ,
. Tragacantha denudata. , , Fumana procumbens, Stipa caucasica, Psephellus
prokhanovii, Hypericum asperuloides, Symphyandra pendula, Gypsophila tenuifolia.
, , .
. .
117
______________________________________________________________
. Querqus petraea, Tilia caucasica,
Fraxinus excelsior, Acer platanoides, A. campestre, Carpinus caucasica, Fagus orientalis, Albovia tripartita, Doronicum macrophyllum,
Convallaria transcaucasica, Euphorbia macroceras, Polygonatum glaberrimum, P. verticillatum, Viola odorata, Lilium monadelphum.
, - ,
- ,
.
, .
.
-
. . , , Fraxinus exelsior, Quercus petraea, Tilia caucasica, Acer campestre, Ulmus
suberosa . Euonimus europaea Swida
australis. Hippophae rhamnoides. ,
Rosa.
. - ,
. .
. - .
. .
,
. :
.
, ,
1420-1625 . ,
. , . . , . . . (
, ), ( . ),
- .
, .
.
. . , .
, . , . .
(, . -, . ),
( ).
. , .
, -
-- Paliurus spina-christi, Berberis vulgaris,
Cotoneaster meyeri, Rhamnus pallasii, Spiraea hypericifolia, Rosa canina, Crataegus
118
_______________________________________________________________
monogyna, Quercus robur.
. . (1974), Colutea orientalis, Cotinus coggygria, Celtis
glabrata, Ephedra procera, Cerasus incana, Cotoneaster integerrimus.
.
, -, . 750-850 .
,
. -
Spiraea hypericifolia
Rhamnus pallasii, Berberis vulgaris, .
-
.
- Cotinus coggygria,
. ,
, .
1400
Tragacantha denudata.
[17]. (Tragacantha aurea)
. , . - ( ) 1400 . ,
. ( . , .).
(, , ). Festuca varia Bromopsis variegata;
Calamagrostis arundinacea Phleum montanum. Betonica macrantha, Pyrethrum coccineum, Inula orientalis, Heracleum
asperum, Anemone fasciculata, Pulsatilla albana, Trollius ranunculinus, Scabiosa
caucasica, Cephallaria gigantea, Ranunculus caucasicus, Primula macrocalyx,
Polygonum carneum .
Rhododendron luteum, [18] . . (Betula
raddeana, B. litwinowii, B. pendula), Sorbus aucuparia. ( . )
- ( . ),
, -.
Rhododendron caucasicum.
, -, - - .
- (
), -,
, .
, . .
.
,
. . -
119
______________________________________________________________
[19], , ( ), , . , ..
(1974) , , , , Red Data Book - [19]. , , , , , , ,
locus classicus ,
, . 87. , :
;
;
, , ;
, ,
.
, . , , Amberboa glauca, Campanula ossetica,
Caragana grandiflora, Cerasus incana, Colutea orientalis, Gentiana grossheimii, Medicago daghestanica, Rhaponticum pulchrum , ,
. (Dentaria bipinnata, Primula bayernii,
Saxifraga columnaris), ,
.
,
.
, , Anacamptis pyramidalis, Betula
raddeana, Fritillaria orientalis, Orchis coriophora, Orchis ustulata, Petrocoma hoefftiana,
Saxifraga columnaris, Stipa pinnata, Stipa pulcherrima, Traunsteinera sphaerica , .. .
7 (Isatis pseudoararatica, Jurinea annae, Primula
darialica, Primula zeylamica, Psephellus pseudoandinus, Rosa valentinae, Saxifraga charadzae). . ,
. ,
.
, , , . , Botrhiochloa caucasica, Iris colchica, Petrocoma hoefftiana, Pulsatilla andina, Clematis integrifolia, Saxifraga columnaris, Medicago daghestanica, Caragana
grandiflora, Omphalodes rupestris, Rhaponticum pulchrum, Psephellus prokhanovii
. 37 .
,
, : Aetheopappus
pulcherrimus, Allium oreophilum, Aquilegia caucasica, Botrychium lunaria, Dentaria bipinnata, Fritillaria orientalis, Orchis sanasunitensis, Orchis triphylla, Primula bayernii,
Rhamnus depressa, Saxifraga columnaris 24 . ,
, , ,
120
_______________________________________________________________
.
, , 19 (Adoxa moschatellina, Amberboa glauca, Artemisia
salsoloides, Campanula ossetica, Caragana grandiflora, Cephallaria balkharica, Cephallaria dagestanica, Gadelia lactiflora, Gentiana grossheimii, Medicago daghestanica, Omphalodes rupestris, Onosma armeniaca, Psephellus prokhanovii, Rhamnus depressa,
Rhaponticum pulchrum, Rhododendron caucasicum, Tephroseris subfloccosus).
, , .
. , .
: , , . .
, . .. [20] . ,
Rhododendron caucasicum Betula pendula, Pentaphilloides
fruticosa.
.
11 , locus classicus : Campanula argunensis, Isatis pseudoararatica,
Mandenovia komarovii, Merendera ghalghana, Potentilla ghalghana, Primula darialica,
Primula zeylamica, Psephellus prokhanovii, Rosa valentinae, Saxifraga charadzae, Scutellaria leptostegia, Sedum argunense, Taraxacum ghalganum.
, 87 , . 7 , 19 , 37 , , 24
, 11 locus classicus .
, c,
. ,
.
1. .. . .1. //
-. 1894. . 27. . 123-332.
2. . . , ,
// . . .
. ., 1894. . 28. 267 .
3. .. - . . , 1923. 23 .
4. .. - . .-.: - , 1935.
108 .
5. .. - // : . / . .. . - , 1974.
. 5-22.
6. .. -. : . -, 1975. 117 .
7. ..
// . . . .1, .3, 1975. . 201-209.
8. .. . .: - , 1948. 267 .
9. .. // : .
. .: - , 1960. . 5. . 115126.
121
______________________________________________________________
10. .. ,
// . .5. .; ., 1960. . 115126.
11. .. - //
, 1891. 190 .
12. .. - // .-. . .24, .1, 1909. 174 .
13. .. . - . - .: - , 1953. 396 .
14. .., .. // .: . .2. , 1960. . 8-88.
15. . . //
, . 2. , 1956. . 93-105.
26. .. . . . 46, .6-7, 1910. . 213-280.
In article data on wood foothill flora of East caucasus, and also data
on 87 kinds needing protection are resulted. Among them 7 rare plants,
19 relicts, 37 kinds for which in this area there passes area border, 24 are
global rare species, 11 kinds with locus classicus in investigated area.
Key words: East caucasus, relicts, rare plants.
122
_______________________________________________________________
634.75; 631.526(571.56)
.. 1
.. 1
.. 2
1
,
677001, ., ., 23/1
e-mail: agronii@mail.ru
, 308015, . ,
. , 85
e-mail: sorokopudov@bsu.edu.ru
2
.
, ,
, , .
,
[1, 3]. , ,
, , .
- , .
75 . .
- , -, , (3,5%).
, (1999).
123
______________________________________________________________
1
Fragaria orientalis Los.
(. I, II, III,
. ,. ,
. )
(. ,. I, II)
-
(. )
-
(. ,
. )
(. )
(. )
(. )
, t
124-130
10
1200-1400
200-250
120-130
10,212,2
1400-1600
200-220
115-120
9,211,2
1000-1400
210-280
124-130
5,76,7
1400-1600
240-380
123
124
_______________________________________________________________
6 - . , (), .
.
+10o 90-100 , 1200-1400o.. 124-130 .
65 151 . 200250 , 90-130 .
. . +10o
80-90 .
1000-14000. 115-120 . 100-129 .
. 210-280 .
, - 6 .
. - . +10o , 1400 1600o. 120-130 , 65-90 .
200-220 , 105-160 .
-
. - - . ,
.
124-130 . 78-103 . +10o 1400 1600o.
240 380 . (195-200 ).
.. [2], 20 , ,
, , ; , - .
, , . , . :
(7-8),
2 . .
.. [2], 20 , ,
, , ; , - .
, , . , .
:
(7-8), 2 .
.
125
______________________________________________________________
1- ,
2-5 ,
. 30, 100 .
,
11-20 , -, -
18-23 .
, - (, , -, ).
,
, (, , ), (), ,
, , .
, (
7,0:4,2 ).
, . (, ), (),
, . .
().
.
. ,
(, , ), (), , , , -;
(,), (), - ()
- () - . - () , .
:
, .
.
(, , ),
- ; (, ), ,
, - () ;
- () . - :
, -, . -
3,7, - () 1,4 .
.
(), ,
(, , ) ;
(), - (, ), () ; - .
, . (, ), () , .
, , , .
. . . , .
126
_______________________________________________________________
, ,
, .
1. .. . ., 1985. 272 .
2. .. - , 1950. 350 .
3. .. . ., 1984 278 .
e-mail: agronii@mail.ru
2 Botanic
e-mail: sorokopudov@bsu.edu.ru
127
______________________________________________________________
634.75; 631.526(571.56)
FRAGARIA ORIENTALIS LOS.
..
..
,
677001, .,
.-, 23/1
e-mail: agronii@mail.ru
128
_______________________________________________________________
43,20 ().
+100 90-100 ,
1200-14000. +18.+190,
.
100. 124-130 . 65 151 (. 1).
.. [3] () .
11 1 , 2-6 , 8 12 , 7 27 .
5 48,0-195,1,
185,1-272,5. 10 171,8-740,4,
637,2-1090,7.
.
,
, .
. 38,20-40,90; +16-180; 9,2-11,10. +100 80-90 . 1000-14000. 115-120 .
100-129 .
. 210-306 .
() 11-45 ,
172-176 . 20-22 , 5 50, 10 190.
41.
, - 6 .
. - . , , , , . : . , . +17+180,
10,212,20. +100 , 1400 16000.
120-130 , 65-90 . 200-220 , 105-160 .
-
().
5 141, 271, 684, 19, 29,
70. ,
.
4-5 , 60-62 .
129
______________________________________________________________
1
Fragaria orientalis Los.
( )
,
/
, o
151
2102
-63
261
-10,2
151
-8,6
151
-10,2
151
-11,1
150
-11,1
150
-
-10,3
150
-
-11,6
-9,8
129
-11,1
-6,7
-
2102
2102
2102
-63
-59
-63
261
304
261
-65
-65
268
268
2294
-62
251
-66
-
262
262
2091
-
-63
-65
306
272
1812
-59
327
, o
116
F. orientalis Los.
.
.
147
F. orientalis Los.
147
F. orientalis Los.
.
.
140
F. orientalis Los.
100
210
F. orientalis Los.
223
F. orientalis Los.
-10,2
.
I, II, III
.
.
.
129
130
_______________________________________________________________
-
. - - . ,
.
30,1033,90. +17,50+18,70.
5,706,70. 124-130 .
78-103 . +100 1400 16000. ,
. , 6043 ..
240 380 . (195-200 ). 180 , ..
.
.
, Fragaria orientalis Los. - . , - - . - - .
, ,
- .
,
, , , , . .
, , .
,
(. 2). 2009 . - (82,7 /), (65,4 /) , (. 2).
(2010 .) (157,2 /), (143,3 /), (134,8 /) - ()
(134,8 /) . - , 2010 . .
2
, 2009-2010 .
, .
, .
, .
, .
, .
- , .
, .
, 1
2009 .
2010 .
28,2
77,8
53,0
33,8
64,3
49,0
26,2
88,1
57,1
30,5
96,2
63,3
44,3
157,2
100,7
65,4
143,3
104,3
42,5
134,8
88,6
27,2
118,4
72,8
16,7
99,8
58,2
131
______________________________________________________________
, .
- , .
, .
- , .
, .
, .
18,2
45,6
17,4
82,7
26,9
18,8
121,1
133,4
78,0
121,6
47,2
61,2
69,5
89,5
47,7
102,1
37,0
40,0
1. Fragaria
orientalis Los. - : , - ; - - ;
, , - .
2.
, - ,
100 .
1. .. . ., 1985. 272 .
2. .. . ., 1984. 278 .
3. ..
. //
( .-. . . 15- ,
, 7-8 2008 .) : - , 2008. 365-370 .
It is established, ecological resources of areas of selection of wildgrowing populations Fragaria orientalis Los. Differ warmly - in vegetation of plants: Verhneviljujsky and Mountain
area differ rather high moisture and heat lack, Amginsky - moderated warmly Heat oisture Area; in Megino-Kangalassky and UstAldan area it is noted high heat at a moisture lack; Olekminsky in
comparison with Hangalassky area where take place tests introduction,
has the highest warmly - and. For years of researches on productivity
samples of populations from Verhneviljujsky, Megino-Kangalassky and
Mountain area, having on the average productivity 100 from a bush
and above were allocated.
Key words: Fragaria orientalis, selection, Yakutia, an introduction, productivity.
132
_______________________________________________________________
633.31/37:633.366:581.41(47+57)
MELILOTUS MILL.
..
..
. .. ,
190000, -,
. , 42-44
e-mail: g.talovina@vir.nw.ru;
t.smekalova@vir.nw.ru
13 Melilotus
Mill. , , ,
.
. 2 ,
.
, , ,
.
,
, . ,
, ,
.
: ,
Melilotus, , , .
,
, [3, 8, 17 .].
[8, 9 .].
Melilotus ( )
13 , 2 : 11 (M. albus Medik., M. altissimus Thuill., M. arenarius Grec., M. dentatus (Waldst. &
Kit.) Pers., M. hirsutus Lipsky, M. officinalis (L.) Pall., M. polonicus (L.) Pall., M. scythicus
O.E. Schulz, M. suaveolens Ledeb., M. tauricus (Bieb.) Ser., M. wolgicus Poir.), 2 Micromelilotus O.E. Schulz (M. indicus (L.) All., M. spicatus (Sm.) Breistr.). ,
.
-
. Micromelilotus ;
( )
[6]. , [13].
.
(). . ,
; ; ; , , , ; (
); ;
, . , , (. indicus), , (M. spicatus).
133
______________________________________________________________
. 2-5 ( 15-20) , , .
, ( M. tauricus), ( M. albus, M. officinalis).
. , , ,
. , .. [7] , M. albus M. officinalis , , ,
(, , , ..).
. ,
, ,
. , , , ,
, - , , .
,
; .
-. , , . , M.
dentatus , - , . polonicus 2-6 , , , . ,
, .
,
; , . , -
; .
Micromelilotus , -, , , , ,
- M. dentatus; - M. suaveolens, M. officinalis, M. albus, M. altissimus.
; - - M. hisutus,
M. altissimus ., - M. arenarius, M. polonicus, M. spicatus .; - M. altissimus, M. indicus ., -- M. arenarius, M. spicatus .;
(M. albus, M. altissimus, M. dentatus, M. officinalis, M.
suaveolens .) .
2,5 8 , , ,
(M. spicatus).
, M. wolgicus -- . ,
, , , ,
, . ; , . , M. hirsutus , , .
(, ),
[1; 16].
, , , 2-8
, , ; (M. officinalis, M. hirsutus,
M. tauricus, M. scythicus) ,
[1, 2]. . 10, . . : -
134
_______________________________________________________________
1,5 ; M. polonicus 1,5 ; M. indicus,
M. dentatus, M. arenarius .
M. hirsutus, M. tauricus, M. altissimus, M. spicatus.
. M. officinalis M. suaveolens [16]: M. officinalis 5-7 (
4 8), M. suaveolens 2-4 ( 1).
, , , : 19,5 31,1 ; 13,0 19,5
[5, 12]; . Melilotus
,
[5, 12]. , Melilotus, , , . ,
Melilotus,
(M. albus, M. indicus, M. messanensis, M. segetalis, M. sulcatus), ,
M. albus , [12];
, - ,
,
Micromelilotus. M. albus , , ;
, . 9
(M. albus, M. dentatus, M. hirsutus, M. officinalis, M. polonicus,
M. spicatus, M. suaveolens, M. tauricus, M. wolgicus) ,
M. albus M. polonicus, M. polonicus [5]. , . , , , Melilotus.
, 2-3 7-8 , , , - , . -, -; -,
-, -, -.
,
[4, 7, 14, 15, 16].
,
: 3-7 , , , - -. Micromelilotus 2,5-5(8)
, -, -. : M. dentatus;
M. spicatus. , ; 1-, 2-3-.
. Fabaceae. , . , , . 100%
. .
, . , ,
, 1,5 3 . M. polonicus, M. arenarius, M. wolgicus, M. indicus, M. spicatus.
135
______________________________________________________________
, , , , , ;
, .
,
, ( - )
, [2].
, , 900 (M. albus, M. officinalis, M. indicus); , 900 (M. altissimus, M. hirsutus, M.
suaveolens) , 900 (M. dentatus,
M. polonicus, M. tauricus).
; ,
( Melilotus) ( Micromelilotus) (.
24 (. 1*).
. [10, 11] Statictica 6.0. ( ), ..
- . 2 , ,
, , ().
(. 1).
(Factor 1) 36% . 9 : ,
, , ,
, .
,
,
, , , . , , M. polonicus, M. arenarius; M.
indicus, M. spicatus.
(Factor 2) , , .
21% .
, . ,
, M. hirsutus, M. tauricus M. altissimus. M. dentatus M. indicus,
.
2 Melilotus
.
Melilotus , , .
136
_______________________________________________________________
. 1. *
*(zhc ; hr ; kst
; hkst ; lctn . ; lcv .; okr
; otnc ; opz ; st/z ; nsp ; gif ; fb ; fvb ; opb - ; kil ;
hrpb ; hrps ; rs ; fs
; fl ; fvl - ; fkrl ; prls ).
137
______________________________________________________________
. M. officinalis, M. albus, M. dentatus, M. suaveolens M. hirsutus M. tauricus, , . 7- .
. 2. Melilotus
138
_______________________________________________________________
5. .. Melilotus Adans. // , , . 13, 1957. . 139-142.
6. .. . .: , 1962.
277 .
7. .. Melilotus // , . 13, . 1 .; ., 1950.
. 345-502.
8. .. . .; .: , 1966.
611 .
9. .. . .: , . ., 1970. 145 .
10. .., .. . ., 1977. 152 .
11. .. : . . .: - . -, 1984. 288 .
12. Gazar M. Pollen morphology of the three genera of subfamily Papilionoidea in Egypt (Melilotus, Trifolium and Trigonella) // Acta Botanica Hungarica, 45 (3-4) 2003. P. 279-296.
13. Raunkiaer C. The Life Foms of Plants and Statistical Geography. Oxford, 1934. 632p.
14. Schulz O.E. Monographie der Gattung Melilotus. // Engler's botanischen Jahrbuchen.
29 Band, 5 Heft, 1901. P. 660-735.
15. Seringe. Melilotus // Prodromus Systematis Naturalis Regni Vegetabilis, auct. A.P. De
Candolle. Pars 2, 1825. P. 186-188.
16. Stevenson, G. A. An agronomic and taxonomic review of the genus Melilotus Mill. // Canadian journal of plant science, 1969. Vol. 49. N 1. P. 1-20.
17. Zohary M. Plant life of Palestine, Israel and Jordan. New York, 1962.
G.V. Talovina
T.N. Smekalova
All-Russian Vavilov Institute of Plant
Industry, 190000, Saint-Petersburg,
Bolshaja Morskaja str., 42-44
e-mail: t.smekalova@vir.nw.ru;
g.talovina@vir.nw.ru
Morphological characters of 13 Melilotus Mill. species were investigated for revealing of taxonomically significant characters and the diagnostic treats. It is necessary for differentiation of close species. For revealing of structure of morphological characters interrelation the factorial analysis on a method of the main a component was carried out. 2 factors which cover more than half of characters variability are allocated.
The first factor has the greatest correlation with the seed and bean form,
brush density, the form of a leaflet and its top, relative height of a plant
and duration of life cycle. The second factor is shown through "leg" presence ovary, a parity of corolla parts lengths. It is established, that the
studied species form groups which essentially differ from each other as
for investigated morphological characters. It gives the chance to specify
systems.
Key words: morphological diagnostic characters, Melilotus species,
factorial analysis, systematics, taxonomical importance of characters.
139
______________________________________________________________
635.65:575
IN VITRO
..
..
. ..
, 396030
,
, . , 8(47340) 5-33-27
e-mail: vniiss@mail.ru,
samani84@mail.ru
, , in vitro, .
: , ,
, , .
, , ,
. [4].
. .
.
.
,
,
[2, 5].
in vitro.
:
,
;
,
.
. : , , . 30 .
, , [1], t = +23-25, 16-
.
140
_______________________________________________________________
, . ,
.
10 % , 90,9 %, 39,4 %
51,5 % (. 1).
1
in vitro
Domestos
,
%
10
0,05
0,01
10
10
, .
99
384
132
141
207
%
%
39,4
51,5
12,5
10,4
2,3
2,3
2,1
2,1
8,7
10,1
%
9,1
77,3
95,5
95,7
81,2
2
5
3
3
4
9,1 %. .
1-3 . . 1-2 .
5 2 . , 10
.
77,3 %. in vitro 12,5 % 10,4 % .
.
. 7-8 .
5-6 . . 5 .
, .
domestos 95,5 % 95,7 % .
4,6 %, domestos
4,2%.
,
domestos 2,3 % 2,1 %, . , . , 5-7 . 1 .
( 1,5 ). , 3 .
207 . 81,2 %. 3-4 . . 5-6 . 4 .
141
______________________________________________________________
,
0,05 %.
, ,
.
.
, 7 10
.
, 2,0 4,0 , 1 (. 2).
2
1
2
3
4
5
6
()
(/)
, ,
0,1
, ,
0,2
0,3
+ 0,1
0,4
+ 0,1
0,5
+0,1
0,5
+ 0,1
-
1 (),
36
12
46
12
27
12
26
23
36
36
26
36
24
1 4, , , ,
2 7 . 1 ,
, 1 3.
, 4 , , . 4.
5, 6 , /
, .
26 . 1 36 , 5. ,
.
( ) 1.
2 4 .
, 0,5 / () 0,1 /
, in vitro
5, 6. 34
. .
142
_______________________________________________________________
in vitro .
in vitro
, . ,
[3, 6]. in vitro,
.
, ,
.
,
0,5 /.
30%.
1-2 , 1-2 . 2 4 .
, 1,0 /
, 15 %.
(42%) 1,5 /,
2-3 , 2-3 . 3 5 (. 3).
3
()
,
/
0,5
1,0
1,5
2,0
0,5
1,0
0,2+0,2
0,5+0,5
1,0+1,0
1,0
, .
, %
60
30
15
42
2
0
0
0
0
0
0
0
,
;
1-2; 1-2
,
2-4
2-3; 2-3
1-2; 1-2
3-5
1-2
( 2,0 /)
,
2 %.
. .
, in vitro 1,5 /.
,
, ,
in vitro, .
1. .. , / .. . .: , 1970. 342 .
143
______________________________________________________________
2. .. in vitro. ., 1988. 15 .
3. .. . .: , 1974. 253 .
4. .. . .: , 1973. 312 .
5. .. ( ) / .. , .. , .. . ., 2005. 20 .
6. .. // . .: , 1975. . 126-145.
144
_______________________________________________________________
: 58. 032.3: 58. 036.1
JUGLANS
..
..
, 308015,
. , . , 85.
e-mail: sorokopudov@bsu.edu.ru
, J. manshurica, J. siboldiana, J.
cordiformis,
J. regia, J. nigra. J. regia J. nigra, J.
cinerea . J.
manshurica, J. siboldiana, J. cordiformis
35 .
, . J.
manshurica .
: Juglans, , , , , .
, , , .
. ,
, , , .
[6].
, . ,
[4]. .
, , . Juglans ,
, [2, 3].
,
.
Juglans.
Juglans, , 2001 : J. regia L., J. manshurica Max., J. siboldiana Maxim., J. cordiformis Max., J. nigra L., J. cinerea L., J. rupestris Engelm. . J. regia L.,
. .
145
______________________________________________________________
27, 30, 35, 40 40 54%
. , . (, , 1970).
-
(7- ), , , . : , , (1:1:1).
[8].
13,
-5, 10 Quanta 200 3D , -.
,
50- .
, , () [5],
. ,
,
.
.
. . (1984), Microsoft Office.
Juglans (. 1 ).
J. manshurica, J. cordiformis, J. siboldiana
( 30% ), J. regia, J. nigra ( 14% ). J. manshurica, J. cordiformis, J. rupestris
40,
45 . J. siboldiana, J. regia, J. nigra, J. cinerea 45.
. 1. Juglands
146
_______________________________________________________________
. J. manshurica, J. crdiformis, J. siboldiana, J. cinerea , ( 1 ).
Juglans
, . ,
. J. regia, J. nigra, J. rupestris , J. manshurica,
J. crdiformis, J. siboldiana, J. cinerea
[7].
(54%). 2, .
. , 2, [9].
. 27 J. manshurica, J. siboldiana, J. cordiformis (. 2).
. 2.
147
______________________________________________________________
40 (. 3 ). 35 40 (. 3 ). , J.
regia , [10],
27-35. , [11]. .
27-35 (. 3).
(45)
, (. 3 ).
27-45 (. 2).
20%, 45.
( ),
(. 3). , 27-35
, .
. 3.
J. siboldiana
( 30% ) (. 1 ). 8
27-35 40-45 (. 3).
J. cinerea
( 22% ) (. 1 ).
( 2). 40 (. 3).
(27-45). 40 (30% )
(. 1 ). , J. regia (. 2). 40, 45 .
35 (
). 40-45 (. 3).
148
_______________________________________________________________
J. cordiformis 30%
(. 1 ).
(. 2). 35 4045 (. 3).
. J. siboldiana, J. nigra
,
.
(54%).
[4]. , . , , . 40-45.
J. manshurica
35,
. , [7].
J. siboldiana, J. nigra,
J. cordiformis, J. rupestris 35 (. 4).
[7], .
. 4.
35 , 40-45.
J. manshurica, .
149
______________________________________________________________
(. 5).
. 5.
.
J. manshurica, .
, , .
, , J. manshurica.
. , , , , . .
150
_______________________________________________________________
. J. manshurica
( ).
1. .. Juglans L. /
.. , .. // . .
(23-25 2003 ., ). 2003. . 179-180.
2. .. //
. 2002. . 274. . 444-448.
3. .. //
. . 2009. 5. . 60-62.
4. , ..
, /
.. , .. , .. , .. , .. , .. // , 2007, 54, 1, . 54-58.
5. , ..
/ .. , .. //
( . 19-22 2006). : , 2006. . 344.
6. .. /
.. , .. : , 1991. 307 .
7. .. Juglans / .. , .. , .. //
. , -
9 (104) 2011, . 15/1. .302-307.
8. .. - Ribs
americanum Mill. / .. , .. //
: , ,
, .-. ., .:
2008. .133-135.
9. , .. -
/ .. , .. //
. 1992. 2, . 24. . 107-116.
10. Cochard Herv Putative Role of Aquaporins in Variable Hydraulic Conductance of
Leaves in Response to Light / Herve Cochard, Jean-Stephane Venisse, Tete` Severien Barigah,
Nicole Brunel, Stephane Herbette, Agne`s Guilliot, Melvin T. Tyree, and Soulaiman Sakr // Plant
Physiology, January 2007, Vol. 143, pp. 122133. (10).
11. Cochard Herv Unraveling the Effects of Plant Hydraulics on Stomatal Closure during
Water Stress in Walnut / Herve Cochard, Lluis Coll1, Xavier Le Roux, and Thierry Ameglio // Plant
Physiology, January 2002, Vol. 128, pp. 282290. (11).
It is established that the least water-retaining ability leaves J possess. manshurica, J. siboldiana, J. cordiformis, the greatest waterretaining ability leaves J possess. regia, J. nigra. J. regia J. nigra, J. cinerea possess high thermoreactance. J. manshurica,
J. siboldiana, J. cordiformis are characterized by the big conductivity at
action 35 With. The basic cages abaxial and at moisture loss reduce the area, the factor of tortuosity of anticlinal
walls increases. The basic cages J. manshurica differ the least stability to
water deficiency.
Key words: Juglans, a sort, kinds, coma, a sheet surface, Water.
151
______________________________________________________________
(581.8 + 581.4):582.682 + 615.322
-
PAPAVER SOMNIFERUM L. - P. RHOEAS L.
.. 1, .. 2,
.. 1, .. 1,
.. 3
1
, 440026,
. , . , 40.
e-mail: sef1957@mail.ru
2
. .. ,
440026, . , . , 37
, .
.
: , -, ,
,
"", 107139, . ,
. , 1/11
Papaver
, , .
P somniferum L. 1
, , , . ,
, . , ,
, . , , , - , . , .
-. , , , . .
, .
P. somniferum L. 20% , (20-30%), , , , , , , , (20-30%) 40-50% . ,
[1].
40-60% 20% , . (2,6%), (28,3%), (4,6%), (64,5%),
75,5-76,63% 11,2-12,3% [2].
P. rhoeas L. 30-80 . , , . , -, -, . , -, . .
- , . . (, , , ),
(), . ,
152
_______________________________________________________________
.
.
. ,
, , . , , -, , , , ;
, , [3].
,
,
.
Papaver, : P. somniferum L. P. rhoeas L.
(1:3) 6% ;
[4-5]. 0,1%
.
-1, - SONY DSC-T20.
( 1 , ,
) [6-10].
- P. somniferum
L. , . ,
, 1:5 1:20.
.
, , . , .
,
, (.2, 3; .1, ).
.
(.2, 3).
, , .
, .
, 4, . , 5- (. 2, 3;
.1, , ). , , .
.
. , (.1, .2). : , , .
,
. , .
153
______________________________________________________________
. 1. P. somniferum :
. 157; , . 1540
: , ,
, , , , .
154
_______________________________________________________________
1
.2
P. somniferum
(. 1 - 157, 2 - 1540, 3 - 1520)
(. 1 - 157, 2 - 1540)
:
,
, , , , , .
,
(. 2, 2, 2; .1, ). , , , .
, (
) . -
155
______________________________________________________________
. , .
.
. 3.
P. Somniferum :
. 157; , . 1540
:
; ,
,
, ,
- ,
P. rhoeas L. P. somniferum L.
(. 4 , ). , P. rhoeas L. , , ,
156
_______________________________________________________________
. ,
.
. 4. P. rhoaes L.
(.157), - ( . 1520),
- (. 1540)
: , ,
, , ,
, ,
, . . , .
. ,
.
- [7] .
157
______________________________________________________________
, -
, . , - , :
, - [11].
1. -
P. somniferum L.:
, ,
, ,
2 , ,
.
2. - P. rhoeas L. P. somniferum L.; : P. rhoeas L. ,
.
1. . . : , 1971. 270 .
2. .. . .: -
, 1973. 412 .
3. .., .. . . 2 /
.: ; .: -, 1999. 816 .
4. .. . .: , 1960. 206 .
5. .. . .: , 1970 255 .
6. .. . .: ,
2000 528 .
7. .., .. . . .: -, 2007.
.1. 192 .
8. / . .. . , : - / , , 1963. 504 .
9. .. - .
.: , 1979. 154 .
10. . . 2- . . .: , 1980. - 558 .
11. .. (, , ) // , . .: , 1975. .55, . 1. - .175 221
e-mail: sef1957@mail.ru
2 The
Penza State Pedagogical University of Belinsky name, 37, Lermontova st., Penza, 440026
3 State
The anatomic research of two poppy species peduncles, differing taxonomic place, was carried out. The micromorphological
features of studied structures during budding - wax ripeness - are
revealed in comparative aspect.
Key words: opium poppy, corn poppy, anatomic features, peduncles
158
_______________________________________________________________
634.722
,
(Mahonia aquifolium (Pursh) Nutt.).
: Mahonia aquifolium, , , ,
, , , ,
1806
1818
(1775-1816).
(Mahonia aquifolium (Pursh) Nutt), ,
.
,
, - .
,
1-1,2 . . 20 ,
, , -,
. 5-9
8 . , ,
-, , -,
- .
,
, -
. e - . ,
5-8 . -
.
, . , -
- . , .
,
, .
.
: , , , ,
. ,
.
[1]
[2,3, 4].
159
______________________________________________________________
, : , , . 24 , . , .
.
I.
Mahonia aquifolium
(Pursh) Nutt. ,
Mahonia aquifolium (Pursh) Nutt. RTG/01/3 " , " 22.07.2002 . 12-06/52 ( 6, 2002 .).
II.
1. , , .
2. ,
, .
3. , . , .
4. , , .
III.
1. , , , . -
, . .
2. ,
, .
3. , 6 .
.
4. .
IV.
1. 6 (24),
6 ( 4).
2. 1% 95%, 1- .
160
_______________________________________________________________
V.
. , , ,
,
.
12 (: ).
VI.
, , , VII. (*)
,
, ,
. (+) ,
() .
(1 - 9) .
(a)-(d) VIII.
VII.
1.
(*)
(a)
2.
(a)
3.
(*)
(b)
4.
(b)
5.
: - (b)
6.
: - (b)
: - (b)
7.
8.
9.
: - (b)
: - (b)
10. :
(*)
(b)
11.
(*)
(b)
12. :
(*)
(b)
1
3
5
3
5
7
1
2
3
4
3
5
7
3
5
7
1
9
3
5
7
1
9
3
5
7
1
3
5
7
9
1
2
3
1
3
45
45
(90)
90
5
5-7
7-9
9-11
11
161
______________________________________________________________
(+)
13. :
(b)
14. :
(b)
15. :
(b)
16. :
(*)
(b)
17.
(*)
:
()
(b)
18. :
(*) ()
(b)
19. :
(*)
20. :
(c)
21. :
(c)
22. :
(*)
(c)
23. :
(*)
(+)
(d)
24. :
(*)
(d)
5
7
3
5
7
1
3
5
7
1
3
5
7
1
2
3
1
2
3
4
1
2
3
4
1
2
3
4
3
5
7
3
5
7
1
2
3
1
2
3
4
5
1
2
3
4
-
-
-
-
-
-
-
VIII.
, (a)-(b)-(c)-(d)
, :
(a) : ;
(b) : ;
(c) : ;
(d) : .
162
_______________________________________________________________
12. :
163
______________________________________________________________
22. :
4
-
164
_______________________________________________________________
1.
Mahonia aquifolium
(Pursh) Nutt.
( )
( )
2. ___________________________________________________
( )
3.
___________________________________________
_______________________________________________
4. ,
4.1
4.1.1 :
[ ]
(a)
[ ]
( -)
(b)
[ ]
( () ()-())
(c)
[ ]
4.1.2
( -)
[ ]
_____________________________________
4.1.3
(, , )
[ ]
4.1.4
[ ]
( ) ___________________________________________
4.2
4.2.1
{......}
[ ]
165
______________________________________________________________
4.2.2
( )
[ ]
5. ( ). .
5.1 :
(1)
1[ ]
3[ ]
5[ ]
5.2 :
(10)
1[
3[
5[
7[
9[
]
]
]
]
]
5
5-7
7-9
9-11
11
5.3 : 1[
(12)
3[
5[
7[
]
]
]
]
5.4 :
(16)
1[ ]
2[ ]
3[ ]
5.5 :
(23)
1[
3[
5[
7[
9[
5.6 :
(24)
1[ ]
2[ ]
3[ ]
]
]
]
]
]
6.
____________
____________
____________
____________
____
_____________________________________
_____________________________________
_____________________________________
_____________________________________
____
-
__________
__________
__________
__________
__________
__________
__________
__________
____
____
7. , .
7.1. , 5 6,
, ?
[ ]
[ ]
, .
7.2 .
7.2.1 ?
[ ]
[ ]
, .
7.3 .
(912 1318 ) , , , , .
166
_______________________________________________________________
8.
,
- 5 1996 ?
[ ]
[ ]
?
[ ]
[ ]
, .
9. , .
9.1
,
(.. ),
, , ,
, ..
9.2 ,
,
. ,
.
, ,
:
(a) (.. , , )
[ ]
[ ]
(b) (.. )
[ ]
[ ]
(c)
[ ]
[ ]
(d)
[ ]
[ ]
, .
9.3
?
[ ]
..
[ ]
,
,
, .
1. .., .., ..
(Mahonia aquifolium (Pursh) Nutt.). : , 2008. 22 .
2. Ahrendt L. Berberis and Mahonia. A taxonomic revision // J.Linn. Soc. (Bot.) 1961.
Vol. 57 (369). P. 1-410
3. Lee M.K., Kim H.S. Inhibitory effects of protoberberine alkaloids from the roots of Coptis
japonica on catecholamine biosynthesis in PC12 cella. Planta Med. 1996. Vol. 62. P. 31-34.
4. Hansel R. Mahonia aquifolium Ein pflanzliches Antipsoriatikum. Dt. Apoth. Ztg. 1992.
Bd. 132- 140. S. 2095-2097.
167
______________________________________________________________
WORKING OUT OF THE TECHNIQUE OF CARRYING OUT
OF TESTS FOR DISTINGUISHABILITY, UNIFORMITY AND STABILITY
MAGONIJA (MAHONIA AQUIFOLIUM (PURSH) NUTT.)
V.N. Sorokopudov
O.U. Zhidkih
O.A. Sorokopudova
Belgorod National Research University, Belgorod,
street of the Victory 85
e-mail: sorokopudov@bsu.edu.ru
168
_______________________________________________________________
581.9:(470.23)
..
..
. .. ,
190000, -,
. , 42-44
e-mail: t.smekalova@vir.nw.ru;
l.shipilina@vir.nw.ru
. ,
.
.
: , , .
, ,
, ,
, ,
.
(), () (),
( ). , , , . - ,
(
), , . ,
: , , , ,
(-, .).
,
(, ).
, (
- , )
, ,
, .
( , ) (, ) , , ,
[3].
- [4, 5,6,7].
() [1].
( ) ( ), .. -
169
______________________________________________________________
.
, , ,
.
-
,
. 1294 .
, 716 (94,6%), Liliopsida - 231
(30,6%), Magnoliopsida 485 (64%); 5,4%
: Lycopodiophyta 9 , Equisetophyta 8 , Polypodiophyta 19 , - 5,2%; Pinophyta 2 (0,4%). , 7,55% 57 .
539 309
71 . , 526
(97,6%), Liliopsida - 96 (17,8%), Magnoliopsida
430 (79,7%); 2,4% Pinophyta 13
. ,
7,59% 41 . 19
(huja occidentalis, Colchicum speciosum, Schisandra chinensis, leretum bellidiforme,
Portulaca grandiflora, Fagus sylvatica, Reseda lutea, ropaeolum majus .). : Asteraceae (55); Brassicaceae (47); Rosaceae (44); Poaceae (42); Fabaceae (35); Caryophyllaceae (23); Liliaceae
(13). Rosa (8); Vicia (8); Lilium (7);
Atriplex (6); Crocus (6); Rumex (6). , : Rosaceae, Fabaceae.
103 , 14%
, , 6% , (, , 2005). , - , .. (, .).
, ,
, 116 23 . () Poaceae (27), Fabaceae (22), Rosaceae (15), Brassicaceae (12), Polygonaceae (9), Chenopodiaceae (5).
1.
2. FABACEAE
3. ROSACEAE
4. POLYGONACEAE
5. PAPAVERACEAE
6. AMARANTHACEAE
7. BRASSICACEAE
3
2
2
1
1
1
1
14
10
10
6
3
3
3
romus, Avena, Lolium
Vicia, elilotus
Rosa, alus
Rumex
Papaver
Amaranthus
Lepidium
, , -
170
_______________________________________________________________
.
.
:
(71,84%),
, .
Poaceae.
. Ericaceae, Rosaceae, Fabaceae, Lamiaceae. : Alliaceae, Polygonaceae, Betulaceae, Hypericaceae, Cannabaceae,
Urticaceae, Grossulariaceae, Linaceae, Viburnaceae, Caprifoliaceae, Solanaceae, Asteraceae, . Poaceae, Apiaceae. Ericaceae,
: . (Vaccinium myrtillus, V. uliginosum) , , . (xycoccus palustris, . microcarpus, Rubus chamaemorus) , .
Fabaceae (18), Poaceae (17), Rosaceae (14), Polygonaceae (4), Grssullariaceae (4).
, - , , .
,
. :
Allium schoenoprasum, Vaccinium uliginosum;
Festuca rubra, F. ovina, Poa pratensis, P. nemoralis, P. palustris, Phalaroides arundinacea, Alopecurus aequalis, Rumex maritimus, Rhodococcum vitis-idaea, xycoccus palustris, Mentha arvensis;
Agrostis canina, Alopecurus geniculatus, Vaccinium myrtillus, umulus lupulus, Rubus
arcticus, R. chamaemorus, Rosa acucilaris, athartolinum catharticum, ulgedium sibiricum; Allium angulosum. lymus caninus, Arrenatherum elatius,
Trisetum flavescens, Agrostis gigantea, A. capillaries, A.stolonifera, Festuca polesica,
Schedonorus giganteus, S. pratensis, Poa angustifolia, P. remota, P. trivialis, P. compressa, Dactylis glomerata, Cynosurus cristatus, Beckmannia eruciformis, Phleum nodosum,
P. pretense, Alopecurus pratensis, Rumex hydrolapathum, R. pseudonatronatus, R. aquaticus, ypericum perforatum, . maculatum, xycoccus microcarpus, Urtica galeopsifolia, Ribes spicatum, R. scandicum, R. alpinum, R. nigrum, Rubus idaeus, Rosa majalis, R.
mollis, Fragaria viridis, Padus avium, nobrychis arenaria, Vicia sylvatica, V. cassubica,
Lathyrus vernus, L.sylvestris, Trifolium montanum, arum carvi, enanthe aquatic,
Daucus carota, Lonicera xylosteum, Solanum dulcamara, Dracocephalum ruyschiana,
Salvia verticillata; Allium oleraceum, Bromopsis riparia, Festuca unifaria, F. brevipila, Poa humilis, Rubus nessensis, R. saxatilis, Rosa dumalis, Fragaria moschata, alus sylvestris, Anthyllis arenaria, A. schiwereckii, A. macrocephala, Anthriscus
sylvestris,
Lonicera
baltica;
-- Setaria viridis; -- orylus
avellana; Fragaria vesca, Lathyrus pratensis,
Viburnum opulus.
171
______________________________________________________________
:
- Agrostis capillaris, Festuca rubra, F. ovina, Poa palustris,
P. palustris, Phalaroides arundinacea, Rumex aquaticus, Vaccinium myrtillus, V. uliginosum, Rhodococcum vitis-idaea, xycoccus palustris, . microcarpus, Ribes nigrum, Rubus idaeus, R. saxatilis, R. arcticus, R. chamaemorus, Lathyrus palustris, ulgedium sibiricum; (75) - Allium angulosum, A. schoenoprasum, lymus caninus, Arrenatherum elatius, Agrostis gigantea, A.stolonifera, Schedonorus giganteus, S. pratensis,
Poa pratensis, P. humilis, P. angustifolia, P. remota, P. trivialis, P. nemoralis, P. compressa, Dactylis glomerata, Rumex hydrolapathum, R. maritimus, orylus avellana,
ypericum perforatum, . maculatum, umulus lupulus, Ribes spicatum, R. alpinum,
Rosa majalis, R.acucilaris, nobrychis arenaria, Lotus corniculatus, Lathyrus vernus,
athartolinum catharticum, Anthriscus sylvestris, arum carvi, enanthe aquatica, Solanum dulcamara, Dracocephalum ruyschiana, Salvia verticillata .;
- Lonicera xylosteum; - Beckmannia eruciformis,
Urtica galeopsifolia, Daucus carota, Viburnum opulus; - Setaria
viridis.
,
(Allium angulosum, lymus caninus,
Arrenatherum elatius, Trisetum flavescens, Agrostis gigantea, A.stolonifera, Festuca
polesica, Schedonorus giganteus, S.pratensis, Poa angustifolia, P.remota, P.trivialis,
P.compressa, Dactylis glomerata, Cynosurus cristatus, Phleum nodosum, P. pratense,
Rumex hydrolapathum, R. pseudonatronatus, ypericum perforatum, . maculatum,
Ribes spicatum, R. scandicum,R. alpinum, Rubus caesius, Rosa majalis, R. mollis, Fragaria viridis, Padus avium, nobrychis arenaria, Vicia sylvatica, V. cassubica, V. ,
Lathyrus vernus, L.sylvestris, edicago falcata, . lupulina, Trifolium montanum, . hybridum, . repens, . medium, arum carvi, enanthe aquatica, Solanum dulcamara,
Dracocephalum ruyschiana, Salvia verticillata).
: (Allium oleraceum, Bromopsis riparia, Festuca unifaria, F. brevipila, Poa
humilis, Rubus nessensis, Rosa dumalis, Fragaria moschata, alus sylvestris, Anthyllis
arenaria, A. schiwereckii, A. macrocephala, Lotus corniculatus, Lathyrus linifolius,
Anthriscus sylvestris, Lonicera baltica); (Agrostis
capillaris, Alopecurus pratensis, Rumex aquaticus, xycoccus microcarpus, Ribes
nigrum, Rubus idaeus, Vicia sepium, Lathyrus palustris); (Festuca rubra, F. ovina, Poa palustris, Phalaroides arundinacea, Rhodococcum
vitis-idaea, xycoccus palustris); (Poa pratensis, P.
nemoralis, Alopecurus aequalis, Rumex maritimus, Mentha arvensis); (Agrostis canina, Alopecurus geniculatus, umulus lupulus,
Rosa acucilaris, athartolinum catharticum); - (Vaccinium myrtillus, Rubus arcticus, R. chamaemorus, ulgedium sibiricum); (Beckmannia eruciformis, Urtica galeopsifolia, Poterium
sanguisorba); (Vaccinium uliginosum); (Allium schoenoprasum); -
(Lonicera xylosteum); (Rubus saxatilis); --- (Setaria viridis);
- (orylus avellana); -
(Lathyrus pratensis), -
172
_______________________________________________________________
2. .., .. .
, . 76, , 2005, 54 .
3. .. . : , 1978, 319 .1978
4. .. . .,
1977, .60, 1, . 69-73.
5. .. // ., ..,1982, .87, 4,
.3-22.
6. .. : ,
// . ., 1987,
.3-28.
7. - .. //
., 1980, .6, .761-774.
Crop Wild Relatives (CWR) in adventive and native fractions of Luga area
flora from Leningrad region are investigated. On features of the geographical
distribution, all CWR species can be carried to several groups. The allocated
speciess with related types of areas of distribution areas are united in larger
chorological groups.
Key words: Crop Wild Relatives (CWR), geographical distribution, types
of areas of distribution.
173
______________________________________________________________
633.31/37 : 633.366 : 502.171 (470+571)
..
..
. .. ,
190000, -,
. , 42-44
e-mail: t.smekalova@vir.nw.ru;
g.talovina@vir.nw.ru
in situ ,
; (),
. Melilotus
56- . M. scythicus . M. hirsutus,
.
in situ. M. scythicus , . M. scythicus
.
, , . ( in situ ex situ
).
: in situ , Melilotus (),
.
Melilotus Mill. ,
, , -
, , -
.
() , , (). , . , , ,
.
-
,
, [5].
. . . ()
,
(ex situ)
(in situ).
[1] in situ .
in situ .
,
. 1996
( ), in situ (4 ) ,
.
174
_______________________________________________________________
, , , ().
in situ ex situ
()
EX SITU
, - : , , ,
;
;
-
;
- ;
, -
;
;
;
,
, ;
.
.
IN SITU
; ; - ;
;
;
; - , - , ,
., - ;
, . . ;
;
-
(- ;
-, - .);
- .
;
, .
in situ ,
;
(),
. . , , . , .
, ( ),
.
[6], (
), , ,
.
.
, , .
, . , -
175
______________________________________________________________
,
.
,
in situ, [7]. :
1. ;
2. .
, :
1.
:
,
( , ,
, ..);
.
,
, 5 :
I. , , ;
II. , ;
III. , ( , );
IV. , ( );
V. .
2.
, ,
. . [7], [2] ,
:
(Endangered),
(Vulnerable),
(Rare).
in situ. , , .
, ,
( ) (in situ),
, :
- ,
, - ;
- , 1 2
.
: M. hirsutus Lipsky, M. scythicus O.E. Schulz. (
). , M. albus Medik., M. officinalis (L.) Pall., I , M. dentatus
(Waldst. & Kit.) Pers., M. suaveolens Ledeb., M. polonicus (L.) Pall., M. wolgicus Poir.
III .
, , . . -
176
_______________________________________________________________
, ,
,
(in situ).
, , ,
, , . , , , . .
.
,
, ,
, - .
,
,
(,
.).
, 1- , [3, 4]. Melilotus 56- : M. albus (43 ), M. officinalis (39), M. dentatus (9),
M. suaveolens (15), M. polonicus (3), M. wolgicus (2), M. hirsutus (1).
, Melilotus 4 8 ,
; , , , 3 . , .. .
. , ,
, . ,
,
. M. albus, M. officinalis -, , ; M. albus M. suaveolens ; M. wolgicus , ,
. M. dentatus, ,
. , , , .
M. hirsutus, in situ,
; M. scythicus . , , , .
, M. hirsutus , - , , [8]. M. scythicus
, ;
(MHA, LE) .
177
______________________________________________________________
M. scythicus , , . , 1933 .
. , M. scythicus
.
, ,
. , .
,
( in situ ex situ ). ex
situ ( , ) .
1. . ( , . .) // The Interim secretariat for the CBD. Geneva, Execitive Center, 1992. 34 c.
2. . . .: , 1988. 592 .
3. .., .., ..
/ . . . . .: , 2005. . 102-113.
4. .., .., ..
: / . . .. . .-, 2005. 85 .
5. .., .., .. (in situ) . // . , ,
/ - , 13-16 2001. ., 2001. 57-59.
6. .., .., ..
// / - 1 . .-. ., , 2002. . 265-271.
7. .., .. .
, . 76, , 2005, 54 .
8. IUCN Red List of Rare and Endangered Animals and Plants, which Particularly Protected
in Russia. Part 3.1. Seminal plants. Red data book Laboratory of All-Russian Research Institute of
Nature Protection; Res. Editor Prisyazhnyuk V.E. Moscow, 2004 (2005). 352 p.
The method of in situ conservation assumes of plant species preservation in natural, rather not broken ecosystems; such ecosystems are characteristic for especially
protected natural territories (NPT), first of all - reserves with a rigid mode of protection. Preservation of priority Melilotus species in territory of Russia is possible in limits of 56 reserves. M. scythicus it is not found out in territory of any of country reserves. M. hirsutus, grows in territory of the Caucasian reserve. Both species are priority for preservation in situ. For M. scythicus we offer studying of a condition of species
populations around growth, with the subsequent regular monitoring. Introduction M.
scythicus in the Red Book of the Astrakhan area is necessary. In case of species detection within existing NPT, located near known species sites, it is necessary to include it
in system of NPT monitoring and to apply measures of rigid protection. For both species application of complementary preservation (a combination in situ and ex situ preservation ways) is optimal.
Key words: in situ conservation, Melilotus species, criteria of priority for conservation.
178
_______________________________________________________________
635.9:582.579.2
-
..-.
.. , .. ,
.. , .. ,
. . , .. ,
..
, 364037, . ,
, . , 33
- ,
.
: , , .
( ) -. , ..
, .. , 1926; .. , 1947;
.. , 1966 .
, , , .. , .
-
,
. , , .. (1941) : ,
, ,
, ,
XVIII ,
. , . .
.. , .. , ..
(), .. , . .
.. , (1770-1773) , 1770 . .
. , .
1773 . ,
. ,
. 108 , .
20 . .. . -. 86 .
., , .
(1824,1825) (. .. ,1962) , , , .
, , .
179
______________________________________________________________
-
XIX .
. - .. . 1798 ., . ,
, ,
, .. .
. ,
, . . Flora Taurica-caucasica (1808-1919) (. .. , 1962).
.. (1938) ...
, , .
, .
.. 1829 . ,
,
(Corydalis emanuelii, Eunomia
rotundifolia .). (1836-1838 1843-1844 .) . . , 2500 ,
. Reise durch Russland
nach dem Kaukasischen in der jahren 1836, 1837 und 1838 (1842-1843 .).
1860-61 . () .. . ( ) .
.. .
(1901), ,
Nepeta supina, Lamiuin tomentosura, Gystopteris fragilis
.
,
.. , : (1890) 1887 . (1890), . , , , : ... ,
(Gentians
pyrenaica), (Saxifraga), (Cerastium), Draba scabra, (Potentilla nivea), (Alsine imbriata), (Alchemilla vulgaris), .
-
, .
XIX . . .. ,
20 .
, .
, : (1893);
(1894), 759 , Draba Saxifraga - ; (1898); - (1899), 158-160
88 .
.. . . 1896 . . - Daphne
baksania, .
180
_______________________________________________________________
1888-1889 . . .. - , . ,
. :
(1889); (1909); . Rindera Pall. (1909); (1910).
.. , , , ,
. , , , , . , , (, 1910). ,
, .
,
.
, . , , .., .. (1973).
.. (1970), : , , . , , , .
.. (1910) - . . - , ..,
. , , .
.. 1889 1893 . (1889), . Draba scabra, Campanula besenginica, Thymus lypskyi (.
.. , 1962).
.. .. . 47 ..
.
(1905) (1905). 3
1. - ( - .. ). 2. . 3. .
.
, Draba siliguosa, Thalitrum foetidum . -
Salvia canescens, .
181
______________________________________________________________
1909. - .. .
, , , . ,
Draba bryoides, ,
. - () .
1911 . .. ..
, -
. 1913 .
, -.
..
, , . Corydalis pallidiflora, Campanula besenginica
., . , .. , , , . , ,
, .
1917 ,
, . - ,
, .
1922 . -
- . .. .. (1926), 2 .
, .. . 4 : I. ; 2. , ; 3. ; 4. .
20- , . . .
. . (1925) (, , ), .
(1928)
: , , , -
. - . .. , , ... .
.
.. .. . 1925 . ,
...
: , 3500 . , , , (.. .. , 1926,
. 163). , , , , , , . -
, , , -
182
_______________________________________________________________
- -
, , ....
. 1898 . .. (1925) . , , 1894 . .. .
(1926), .. ..
Ranunculus balkharicus- Ranunculus
suukensis, .
-Papaver lisae, .
.. , .. .. 1911, 1913 1925 (1927),
.
. 1932 . .
, . - ,
-, , ,
,
. .
- (1935) .
. . : , ,
.
, , , . ..
( ) (1931); ..
(1932).
(1938) ..
. .
Gnaphalium supinum, Gorydalis alpestris, Jurinea subacaulis. Ha
microglochin, Colpodium versicolor.
,
.
( 1913 1932 .) .. 1938 . :
. 3793 , 170 , , .
, . .. , :
(1933-1939); (1936);
(1949); (1948);
(1949).
(1948) .. ,
... - ,
. -
183
______________________________________________________________
- , , -
,
, .
, .. -
.
.
.. (1948) -
: 1. ; 2. ,
- ; 3. ; 4. . - : 1 , , - ;
2 .
.. (1925), 1915 . ,
, :
; ; ; . , .. -
.
- , .. (1936)
.
.
..
- (1953). , , ,
. , . 2 : , ; - , , . , - (, ) .
.. :
1. , , -
; 2. , ; 3. , , ,
.
.. (1941, 1946, 1951 .),
, - .
, .. (1940); .. (1941); .. (1947); .. .
(1954); .. .. (1959), -
184
_______________________________________________________________
- . , , .
60-
,
.
,
(, )
, -
. .. (1957, 1960), .. (1953), .. (1946, 1969); .. (. , . , . .), . .. -
.
,
(1948);
(I960), , .
, .. (1969) 8 ,
, 147
6,4% . : , (1974)
(19746),
, .
- (1975) . , .
.. - (I960) ,
.
.. (1957) ,
.
. 2 : 1.
; 2. , . , , , , . : I.
; 2. -; 3. ; 4.
; 5. .
.. - (1953) 2
: 1. ; 2. . ,
.
- ( . . , ) .. (1968) . , , . :
185
______________________________________________________________
I. , ; 2.
, 40-60.
2 : , .
, .. (1910) . , , , . .. (1969, 1974, 1976, 1982) .. (1965).
20 , 80- 90-
, . , , , 1989 1991 .
.. (1989), , , , .
. , ,
(, 1988, 1989). (.., 1997, .., 2009),
, , .
.. ( ) (1973)
. , 87
30% .
.. (1963). - 140
.
-
.. (1974), 200-250 .
, -
(.. ,1946; .. , 1971;
.. , 1972; A.M. ,1981 .).
.
.. .. , .. , .. (1975).
-
,
,
, .. ,
(1976)
.. (1979, ),
, :
- ,
. , , .
. , , - . (
186
_______________________________________________________________
) -
.
,
.
1. .. . .1. , 1976.-215 .
2. .. . .
. 1947. 214 .
3. .. . 1966. 319 .
4. .. . M.-JT.: - , I94I. 275 .
5. .. -. , 1962. 145 .
6. .. . - ., 1938, 1, .46-55.
7. .. . , 1901, 199 .
8. .. . - .., 1890, .14, .1,
.9-24.
9. .. 1887 . - . .. 18906, T.I4,
.1, . 26-35.
10. . . . , 1893, 30 .
11. .. . ..
.-. , 1894, .27, 212 .
12. .. . , 1896, 36 .
13. .. - . - ... , 1899, .3, 186 .
14. .. . - .
, 1889, .1-19.
15. .. -- . - .. ., 1909, T.2I, I. 174 .
16. .. - .
Rindera Pall. - . , I9Q96, .7, .20-68.
17. .. . -, 1910. 48 .
18. .. . - . , .V, 1973, 3, .57-62.
19. .. .- ..
., 1956, .24, .261-311.
20. .. . -, 1905. - 47 .
21. .. . ., 1915, .15, .5, .1-17.
22. .. . - . , .,
1931, .23. 21 .
23. .. - . .-.: - , 1935.
107 .
24. .. .. 1925 . - . . ., 1926, T.I9, C.I63-I8I.
25. ., . .- . ... , 1956, T.I, .4. 48 .
26. . ( )., 1925, .27,.1-2, .52-62.
27. .. . - .- .. .
, 1936, .1, 256 .
28. .. - . - .;.: - , 1953. 368 .
29. .. .- , 1941, 3, .73-79.
187
______________________________________________________________
30. .. // .. . , 1947, 2, .159-167.
31. .., .., .. -
. , 1954, 136 .
32. .., .. // . . , 1959, .6, .119-130.
33. .. .-
., , 1957, .1, .85-112.
34. .. ,
. - .: . .; .: ,
1960, .5, .140-147.
35. .. - . -
.. -, 1953, .15, .251-282.
36. ..
(). - .. -. 1946, .9, .131-153.
37. .. - ( . ). . . . ., 1969, .84, .6, .86-92.
38. .. () . ... .. ., 1969. 42 .
40. .. -
( . . ., ). . ..., 1968. 23 .
41. .. . - .: . , 1979, .3, .92-94.
42. .. ( ). .: . , 1979 , .3,
.96-100.
This article describes the historical facts of studying rockscree flora of the Eastern Caucasus, as well as new species are
described by various authors.
Key words: Eastern Caucasus, rocks, scree, flora, vegetation.
188
_______________________________________________________________
:615.235:665.57.92
..
..
, 305041
. , . ., 3.
e-mail: fg.ksmu@mail.ru
,
Inula salicina,
, .. . .
, ,
.
.
: , ,
189
______________________________________________________________
1:20 95 1 . 1:10. , 1/5 .
96%
. , 70%
, . [3].
2 . (0,05)
5-10 , 2,5 ,
100-105 6 .
, . 2 .
--- (6:4:3) . (18:1:4:3). , 100 ,
10-15 [10].
, .
.. . Rana
Temporarea.[4].
.
, (),
6,87% 8,86%. , , - ; ( 1% 56); . , , [10].
, , , , , ,
, , .
, , , , ().
,
Inula helenium .
190
_______________________________________________________________
/ /
/ /
//
0,670,01
, %
32,060,53*
0,670,01
32,331,27*
0,810,02
18,751,73*
: *
0,05, n = 6 .
,
. , , , . ,
, .
.
1. , ..
. .-. . 14. 1980. . 74-81.
2. , . . 1878. . 46.
3. ..
// ... ., 1991. . I. .97-102
4. , .. .: , 1974. - 143 .
5. , .. .- . . - 1967. . 2. - . 207-213.
6. , .. . . ., 1959. . 124-129.
7. , .. . - : . ., 1990. - .212.
8. , .. . - . .-..-. 1941. - .4. - . 131-226.
9. .., .., .. Malva neglecta L. //
. . 1989. 2. . 280-281.
10. : , , ; Hippuridaceae - Lobeliaceae. ., 1991. - 200 .
11. , .. . , 1985. .86.
12. , ..
. - . : . . . . - 1973. - 3. . 76-79.
191
______________________________________________________________
: 615:322:547.56:577.114
VIOLA ODORATA L.
..
, 305041,
. , . ., 3
e-mail: fg.ksmu@mail.ru
(Viola odorata L),
.
1-2 , 70% - (1:100) 45 .
1,15
3,21%.
: (Viola odorata L), ,
, .
(Violaceae), , [7].
, , , [5], I ,
: , , , . [7].
, , , .
[3, 7].
,
, ,
. , , [3].
.
.
-
, 2009 . (. . , .
) .
, [1, 6]. , ..
.
70%
410 .
, .. , ,
[2, 6].
192
_______________________________________________________________
[2, 6].
. , 1-2 (. 1).
.
70% (. 1).
, 45 [6].
1
, :
0,5
1,0
2,0
3,0
: , %
30
50
70
96
, (70% ,
- 1:100):
30
45
60
,%
2,63
2,79
2,83
2,21
2,54
2,71
3,20
2,10
2,60
3,20
3,21
. ,
, ( 23.2,2068-89) 1 .
1.0 ( ) ,
250 , 100 70% +0,01 . , 45 ,
.
, 70%. ,
, 10 .
2 25 ,
5 5% 70% 10 1 3%
. 30 .
410 10 .
, 2 , 1 3% 70%
25 .
(), .
() :
193
______________________________________________________________
D . m . 1 . 100 . 100 . 100
;
=
D . m . 2 . 100 . (100 - W)
D ;
D () ;
m ;
m ;
W .
[4]. , 95% 3,63% (. 2).
2
N /
1
X
2,02
2,00
2,03
1,99
2,01
2,76
2,70
2,78
2,72
2,74
1,82
1,82
1,84
1,88
1,84
2,29
2,35
2,33
2,30
2,33
2,40
2,38
2,41
2,44
2,42
S2
Sx
P,% t(Pf)
1 ,
15,3
15,3
15,3
15,3
,
3,2
7,2
14,4
,
15,3
18,5
22,5
29,7
15,30
18,34
22,26
30,06
, %
- 0,87
- 1,07
+ 1,21
(. . 3).
5 . (. . 2),
1,15 3,21%.
194
_______________________________________________________________
,
.
(Viola odorata L), . 1-2 , 70% - (1:100) 45 . 1,15 3,21%
1. .., .. // . 1970. . 1. . 68-72.
2. .., ..
// . - -, 1973. . 168-172.
3. ..
: . . . . . , 2002. 24 .
4. . 11- . .: , 1987. . 1. 277 .
5. : , , ; Paeoniaceae Thymelaeaceae. ., 1985.
6. .., ..
// . . . . XXIX. ., 1991. . 150-155.
7. : 30- . ., : - , 1934-1964. . VI, 1941.
195
______________________________________________________________
: 582.949.27
..
..
. .
,
305041, . , . ., 3
e-mail: fg.ksmu@mail.ru
. . , , .
: , , ,
196
_______________________________________________________________
[5].
, , , , .
(5,9%) (6,3%).
1
,
-
, %
5,86
, %
6,3
5,9
0,6
0,4
4,9
. ,
96% .
: .
- (1:10). 96% [1].
( , )
( ).
,
(. 2).
2
,
, :
96%
1
2
3
30
60 (302)
90 (303)
120 (304)
1:2
1:3
1:4
1:5
, %
6,07
4,76
5,38
3,75
7,20
7,91
6,62
5,62
6,80
7,91
7,89
, ,
1 , 3- 90 (3 30 ) - (1:10) 4- 96%
.
197
______________________________________________________________
.
,
1 . 10 ( ) 250 , 100 , 30 . 100
30 . , 5000 / 10
500 5 ,
66 .
( ).
25 , 100
96% , , 60 5 . 30
5000 / 30 .
13-16 100-105 16 40 .
15 96% (3:1).
, 100-105 .
() :
(m2-m1)500100100
X (%) = --------------------------------,
m25(100-W)
m1 , ;
m2 , ;
m , ;
W , %.
,
5,15% 8,12%.
1.
.
2. .
1. , .., .., .. - // . 1984. - 3. . 13-16.
2. .. , .. , .. , .. 3 (: ) - - . ., 2004. 520 .
3. .. . ., 1970.- 378 .
4. : , , ; Asteraceae (Compositeae). .: , 1993. . 140.
5. .. (). ., 1978. 256 .
6. ..
// . : . . . . 1973. - 3.- . 76-79.
198
_______________________________________________________________
STANDARTIZATION OF LAPSANA COMMUNIS L. HERB BY CONTENT
OF POLYSACCHARIDE COMPLEXES
V.N. Bubenchikova
S.V. Logutev
S.N. Redkina
Kursk State Medical University,
Kursk K. Marks St., 3, 305041
e-mail: fg.ksmu@mail.ru
The paper gives the results of the isolation and chemical study of
polysaccharide complexes of Lapsana communis L. Their qailitative and
quantitative monosaccharide composition has been established. To standardize the herbs of Lapsana communis L. by content of polysaccharides
the gravimetric method with following studies have been proposed: extraction of polysaccharides from herbs; precipitation of polysaccarides.
Key words: Lapsana communis L., herb, polysaccharide complexes, gravimetric method
199
______________________________________________________________
: 633.812:581.192
..
..
,
305041, . , . . 3
E-mail: fg.ksmu@mail.ru
-. : 70% , , 0,5%
(1:1), 10% ; . : , Geranium sanguineum L.
, , ; .
- Ni, Co, V, Mo, Cu,
Pb, Zn, As, Sb, Sn, P, Ga, Bi, Tl, Ge, Ag, Ti, Mg, Cr, W, Be, Ba, Li, Cd, Sr, Zr,
Sc, La, Hf, Nb, Ta, Y, Yb, Hg, In.
: -, ,
200
_______________________________________________________________
1:10,
96% [3,4].
10%
12
. .
.
. -
. 96% . .
, [6].
, ,
2 . (0,05) 5-10 , 2,5 (1 /), 100-1050 6 ( ), 24 ( ) 48 ( , ).
, . -2 .
--- (6:4:3) . -
-- (18:1:4:3). - , 1000 , 10-15 [7].
.
,
t = 5500 2 .
8-1.
2% [5].
, - 4 : ,
5,74%, 7,24%, 1,1%, 2,28% .
, , , , , , . ,
, , .
, , .
, .
,
35 . :
(. , , , , );
(, ); (, , ,
); (, , , , , ,
).
, (0,0001%), (0,008%), (0,08%),
(0,1%).
, . ,
201
______________________________________________________________
, , ; .
(, Mn), (Mn, Cu, Zn, Cr, Mo .), , .
1. ..
//.. .-.,1991.-.XXIX.- .97-102.
2. .. // . ., 1992. - 477.
3. .., .., .. //
. . - 1993.-3.- . 355-357.
4. .., .., .. lva neglecta L. //
. .- 1989. - 2. . 280-281.
5. .., .., .. // . 2000. - 1.- . 25-27.
6. .., .., ..,
- // . . 1994. - 5. . 597-600.
7. .. / . - . 1978. 256 .
V.N. Bubenchikova
Zh.A. Bulatnicova
Kursk State Medical University,
305041, Kursk, Rassua
E-mail: Fg.ksmu@mail.ru
202
_______________________________________________________________
635.939.73
.. 1,
.. 2
1
, 393760,
, . - ,
. , 101
e-mail: mgau@mich.ru
2
, 393760, , . , . , 274
e-mail: mgpi_lab@mich.ru
. , 10 . 5-7 .
: ,
, , , , ,
.
- ,
, , , - [1]
.
. , , , .
- . , - , [2].
. [3, 4] : .
,
, . ,
,
.
:
, , 75% .
, ,
, , .
75% , ,
.
.
203
______________________________________________________________
.
1988 - 2009 .
.
59 ( )
- . -
( ,
), ( , ), ( , ), 14-9 ( , ). [3, 4].
STATISTICA 6.1. StatSoft StatGraphics Plus 5.0 .
(.1.).
, [4]
- 25% . 10 ( 3,56% 2,05%), ( 10,50% 13,05%), ( 4,59% 7,27%) - (
0,42% 1,05%). :
5 ( 0,31% 0,35%),
, 5 0,35%
0,26%.
, , , ,
.
.
, ,
- , ,
10-11 . 5
.
204
_______________________________________________________________
. 1.
15-
, .
, , .
10 . ,
( 10% 20%). , - , , . (. 2-4) .
205
______________________________________________________________
. 2.
, ,
,
5 , ,
.
. 3.
206
_______________________________________________________________
. 4. 14-9
14-9 (. 3, 4)
5-6 , ( ) , 5 , . , 14-9,
7-8 ,
- .
. , - , 10 , ,
.
5-7 .
1. , .. Lni (), / .. , .. , .. //
. . - 1981. - .17. - .4. .565- 568.
2. , . . / . . // : -
, 1981. 166 .
3. , /
: , 1995. 502 .
4. , / : , 1999. 608 .
207
______________________________________________________________
METHOD OF DETERMINING OF HONEYSUCKLE RIPENING BASED ON ANALYSIS
OF CHANGES IN BIOCHEMICAL COMPOSITION
F.G. Belosohov1
O.A. Belosohova2
1 FGOU
e-mail: mgau@mich.ru
VPO Michurinsk State Pedagogical Institute, 393760, Russia,
Michurinsk RF-Science City, Sovetskaya St., 274.
2 FGOU
e-mail: mgpi_lab@mich.ru
Key words: honeysuckle, the biochemical composition of fruits, vitamin C, vitamin P, glucose, organic acids, dry substances.
208
_______________________________________________________________
543.54:547.973
AMELANCHIER SP.
..
..
..
..
, 308015
., . 85.
- (ESI/MS) 6
. , -3, -3 -3-. ,
, 3- .
, (0.250 0.275 100
), - -
(0.102 100 ).
e-mail: deineka@bsu.edu.ru
: , , Amelanchier sp., ,
.
c, [1], - (
) .
[2].
- (-) . ,
[3-6] .
, Amelanchier, (Maloideae) (Rosaceae), - 18 23
[7]. , , , .
15 15 .
. [6] , . , , , . 7 (A. spicata (Lam.) C. Koch., A. alnifolia Nutt., A. oligocarpa Roem., A. sanguinea DC.). 3,5-, 3-
-3-; . - (ESI/MS) 1- [8] - (A. alnifolia, A. arborea and A. canadensis) -3- (155, 390 165 100
, ). A. alnifolia and A. canadensis
(54 48 /100 ) -3-. [4]
209
______________________________________________________________
, -3-.
, [9] -3- . , , A. ovate ,
[10]: -3-, -3-, -3- .
Amelanchier: A. spicata (Lam.) . Koch, A. alnifolia (Nutt.) Nutt ex M. Roem, - A. sanguinea
(Pursh) DC, A. laevis Weigand, A. canadensis (L.) Medik
A. florida Lindl. .
2011 .
0.1 . 56 510 -3- [11].
- - ( Agilent
1200 Infinity). : : .
510 . 420-600 .
Symmetry 18 4.6250 40. 1 /.
18 .
- Agilent 6130
LC/MS ESI ( )
250-1200.
200 . 4 . - 2 , 10 /, 350,
250 .
, . .1.
(
) 1 5 .1. 3-
, .2. .. 3,5- , ( ) .
( 7.7 7.9 9.1 9.2 ).
210
_______________________________________________________________
.1.
, -
. 2.
[12]
1-3, .3.
-3- (Cy-3-Gala),
-3- (Cy-3-Glu) -3- (Cy-3-Ara)
[13].
( 4 5) (1.003 1.032, ) [14].
(ESI/MS). , ,
, ,
, , -
211
______________________________________________________________
, . , .4, M/z = 287 . 1
2 , -
(M/z = 449). 3 M/z = 419
-3-.
5
4
lgki
0.8
Cy-3-Rut
Cy-3-Ara
Cy-3-Glu
0.4
Cy-3-Gala
0
0
0.25
lgk(Cy-3-Glu)
. 3.
-3-
-3-Gala
-3-Glu
Cy-3-Ara
4. - ESI/MS
, 4 5,
, . -
212
_______________________________________________________________
,
M/z = 287.
4 M/z, 477, .
5 .
, , , , ,
., . , ,
. . . [3] 16 A. alnifolia Nutt. ( ) 19982000 , 414 1 Forestburg 852
1 Nelson.
. -40
, , . (20 ) (0.1%) l (50 ),
, 200
.
, , [4]. ,
( ) 26.7 ( ) 136.8
100 A. alnifolia Nutt. Smoky, .
, ,
, , , .
A. alnifolia [15], ,
, 100 , 100
(?): 196.7 ( Honeywood) 213.6 (
Martin).
, ( ) [5], Amelanchier ovalis Medik, 4 5 % . ( -3-) , 2011 , .
A. spicata (Lam.) . Koch (1)
A. spicata (Lam.) . Koch (2)
A. alnifolia (Nutt.) Nutt. Ex M.Roem (1)
A. alnifolia (Nutt.) Nutt. Ex M.Roem (2)
A. sanguinea (Pursh) DC
A. laevis Weigand
A. canadensis (L.) Medik
A. floribunda Lindl
, /100
0.199
0.129
0.276
0.273
0.102
0.126
0.255
0.249
213
______________________________________________________________
, , (0.250
0.275 100 ), - (0.102 100 ).
20%, 1 % .
, .
(A. alnifolia (Nutt.) Nutt. Ex M.Roem, A. canadensis (L.) Medik
A. floribunda Lindl) , ,
.
1. Handbook of Antioxidants. Ed. Enrique Cadenas, Lester Packer. LLC Marcel Dekker, Taylor & Francis Group, New York, Basel, 2002. 602 .
2. Flavonoids Chemistry, Biochemistry and Applications Ed. yvind M. Andersen, Kenneth
R. Markham, CRC Press, Taylor and Francis Group. 2006 1212 p.
3. Zatylny A.M., Ziehl W.D., St-Pierre R.G. Physicochemical properties of fruit of 16 saskatoon (Amelanchier alnifolia Nutt.) cultivars // n. J. Plant Sci. 2005. V.85. P. 933938.
4. Ozga J.A., Saeed A., Reinecke D.M. Anthocyanins and nutrient components of Saskatoon
fruits (Amelanchier alnifolia Nutt.) // Can. J. Plant Sci. 2006. V.86. P. 193197.
5. .., .., .. // . .-. . 2006. - 3.
. 29-35.
6. .., .., .. Amelanchier Medic. // . . 1982. - 4. . 522-523.
7. .. (Amelanchier Medik.)
// . . . . 2011. - 1. . 52-59.
8. Adhikari D.P., Francis J.A., Schutzki R.E., Chandra A., Nair M.G. Quantification and
characterisation of cyclo-oxygenase and lipid peroxidation inhibitory anthocyanins in fruits of
Amelanchier // Phytochem. Anal. 2005. V.16. P. 175180.
9. Mazza G. Anthocyanins and other phenolics of saskatoon beries Amelanchier alnifolia
Nutt. // J. Food Sci. 1986. V.51. P. 1260-1264.
10. .., .., .., .., .., .. Rosaceae
// . - 2005. - .1. - . 91 -98.
11. Giusti M.M, Ronald E. Wrolstad R.E. Characterization and Measurement of Anthocyanins by UV-Visible Spectroscopy // Current Protocols in Food Analytical Chemistry (2001) F1.2.1F1.2.13.
12. .. //. . . - 2006. - T.80,
3. - . 511-516.
13. .., .., .., .., .. . // . . - 2006. - 3.
- . 16-20.
14. .., ..
// . . . - 2004. - T.78, 5. - .923-926.
15. Guerrero J.C., Ciampi L.P., Castilla A.C., Medel F.S., Schalchli H.S., Hormazabal E.H.,
Bensch E.T., Alberdi M.L. Antioxidant capacity, anthocyanins, and total phenols of wild and cultivated berries in Chile // Chilean J. Agric. Res. 2010. V.70. P. 537-544.
214
_______________________________________________________________
ANTHOCYANINS OF SIX SPECIES AMELANCHIER FRUITS
.N. Chulkov
V.I. Deineka
L.. Deineka
A.V. Stepanova
Belgorod National Research
University, 308015Belgorod,
Pobeda str. 85
e-mail: deineka@bsu.edu.ru
215
______________________________________________________________
616.314.163-08
..
..
..
..
e-mail: deyneka@bsu.edu.ru
,
.
() 1 2
.
40 % .
: , ,
, , .
, , . . .
. . , , , ,
. 40- . , 70% .
40 , () , ,
, ,
,
. ,
[1].
. ,
.
, , ,
. . ,
[2].
.
, .
1% -, , .
24 , (-
216
_______________________________________________________________
100) 3 .
-1, 100,20 400,65 (
() 1 .2 ), , , .
, , . 3
.
- . .
. , . . 1 2.
1
(/)
1
2
3
4
5
6
7
8
9
10
Streptococcus intermedius
Streptococcus sanguis
Streptococcus milleri
Streptococcus mitis
Streptococcus salivarius
Actinomyces naeslundii
Fusobacterium nucleatum
Staphylococcus spp
Staphylococcus aureus
Candida albicans
6,00,2
3,90,1
5,50,2
5,00,2
3,10,1
5,50,2
4,90,2
5,00,2
3,90,1
3,20,1
2,20,1
1,50,1
2,00,1
2,00,1
1,10,1
2,00,1
2,00,1
2,30,1
1,10,1
1,70,1
2
(/)
1
2
3
4
5
6
7
Streptococcus intermedius
Streptococcus sanguis
Streptococcus milleri
Streptococcus mitis
Streptococcus salivarius
Staphylococcus aureus
Candida albicans
5,90,2
10,20,2
7,30,2
5,70,2
6,70,2
3,00,20
3,90,25
2,00,2
4,10,2
4,20,2
2,20,2
3,30,2
1,00,2
1,00,2
, ,
40 % .
.
217
______________________________________________________________
1. . ., . ., . . . // . 1999, 3.
. 29 31.
218
_______________________________________________________________
543.544:637.414
..
..
..
..
..
e-mail: deineka@bsu.edu.ru
-
(Gallus gallus, Pavo cristatus,
Phasianus colchicus, Meleagris gallopavo, Coturnix coturnix, Anser
anser, Cairina moschata, Columba livia Numida meleagris). ,
,
.
- 3--,--3'-.
: , , , ,
.
, , , , [1]. ,
,
[2]. . , ,
,
, 2 ; ,
[3]. [4], () [5]
.
. ,
[6] , , . , , , -, , , ,
, , ,
,
. , - [7] - ,
, , [8], , . , , , , .
- , , -
219
______________________________________________________________
( ) ,
.
.
-, . [9] ,
-, ,
, .
, , , - , ,
. [10], ,
2-6%.
(
)
- . , , ,
, - 1%,
- (7%) --8- (34%).
9-11 ( 3-5 )
-,
. ,
. , , , , (Gallinula chloropus), (Fulica Americana) (Larus
fuscus), (47.5, 131.0 71.6 1 , )
- (25-29%, [11]). [12]
, --8- (
), , . , (
-) ; , . ,
[13], , ( )
[14].
.
, ().
220
_______________________________________________________________
. 0.50-1.00 0.250 10
.
.
(l = 1 ) max = 440 - 445 ( -3-01). ( ) ,
%
E11
2550 :
Amax V P 1000
, /
E11% 100 m l
Amax ,
1%
,
E1
V , ,
,
m , ,
l , .
:
1) Beckman 110B, Rheodyne 7215 20 , : LC/9563
( ). 1.5).
2) Agilent 1200 Infinity - ; ChemStation 32.
,
=-
-
: ~5 . ,
[12].
[15, 16]. , , .1.
,
[17]. ().
( ) .
-
- . ,
, , . ,
, - (
0.45 ). ,
, - ,
221
______________________________________________________________
.
.2 .3.
. 1.
mV
. 2. ,
2504.6 600 (5 ); : 30 .% -;
: 445 .
.2
. . ,
, , ,
.
, ,
. , -
222
_______________________________________________________________
( ) .
, ,
. 7 ,
,
, . ,
, - , .3.
2
3
1+2
. 3.
Aves
,
Phasianinae
,
Phasianidae
,
Meleagridinae
,
Perdicinae
, Anserinae
,
Anatidae
,
Columbidae
,
Numididae
,
Anatinae
,
Columbinae
,
Gallus
,
Pavo
,
Phasianus
,
Meleagris
, Coturnix
. ,
G. gallus
. ,
P. cristatus
. ,
P. colchicus
. ,
M. gallopavo
,
C. coturnix
. ,
A. anser .
caerulescens
,
C. moschata
. ,
A. platyryncha
. ,
C. livia
. ,
N. meleagris
,
Anser
,
Cairina
Anas
,
Columba
,
Numida
Lu/Ze*
**,
/
1.6:1
0.012
2:1
0.025
5.5:1
0.005
5.8:1
0.024
1.7:1
0.011
2:1
0.047
3:1
3.5:1
3.7:1
0.022
6.1:1
0.013
Lu/Ze* , ** - .
223
______________________________________________________________
( ) 70% , , , ,
.
, .3. , , .
,
.
, , - 3--,--3'-. ,
, , ,
[18], [19].
,
. - 3--,--3'-. , , , , .
1. Handelman G.J., Nightingale Z.D., Lichtenstein A.H., Schaefer E.J., Blumberg J.B. Lutein
and zeaxanthin concentrations in plasma after dietary supplementation with egg yolk // Am. J. Clin.
Nutr. 1999. V.70. P. 247251.
2. .., .., .., .., .. : .
// . - 2006. - 6. - . 18-19.
3. .. . .: . 2001. - 88 .
4. 18221-99 .
.
5. 52121-2003 . .
6. . . . - , , .
http://www.roskarfarm.ru/preparations4.
7. http://kemin.biosgroup.com.ua/
8. .., .., .., .., ..
// .
: . - 2010. - 15(86), .12. - . 140-144.
9. .., .., .., .. . , , / . 2010. .79, 2.
- . 4-18.
10. Pizzey H., Bedecarrats G.Y. Study of the effects of dietary lutein on reproductive performances in chickens // J. Poult. Sci. 2007. V.44. - P. 409-415.
11. Surai P.F., Speak B.K., Wood N.A.R., Blount G.R., Spark N.H.C. Carotenoid discrimination by the avian embpyo: a lesson from wild birds // Comp. Biochem. B. 2001. V.128. P. 743-750.
12. Schalatterer J., Breithaupt D.E. Xanthophylls in Commercial Egg Yolks: Quantification
and Identification by HPLC and LC-(APCI)MS Using a C30 Phase // J. Agric. Food Sci. 2006.
V.54. P. 2267-73.
224
_______________________________________________________________
13. Lai S.-M., Gray J.I., Flegal C.J. Deposition of carotenoids in eggs from hens fed diet containing saponified and unsaponified oleoresin paprika // J. Sci. Food Agric. 1996. V.72.
P. 166-170.
14. Nys Y. Dietary carotenoids and egg yolk coloration a review // Arch. Geflgelk. 200.
V.64. P. 45-54.
15. Scott K.J. Detection and measurement of carotenoids by UV/Vis spectrophotometry /
Current Protocol in Analytical Chemistry. 2001. F2.2.1 F2.2.10.
16. Biehler E., Mayer F., Hoffmann L., Krause E., Bohn T. Comparison of 3 Spectrophotometric Methods for Carotenoid Determination in Frequently Consumed Fruits and Vegetables // J.
Food Sci. 2010. V.75. P. C55-C61.
17. Lee C.-Y., Lee B.-D., Na J.-C., An G. Carotenoid Accumulation and Their Antioxidant Activity in Spent Laying Hens as Affected by Polarity and Feeding Period // Asian-Aust. J. Anim. Sci.
2010. V.23. P.799-805.
18. Khachik F., Bernstein P.S., Garland D.L. Identification of Lutein and Zeaxanthin Oxidation Products in Human and Monkey Retinas // Invest. Ophthalmol. Vis. Sci. 1997. V.38. P.
1802-1811.
19. Tyczkowski J.K., Schaeffer J.L., Parkhurst C., Hamilton P.B. 3'-Oxolutein, a metabolite of
lutein in chickens. // Poult. Sci. 1986. V.65. P. 2135-2141.
225
______________________________________________________________
541.6
-
..
..
..
, 308015
., . 85
e-mail: deineka@bsu.edu.ru
c
-. 1- 5- .
+. 3- 4-
.
: , , , , -,
, 1-
[1]. , , , 0.5 1
[2]. - ( ,
, [3]), ( )
. , , [3, 4].
( ) ( ),
, [5].
OH
O
OR5
OR1
OR3
OR4
OH
R=
O
OH
R3
R4
R5
1CQA
R1
R
3CQA
4CQA
5CQA
1.
5- . (6863
SCIRUS),
. [6]
, - (CD) (5CQA)
1:1 424
278 /, . -
226
_______________________________________________________________
,
CD.
- (465 /) [7]. 1H- [8] D2O : 504 936 / , . - , [9] 207 351 ( ) 442 /
. ,
- 1:1. -
,
, 1- . , ( )
. , - [10].
- 5- , - ,
.
,
Beckman 110B, - , Pheodyne 7125 200 . (Nicolet LC/9563). 1.5.
-
1:1 (
), ,
:
+ CD [CD];
(1)
+ CD [CD],
(2)
;
CD -;
[CD] , -
;
[CD] , -
.
(
),
, :
() = 1/(1 + 1[CD] + 2[CD]) = 1/(1 + (1+ 2) [CD]),
(3)
1 K2 1 2, .
227
______________________________________________________________
,
:
k() = k0() = k0/(1 + (1+ 2) [CD]),
(4)
k() CD;
k0 CD .
(4) ,
:
k0/k() = 1 + (1+ 2) [CD].
(5)
- 18. - (80:20 .%) , , -
, .1 .2.
2
3
1
. 1. -
, .2.
mV
2
3
40
20
1
4
10
tR,
. 2. -
1504.6 Kromasil 100-5C18, : 7.5% CH3CN, 2 % CH3COOH ;
1 /. 320 . -: 0; 0.001874;
0.003679 /. : 1 3-, 2 5- 3- 4 , 4 .
228
_______________________________________________________________
5-
. [11-13], :
tR(3CQA) < tR(5CQA) < tR(4CQA).
(6)
.2 ,
- , ; ,
- [14].
(5), .3, -, (R2 0.998), 1:1.
k0/k()
(CD), /
. 3. 5CQA CD
-,
, 7.5 .% 2 .% : 107
12, 163 15 291 10 / 3CQA, 4CQA 5CQA. , , -
, . ,
.1.
1
-
1
2
3
4
, %
CH3CN
5.5
6.5
CH3COOH
5.0
5.0
, /
98 6
72 8
3-
51 6
37 4
4-
81 6
60 4
5-
146 12
116 9
(5.5)/(6.5)
1.36
1.37
1.35
1.26
. , 5CQA
, , .
229
______________________________________________________________
. , 4CQA (
) ,
. 20% , . 3CQA, ,
, ,
- , , ,
.
.
,
,
CD.
,
. 5CQA
6.5% 5.5% ,
4CQA, 3CQA (
), . .1. 5CQA , ( )
: Virtual
Computational Chemistry Laboratory (http://www.vcclab.org/lab/alogps/) ALOGPs
-2.46 1.67, .
, 5QCA ,
- -
-,
HyperChem 8.0 +, .4.
. 4. ) - 5CQA CD:
(1), ) - 5CQA CD: (2); ) - 4CQA CD
230
_______________________________________________________________
CD, .4, , ,
.
5QA CD . 1- ,
5QCA, . D2O -
, , .
-
, .
2, [15].
2H
7H
HO
5 eH
HO
6H
OH
O
O H
6e
3 aH
8H
6 aH
OH H2eH
2a
4 eH
5H
OH
OH
2. 5-
, .2, - .
,
. 5CQA -: 2, 6 5,
5CQA (2). , .3, ,
.
2
5CQA
2
5
6
7
8
D2O
7.1918
6.9760
7.1240
7.6089
6.3525
+ CD
7.2089
6.9722
7.1200
7.7078
6.4398
+
7.2607
7.0093
7.1894
7.6882
6.4349
.
+0.0518 / +0.0171
+0.0371 / -0.0038
+0.0694 / -0.0040
-0.0196 / +0.0989
-0.0049 / +0.0873
3
2
5
6
7
8
D2O
7.2124
6.9903
7.1494
7.6342
6.3683
+ CD
7.1473
6.9959
7.0851
7.6172
6.3133
+
7.2525
7.0050
7.1861
7.6691
6.4099
.
+0.1052 / -0.0651
+0.0091 / +0.0056
+0.1010 / -0.0643
+0.0519 / -0.0170
+0.0966 / -0.0550
231
______________________________________________________________
,
5CQA - -. - - 3, 5 ( 4 CD ), , .5.
, ..
3
1
2
.
.
.
0
5 5-
. 05. 1-
D2O, , -
5CQA CD .
,
, .
1. .., .., ..
// .
2010. - .2. . 91-97.
2. Olthof M.R., Hollman P.C.H., Katan M.B. Chlorogenic Acid and Caffeic Acid Are Absorbed
in Humans // J. Nutr. 2001. V.131. P. 6671.
3. Olthof M.R., Hollman P.C., Zock P.L., Katan M.B. Consumption of high doses of
chlorogenic acid, present in coffee, or of black tea increases plasma total homocysteine
concentrations in humans // Am. J. Clin. Nutr. 2001. V.73. P. 532538.
4. Monteiro M., Farah A., Perrone D., Trugo L.C., Donangelo C. Chlorogenic Acid
Compounds from Coffee Are Differentially Absorbed and Metabolized in Humans // J. Nutr. 2007.
V.137. P. 21962201.
5. Lin L.-Z., Harnly J.M. Identification of Hydroxycinnamoylquinic Acids of Arnica Flowers
and Burdock Roots Using a Standardized LC-DAD-ESI/MS Profiling Method // J. Agric. Food
Chem. 2008. V.56. P. 10105-10114.
6. Grnas P., Neunert G., Baczyski K., Polewski K., Beta-cyclodextrin complexes with
chlorogenic and caffeic acids from coffee brew: Spectroscopic, thermodynamic and molecular
modelling study // Food Chemistry. 2009. V.114. P. 190196.
7. Alvarez-Parrilla E., de la Rosa L.A., Torresrivas F., Rodrigo-Garcia J., Gonzalez-Aguilar
G.A. Complexation of Apple Antioxidants: Chlorogenic Acid, Quercetin and Rutin by -Cyclodextrin
(-CD) // J. Incl. Phenom. Macrocycl. Chem. 2005. V.53. P. 121129.
232
_______________________________________________________________
8. Rodrigues E., Vazi S., Gil V.M.S.S., Caldeira M.M., da Silva A.M.G.M. Inclusion of
Polyphenol Oxidase Substrates in -Cyclodextrin: A 1H-NMR Study // J. Incl. Phenom. Macrocycl.
Chem. 2002. V.44. P. 395397.
9. lvarez-Parrilla E., Palos R., de la Rosa L.A., Frontana-Uribe B.A., Gonzlez-Aguilar G.A.,
Machi L., Ayala-Zavala J.F. Formation of Two 1:1 Chlorogenic Acid: -cyclodextrin Complexes at pH
5: Spectroscopic, Thermodynamic and Voltammetric study // J. Mex. Chem. Soc. 2010. V.54(2).
P. 103-110.
10. Zhao M., Wang H., Yang B., Tao H. Identification of cyclodextrin inclusion complex of
chlorogenic acid and its antimicrobial activity // Food Chem. V.120. P. 1138-1142.
11. Maldini M., Montoro P., Kapusta I., Oleszek W., Piacente S., Pizza C. Quantitative
Analysis of Caffeoylquinic Acids and Styrylpyrones in Sweetia panamensis Bark by UPLC //
Chromatographia. 2009. V.70. P. 16211626.
12. Clifford M.N., Zheng W., Kuhneret N. Profiling the Chlorogenic Acids of Aster by HPLC
MSn // Phytochem. Anal. 2006. V.17. P. 384393
13. Cliffford M.N., Johnston K.L., Knight S., Kuhnert N. Hierarchical Scheme for LC-MSn
Identification of Chlorogenic Acids M.N. // J. Agric. Food Chem. - 2003. - V.51. - P. 29002911.
14. .., .., ..
// . 2007. .81, 7. . 1221-1226.
15. Tolonen A., Joutsamo T., Mattlla S., Kmrinen T., Jalonen J. Identification of isomeric
dicaffeoylquinic acids from Eleutherococcus senticosus using HPLC-ESI/TOF/MS and 1H-NMR
methods // Phytochem. Anal. 2002. V.13. P. 316-328.
233
______________________________________________________________
543.54:547.973
,
.., ..
..
.. , ..
, 308015 .,
. 85.
e-mail: deineka@bsu.edu.ru
(Caprifoliaceae), 15
500 , , , [1]. (Lonicera)
180 ,
L. tatarica L. ( )
L. xylosteum L. (Lonicera edulis Turcz. ex Freyn). , (L. caerulea L.), ,
- : (L. kamtschatica (Sevast.) Pojark.)), (L. edulis
Turcz.), (L. turczaninowii Pojark.), (L. regeliana Bochkarn.), (L. altaica Pall.).
.
, 20 .
, . .
0.6251.736 /100 ,
[2]. 0.116 0.339 100 [3].
(0.235 100 ) [4]. 1.081 100 -3 [5]. 0.116-0.593 /100 [6]. , , 0.186
0.961 100 [7].
, , ,
, .
, -
234
_______________________________________________________________
, . .
0.1 HCl
, 98%- .
-56
- .
. -3-, , = 26900 /() [8].
01- 1,3 -
( ..).
0,0022 /. 1,2 3/.
Agilent 1200 Infinity
- . : 2504.6 Symmetry C18, 5 ; 40. : 8 .% 10 .%
, 1 /.
(30 ),
2005 , ,
0.104 ( ) 1,064 ( 3) 100 , 2/3 0.500 .
, , . , , 1 100
. ,
, .
: , ,
0,329 (2006 .) 0,300 (2007 .) 100 ( , 1 ) 0,734
, .
, (2005 .),
: ,
- 0.522 (2006) 0.215 (2007 .); 0,340 0,255 100 .
0,677 (2005 .), 0,337 (2006 .) 0,240 (2007 .) 100 . , , , , ,
.
( ) .
2011
,
.
0.1
HCl ,
235
______________________________________________________________
(0,738 0,025 100 ) , (0,422 0,034 100 .). (. 1).
. 1. ,
(1) (2)
,
: [9], : -3 ( 2); - -3- ( 4)
-3,5- ( 1), ,
3- ( 5) ( 6), . 2.
2
4
6
3
. 2. Lonicera caerulea L.
236
_______________________________________________________________
, -3,5-
-3-
2 , 420 460 .
3 , ,
2 , . , ,
, .
2
5
4
1
. 3.
.2: 1 3,5-; 2 3-, 3 3-
(3-) 5 -3-
,
.
( ) --01.
, [9], [10] , . ,
TROLOX ( 6--2,5,7,8--2- ), [11], , - , .
( ), , ,
. 0.1
0,0022 .
, , ,
13.1 1.5 100 , ,
6.3 0.8 ( 100 ),
1150 550 100 , . ,
, , -
237
______________________________________________________________
[2] . , ,
, [12].
, , ,
,
, .
.
1. .. (Caprifoliaceae) /
. . . . .. , .: , 1981. . 375.
2. Rop O., eznek V., Mlek J., Jurkov T., Balk J., Sochor J., Kramov D. Antioxidant
and radical oxygen species scavenging activities of 12 cultivars of blue honeysuckle fruit // Hort. Sci.
(Prague). 2011. - V.38. P. 6370.
3. Encyclopedia of fruit and nuts. Ed. J. Janick and R.E. Paull. CAB International 2008. P. 233.
4. Ochimian I., Grajkowski J., Skupien K. Yield and Chemical Composition of Blue Honeysuckle
Fruit Depending on Ripening Time // Bulletin UASVM Horticulture. 2010. V.67. P. 138-147.
5. Bakowska-Barczak A.M., Marianchuk M., Kolodziejczyk P. Survey of bioactive components in Western Canadian berries // Can. J. Physiol. Pharmacol. 2007. V.85. P. 1139-1152.
6. Chaovanalikit A. Anthocyanins, total phenolics, and antioxidant capacity of bluehoneysuckles. In Abstracts of The 30th Congress on Science and Techonology of Thailand, 19-21 October
2004. Impact Exhibition and Convention Center, Muang Thang Thani, Bangkok, Thailand.
http://www.scisoc.or.th/stt/30/sec_c/paper/stt30_C0213.pdf
7. .., .., .., ..
Lonicera caerulea L.
// . . . 2011. - 3. C. 39-45.
8. Giusti M.M, Ronald E. Wrolstad R.E. Characterization and Measurement of Anthocyanins
by UV-Visible Spectroscopy // Current Protocols in Food Analytical Chemistry (2001) F1.2.1-F1.2.13.
9. .. -
// . . .
(. . . - . .. ). 2008. T.5. C. 130-135.
10. .., .., .., .., .. :
// 2010 - 9(80). C. 104-111.
11. Huang D., Ou B., Prior R.L. The Chemistry behind Antioxidant Capacity Assays //
J. Agric. Food Chem. 2005. V.53. P. 1841-1856.
12. Chaovanalikit A., Thompson M.M., Wrolstad R.E. Characterization and Quantification of
Anthocyanins and Polyphenolics in Blue Honeysuckle (Lonicera caerulea L.) // J. Agric. Food
Chem. 2004. V.52. P. 848-852.
238
_______________________________________________________________
543.635:634.72
RIBES L.1
..
..
..
..
..
..
..
e-mail: deineka@bsu.edu.ru
R. nigrum
. , ( 100 280 %
).
( 67 345 100 ) , , -3-,
-3- . ,
, Ribes alpinum , (Ribes aureum) -. ,
, (- -) () .
: Ribes, ,
, , , , .
-
2009-2013 ., 508 14.05.2010
.
239
______________________________________________________________
2,13,2%, 3,9
5,4%, 1,52,0% [9].
. , , [10]. , . , ,
, , 98-100%
[11].
. , (. 1).
OH
HO
OH
OH
HO
F3'H
OH
OH O
OH
DFR
ANS
OH
3GT
HO
OH
F3H
OH
HO
HO
OH
OH
HO
OGlu
HO
OCH3
OH
OH
HO
OGlu
OH
-3-, Pn3G
ANS
3GT
+
O
HO
OCH3
OGlu
OH
OH
OGlu
OH
-3-, Pg3G
OCH3
OH
OH
OH O
OH
DFR
OGlu
OH
-3-, C3G
MT
F3',5'H
OH
O
DFR
ANS
3GT
+
O
HO
OH
OH
MT
OH
-3-, M 3G
OH
-3-, D3G
MT
O
OCH3
OH
OH
OGlu
OH
-3-, Pt3G
. 1.
-3- (F3H) , , -
240
_______________________________________________________________
, -
( F3H F35H).
-4- (DFR), (ANS) 3- (3GT) , [12]. 3-
(MT) ( ),
( ).
, , ,
-6-, 3- 3- .
: 270 /100 [13],
83-199 /100 17 [14], 300 /100
[15]. ,
(23-127 % ) [16].
[17]. ,
- , . ,
, , , . . , (, ), [18]. .
[19],
.
, . ,
-3 (15,0%
) [20]. [21],
Ribes 30,5%.
19% - , ,
, , .
38 R. nigrum,
. , .
, -
( ). - - .
-3-01.
241
______________________________________________________________
0.1 [22].
3 - 5 .
,
0.4 . . .
,
, .
,
Altex 110A Beckman 110B, Rheodyne 7125
20 : Nicolet LC/9563
R401 Waters Millipore.
: 2504,0 , -110-18, 5 (, ); 2504,0 ,
Reprosil-Pur C18-AQ, 5 (. ); 2504,6 , 18.
1,5
(Ampersand ltd).
: (...
-1), (... -205, ), (..), (..).
, .. .
2. , R. nigrum
, . , .. [23], , .
3
mV
4
2
200
100
1
0
0
10
. 2. R. nigrum L.
1 - , 2 ,3 ,
4
242
_______________________________________________________________
R. nigrum ( ), , 1,59-2,91%,
2,23%, ,
[23], .
, .
J.G Bordonaba L.A. Terry [24]
40,2 81,5% , 2,6 7,4%.
( ) R. nigrum (
Eucoreosma) 79 312 % (. 1).
1
R. nigrum
6-26-70
6-21-103
6-10-42
, %
129,022,7
117,85,4
133,66,4
, %
123,623,9
281,032,8
211,623,5
168,917,0
109,912,8
100,311,3
167,414,9
215,721,6
123,623,9
133,910,7
226,713,6
161,68,1
142,912,9
175,019,2
210,416,8
145,919,7
129,313,0
166,013,3
c
234,218,7
, %, 183,227,9
225,521,5
148,011,8
156,918,8
264,821,2
117,311,7
142,09,4
168,113,4
161,313,8
131,79,9
112,610,0
151,211,9
253,721,1
179,014,3
148,911,6
167,720,8
214,412,3
( 51%) ( 8%),
( 27 36%) (. 3).
() , ,
30% , [25].
.
R. nigrum , . , ,
: -3- (D3G) -3 (D3R) -3- (3G) -3- (C3R).
. 4 .
243
______________________________________________________________
, %
250,0
200,0
1
150,0
100,0
50,0
0,0
1
2
3
4
5
6
( 7 .)
. 3. R. nigrum
: 1 ; 2 .
mV
4
1
200
100
0
ch
01
5
10
. 4. :
- ; ; - ; ; 1 D3G; 2 D3R; 3 C3G; 4 C3R.
R. nigrum
,
(. 2).
R. nigrum , [24] [26], ,
[25] [15] .
R. nigrum,
-
244
_______________________________________________________________
155 240 100 . R. nigrum ssp. sibiricum
120-160 100 .
R. nigrum
*, %
D3G
D3R
C3G
C3R
4,8
62,8
2,4
30,0
5,0
67,3
2,6
25,1
7,7
54,0
5,5
32,8
9,7
49,2
6,1
35,0
11,8
58,9
6,5
22,8
2005
163
102
194
196
145
**, /100
2006
2007
2008
245
256
325
247 76
92
131
194
130 64
126
143
159
156 38
139
142
155
158 34
215
197
234
198 36
10,0
66,3
2,6
21,1
206
242
184
252
221 31
5,5
6,6
5,9
9,9
63,7
62,8
60,1
50,0
2,4
3,3
4,7
7,6
28,4
27,3
29,3
32,5
188
128
97
176
252
118
58
270
259
298
145
235
345
292
164
311
261 84
209 89
116 29
248 62
9,9
31,8
11,1
46,2
184
214
199
252
212 40
7,7
9,6
8,2
12,9
10,1
9,8
47,8
62,1
54,9
57,9
50,1
59,5
3,9
4,4
7,7
4,1
4,8
4,1
40,6
23,9
29,2
25,1
34,7
26,6
91
118
180
157
68
147
77
183
183
142
67
249
120
133
197
175
92
208
168
204
201
219
76
306
114 52
160 44
190 11
173 46
76 16
228 58
* - ; ** -3-.
- ,
. - , ; ,
.5.
.
190
,
%
150
200
200
110
70
4
(7.)
. 5. R. Nigrum 2007-2008 .
( )
245
______________________________________________________________
, , .
R. aureum
, -,
(. 6).
mV
3
50
2
1
0
0
10
5
. 6.
( -) 8 19 %
.
.. [27] 5 12 %
.
, , , 4 .
, . Ribes -.
-, , R. nigrum 1,3 %,
R. aureum 3,7 7,6 % (
-), R. alpinum 3,3 4,4 % ( ),
Ribesia 0,5 2,0 %.
.
Ribes :
, .. (
- -) . 8-14 22, (16) 18 ,
.
(. 7).
246
_______________________________________________________________
,
Ribes (R. nigrum R. alpinum), , - - (11-16%
), - ( (3-6%
), [28,29]. [11] ,
(
) -
- ( 300 ), 210 .
mV
20
10
0
0
10
20
30
, , .
1. .. (Grossulariaceae) / . . .. . . . .: , 1981. . 169.
2. .. . .: , 1981. 510 .
3. .. , , . : . . -, 1992. 384 .
4. http://www.apte.ru/product/Terra-plant_ChERNAYa_SMORODINA
5. Karjalainen R., Anttonen M., Saviranta N., Hilz H., Srewart D., Mcdougall G.J., Mattila P.,
Trrnene R. A Review on Bioactive Compounds in Black Currants (Ribes nigrum L.) and Their Potential Health-Promoting Properties // Acta horticulturae. 2009. N.839. P. 301-307.
6. .., ..
// . 2010. - 1. - . 13-16.
7. .., .. // . - 2010. - 2. - . 9-12.
8. .. // . . 1961. .6. .153-164.
247
______________________________________________________________
9. .., .., ..
// . 1999. 9. . 64-67.
10. ..
: . . .
. : 05.18.15 / . - . -. ., 1971. 26.
11. .. // . 2003. 2. . 37-39.
12. Jaakola L., Maata K., Pirtilla A.M., Torronen R., Karenlampi S., Hohtola A. Expression of
Genes Involved in Anthocyanin Biosynthesis in Relation to Anthocyanin, Proanthocyanidin, and Flavonol Levels during Bilberry Fruit Development // Plant Physiology. 2002. V. 130. P. 729739.
13. Antal D.-S., Grban G., Grban Z. The anthocyans: biologically-active substances of food
and pharmaceutic interest // The Annals of the University Dunarea de Jos of Galati Fascicle VI
Food Technology. - 2003. P. 106-115.
14. Bordonaba J.G., Terry L.A. Biochemical Profiling and Chemometric Analysis of Seventeen UK-Grown Black Currant Cultivars // J. Agric. Food Chem. 2008. V.56. P. 7422-7430.
15. Maatta K., Kamal-Eldin A., Trrnen R. Phenolic compounds in berries of black, red,
green, and white currants (Ribes sp.) // Antiox. Redox Signal. - 2001. V. 3 (6). P. 981-993.
16. .. . .: . , 2008. 320 .
17. Takashi S., Furihata K., Ishidsu J., Shimada K. Carotenoids sulphates from the aerobic
photosynthetic bacterium Erytrobacteria logus // Phynochemestry. 1991. V. 30. - P. 3411-3415.
18. .., .., .., .., ..
// . - 2005. - 6.
- . 46-48.
19. Velasco L., Goffman F.D. Chemosystematic significance of fatty acids and tocopherols in
Boraginaceae // Phytochem. 1999. V. 52. P. 244-345.
20. Vecera R., Skottov N., Vna P., Kazdov L., Chmela Z., Svagera Z., Walter D., Ulrichov
J., Simnek V. Antioxidant status, lipoprotein profile and liver lipids in rats fed on high-cholesterol diet
containing currant oil rich in n-3 and n-6 polyunsaturated fatty acids // Physiol Res. 2003. V. 52
(2). P. 177-187.
21. Traitler H., Winter H., Richli U., Ingenbleek Y. Characterization of gamma-linolenic acid
in Ribes seed // Lipids. 1984. V. 19(12). P. 923-928.
22. .., .., .., .., .. // . . 2006. . 72. 3. . 16-20.
23. i .. i i // . 2007. 1(6). . 8-14.
24. Bordonaba J.G., Terry L.A. Biochemical Profiling and Chemometric Analysis of Seventeen UK-Grown Black Currant Cultivars // J. Agric. Food Chem. 2008. V.56. P. 7422-7430.
25. Rubinskiene M., Viskelis P., Jasutiene I., Duchovskis P., Bobinas C. Changes in biologically active constituents during ripening in black currants // J. Fruit Ornam. Plant Res. 2006. V.
14. P. 237-246.
26. Kampuss K., Strautina S. Evaluation of blackcurrant genetic resources for sustainable
prodaction // J. Fruit Ornam. Plant Res. - 2004. V. 12. . 147-158.
27. .., .. // : .-. ., 110- .. . , 2007. . 294-300.
28. Barre D.E. Potential of evening primrose, borage, black currant, and fungal oil in human
health // Ann. Nutr. Metab. - 2001. V.45. P. 47-57.
29. Vecera R., Skottov N., Vna P., Kazdov L., Chmela Z., Svagera Z., Walter D., Ulrichov
J., Simnek V. Antioxidant status, lipoprotein profile and liver lipids in rats fed on high-cholesterol diet
containing currant oil rich in n-3 and n-6 polyunsaturated fatty acids // Physiol Res. 2003.
V. 52 (2). P. 177-187.
248
_______________________________________________________________
BIOLOGICALLY ACTIVE SUBSTANCES OF RIBES L. FRUITS
.I. Shaposhnik
L.. Deineka
V.N. Sorokopudov
V.I. Deineka
J.V. Burmenko
V.V. Kartushinskiy
A.V. Tregubov
Belgorod State National
Research University
e-mail: deineka@bsu.edu.ru
A review of some biologically active compounds in fruits of black currants of BSU Botanical garden collection is presented. The main fruits carboxylic acids were citric, ascorbic and malic. Ascorbic acid accumulation as
well as the dynamics belongs upon the varieties ranging from 100 to 280
mg%. The anthocyanin accumulation level varied from 65 to 345 mg per 100
g of fresh fruits between varieties and years of harvesting the four main components being the main for the all samles under investigation delphinidin3-glucoside, delphinidin-3-rutinoside, cyanidin-3-glucoside and cyanidin -3rutinoside. It has been pointed out that some another Ribes genus accumulate high level of -carotene (R. aureum) or lycopene (R. alpinum). The seed
oil of the species was composed mainly with C18 acids, including - and linilenic as well as stearidonic acid.
Key words: Ribes, carboxylic acids, ascorbic acid, anthocyanins, carotenoids, seed oil, HPLC, spectrophotometry.
249
______________________________________________________________
543.555.4+543.556.4+543.544.5.068.7
,
..
.
..
..
..
e-mail: deineka@bsu.edu.ru
(, )
( )
, .
.
: , , , , , ,
, .
, , . [1].
,
.
, .. [2]. (, , , ) ( [3]), , , .
, ,
. , . , . , .
. , 28
1999 . 21,
:
. , ( ), /, D- , L- . ( = 8,1), , D- , /
( ), L- , /, ,
, , , /, , /, , /,
.
250
_______________________________________________________________
. :
.. . ,
.
.
. ,
[4, 7],
[6].
,
() . , [7], . , ,
. , 3, . , ,
, . ().
, , , . , , , (330)
.
(338), , , .
. , . , . , : 8.9 [8]; 8.30 0.01 [9;10]; 8,2 [11]; 8,8
[4]. , ( ) [12].
,
( = 8.1), , .
, [13 16], [17].
: -10601/7 (
-150) / -1 ( 4100) -
251
______________________________________________________________
. , , ( ).
.
( 3 30 ) 50 200 ; , , .
( ).
- ( )
[18].
, ,
.
, ,
.
.
.
, ,
[19, 20], . 1.
1
/100
1.15*
1.43*
0.83*
1.15*
1.19**
1.64**
0.5**
0.78**
0.003-0.0035**
0.5**
0.008**
0.012**
0.008**
0.014**
2.7*
3.2*
0.23***
0.48***
0.96*
1.04*
1.44**
1.58**
0.22-0.76**
0.45-0.97**
20
28
27
36
39-43
32
41
17
52
7.7
7.1
23-51
3.1
2.84
2.05
2.05
2.03-2.05
2.5
2.3
2.8
2
3.1
2.2
2
* ; ** ; ***
- , 1.
. 2 . ,
252
_______________________________________________________________
(, ),
- . ,
, , ,
.
2
mV
20
0
10
tR,
. 1
1 , 2 , 2504 . Reprosil-pur C18-AQ 2 .%
(205 ); , ; ; ; ;
; ; .
, , , 2, , . , ,
.
, . 2, .
,
, -
253
______________________________________________________________
,
, .
2
*
*
*
**
**
4 () **
() **
() **
() **
() **
() **
,
/
10.2
8.5
16.3
,
/
12.8
12.1
20.3
3.0
2.8
2.8
4.5
6.5
2.0
5.1
6.9
2.1
3.1
4.1
6.6
5.0
6.3
6.3
5.4
6.9
9.1
7.9
7.9
8.5
2.1
2.5
2.1
2.1
2.1
2.2
* ; ** .
(. 2).
1
mV
2
3
5
4
1200
800
400
5
. 2.
1 , 2 , 3 , 4 , 5 ( ), 6 . : , , , - , .
, , . , -
254
_______________________________________________________________
, , .
, , . , . [21] , , , 140-1500 .
, ,
.
,
, .. 100 %- .
,
. ,
.
(CocaCola, Pepsi) ,
. , . 3.
3
( 2), /*
**, /
, /
Pepsi light
2.10
0.67
0.93
Pepsi
1.67
0.67
0.77
Coca-cola light
2.04
0.63
1.10
Coca-cola
1.99
0.68
0.98
Coca-cola express
1.11
0.67
0.77
* - ; ** - 2.
: 2.41 3
2.17 2.34 .
, ,
(IV), . , 2
, . 4.
4
2
, 2
Pepsi light
Pepsi
Coca-cola light
Coca-cola
Coca-cola
express
2
42.8
34.5
41.7
40.7
22.7
. /
, 1*
, 2*
, 1**
, 2**
2
4.12
14.7
10.3
19.6
5.41
13.6
7.73
15.7
4.12
13.9
14.4
22.7
4.89
13.9
11.9
22.7
5.92
13.9
8.24
15.7
1.90
2.03
1.58
1.91
25.0
13.4
38.8
38.8
1.91
11.5
:
( 2 :
255
______________________________________________________________
, 2 * *
1 **
( 2 * * 1 * *)
100,%
2 **
2 -
, , - . ( ) .
, Pepsi light, Pepsi, Coca-cola, Coca-cola express
, .. , 2. Coca-cola
light ( = 1.6),
. Coca-cola
Coca-cola light (
).
, , ( [22, 23])
,
, .
1. .., .. - - , - - 2007 46 .
2. ., ., . . .: . 1966. 512 .
3. Munir E., Yoon J.J., Tokimatsu T., Hattori T., Shimada M. A physiological role for oxalic
acid biosynthesis in the wood-rotting basidiomycete Fomitopsis palustris // Proc. Nat. Acad. Sci.
2001. V.98. P. 1112611130.
4 Gomis B.D., Gutierrez M.J., Alvarez G.M.D., Medel S.A. HighPerformance Liquid Chromatographic Determination of Major Organic Acids in Apple Juices and Ciders // Chromatographia.
- 1987. - V.24. - P. 347-350.
5. Gorsel H., Li C., Kerbel E., Smits M., Kader A. Compositional Characterization of Prune
Juice // J. Agric. Food. Chem. 1992 V.40. P. 784-789.
6. Gomis D., Gutierrez M., Alvarez M., Alonso J. Application of HPLC to Characterization
and Control of Individual Acids in Apple Extracts and Ciders // Chromatographia 1988. V.25.
. 1054-1058.
7. Ryan J., Dupont J. Identification and Analysis of the Major Acids from Fruit Juices and
Wines // J. Agric. Food. Chem. 1973 V.21 1 P. 45-49.
8. 3624-92. . .
9. 51455-99. . .
10. Thangavelu S., Chiranjivi Rao K. Titratable acidity of juice of sugarcane genetic stocks
and its association with other characters// J. Indian Sugar. 1996. V.46. P. 391-396.
11. Ayala-Zavala J.F., Wang S.Y., Wang C.Y., Gonzlez-Aguilar G.A. Methyl jasmonate in
conjunction with ethanol treatment increases antioxidant capacity, volatile compounds and postharvest life of strawberry fruit // Eur. Food Res. Technol. 2005. V.221. P. 731738.
12. 5898-87. . .
13. Badoud R., Pratz G. Improved high-performance liquid chromatographic analysis of
some carboxylic acids in food and beverages as their p-nitrobenzyl esters // J. Chromatogr. .
1986. V.360. P. 119136.
256
_______________________________________________________________
14. Yalcin D., Ozcalik O., Altiok E., Bayraktar O. haracterization and recovery of tartaric acid from wastes of wine and grape juice industries // J. Therm. Anal. Calorimetry. 2008. - V.94.
P. 767771.
15. Marce R.M., Calull M., Manchobas R.M., Borrull F., Rius F.X. An Optimized Direct Method for the Determination of Carboxylic Acids in Beverages by HPLC // Chromatographia. 1990. V.29. - P. 54-58.
16. Marconi O., Floridi S., Montanari L. Organic acids profile in tomato juice by HPLC with
UV detection // J. Food Quality. 2007. V.30. P. 4356.
17. Miwa H.J. High-performance liquid chromatographic determination of mono-, poly- and
hydroxycarboxylic acids in foods and beverages as their 2- nitrophenylhydrazides // J. Chromatogr.
A. 2000. V.881. P. 365385.
18. Cunha S.C., Fernandes J.O., Ferreira I. HPLC/UV determination of organic acids in fruit
juices and nectars // Eur. Food Res. Technol. 2002. V.214. P. 6771.
19. .., .., .., ..
//
. . . 2008. - .63, 8. .852-857.
20. .. .- .: , 2004.
528 .
21. Gurrieri F., Audergon J., Albagnac G., Reich M. Soluble sugars and carboxylic acids in
ripe apricot fruit as parameters for distinguishing different cultivars// Euphytica. - 2001 V.117
P. 183-189.
22. Zadernowski R., Naczk M., Nesterowicz J. Phenolic acid profiles in some small berries //
J. Agric. Food Chem. 2005. V.53. P. 2118-2124.
23. Pako P, Sajewicz M., Gorinstein S., Zachwieja Z. Analysis of selected phenolic acids and
flavonoids in Amaranthus Cruentus and Chenopodium Quinoa seeds and sprouts by HPLC // Acta
chromatographica. -2008. V.20. P. 661672.
257
______________________________________________________________
543.422.3:686.862.5
..
..
e-mail: shaposhnik@bsu.edu.ru
22 , 9
10 , , , , , , ,
, . ,
,
. , ,
.
: ,
,
,
,
-.
, 12 15 , 2010 .
, , -
. , , (, ) ,
.
,
, , ,
.
, , - [1]. .
, , ,
[2]. ,
, : ,
. , , , .
[3]: , , ,
, ( ), , ,
.
, . -
258
_______________________________________________________________
, , . - .
, . [4],
[5] - , , .
, ,
, , [6]. .
.
. ,
-.
22 , 9 , 10 (, , , , , , , ), .
,
(
)
. ,
, , , ,
.
-56 ( ) 400 750 . 65% (.) .
, -56 210
240 .
. , .
.
, ,
. . ,
(pH = 2, pH=3, pH=4, pH=5), ,
.
259
______________________________________________________________
,
.
, , .
, , ().
.
, -56 ( )
400 750 .
, 300 650 .
(. 1).
1
5
. 1.
1 - Cello Maxriter (); 2 - Tukzar (); 3 - Berkly (); 4 - BIEFA ();
5 - Cello Pin Point ()
, .
(Cello
Maxriter), (STABILO 030 XF), (Unix 2001 TC POINT)
(BIEFA) -56 300
600 (. 2). 4 ,
,
max=347 , 458 max=528 . -
260
_______________________________________________________________
STABILO 030 XF
, 1 ,
458 . ,
max=350 , 520
max=557 .
(BIEFA), .
(max) 541 .
,
(65% (.) ) -56 - 210 250 .
2
3
. 2. :
1 - Cello Maxriter (); 2 - STABILO 030 XF (); 3 - Unix 2001 TC POINT;
4 - BIEFA ()
(. 3, 4), 3 :
1 : Corvina () 217 ;
2 : Cello Maxriter (), Erich Krause (), Parm (), Stabilo
(), Pensan () 218 ;
3 : (), Stabilo (), ( ).
261
______________________________________________________________
2
1
4
. 3.
:
1 - Cello Maxriter (); 2 - (); 3 - Erich Krause (); 4 - Parm ()
2
1
. 4.
:
1 Pensan (); 2 - Corvina (); 3 Stabilo (); 4 - ( )
,
Corvina (),
.
, 2 .
, 217, 218 219 ,
- .
262
_______________________________________________________________
4
5
. 5.
:
1 - Pensan My Pen (); 2 - Berkly Delta (); 3 - Erich Krause Megapolis ();
4 - Cello Maxriter (); 5 - Corvina ().
() (.5,6), 7 :
1 : Pensan My Pen () 221 ;
2 : BOOM () 222 ;
3 : Stabilo () 223 ;
4 : Erich Krause Megapolis () 224 ;
5 : Berkly Delta () 225 ;
6 : Cello Maxriter (), PILOT () 228 ;
7 : Corvina (), KOOH-I-NOOR () 229 .
,
(Pensan My Pen (), BOOM (), Stabilo (), Erich Krause Megapolis
(), Berkly Delta ())
,
. Cello Maxriter (), PILOT
(), Corvina (), KOOH-I-NOOR () ,
. , 221-225, 228 229 , - .
-
, .
,
, , 5 9 , 1 9
.
263
______________________________________________________________
. 6.
:
1 - Stabilo (); 2 - BOOM (); 3 - PILOT (); 4 - KOOH-I-NOOR ().
.
, ,
.
1. .., .., .. . , . - .:
, 1987. - 96 .
2. .. , // . - .: , 1966. - 16-17. - .39-41.
3. .., .., .. . // . -.: , 1993. - 122. - .12.
4. .. // : . . . .. .- :
. , 2006. - .225-229.
5. Djavanshir Djozan, Tahmineh Baheri,Ghader Karimian, Masomeh Shahidi Forensic discrimination of blue ballpoint pen inks based on thin layer chromatography and image analysis// Forensic Science, 2008. - Int.179. - pp 199-205.
6. ..
// . - .: , 1993. - 38..86-91.
264
_______________________________________________________________
POSSIBILITIES OF SPECTROPHOTOMETRY RESEARCH OF BALLPOINT PEN INKS
OF MODERN MANUFACTURERS
E.I. Shaposhnik
I.G. Evtushenko
Belgorod National Research
University
e-mail: shaposhnik@bsu.edu.ru
265
______________________________________________________________
581.9:582.734.3
CRATAEGUS L.
..
..
..
, ,
. , . , 85.
e-mail: baks922@rambler.ru, sorokopudov@bsu.edu.ru, irina.navalneva@mail.ru
, ,
Crataegus L., .
,
.
: , , , , , , , .
,
, .
- .
( ). -
, , , , .
, - . 3 %
1000 % .
: -. ,
, , . : , ,
.
- [1].
, , , . , ,
. - , , . [2, 3].
() ,
. ; , , . .
(E163). [4].
, - .
266
_______________________________________________________________
, , , , ,
.
. , , . 4 11 % , , , (, ), ,
, , , , , , . 0,26 0,93 %. , .
: 31-108 % ( 257,3), 330-680 %, 2-4 % ( 75). , , [5, 6].
, ,
, , . . 200 . ,
.
, . ,
, . , [6].
, .
20 Crataegus L.: C. arnoldiana (Sarg.), C. canadensis (Sarg.), C. chlorosarea (Maxim.), C. crusgalli (L.), C. douglasii (Lindl.), C. flabellata ((Spach) Kirchn.), C. holmesiana (Ashe), C.
lumaria (Ashe), C. maximowiczii (C.K.Schneid.), C. monogyna (Jacq.), C. pentagyna
(Waldst. & Kit. ex Willd.), C. persimilis (Sarg.), C. pringlei (Sarg.), C. punctata (Jacq.), C.
rotundifolia (Moench), C. sanguinea (Pall.), C. turkestanica (Pojark.), C. ellwangeriana
(Sarg.), C. submollis (Sarg.), C. macracantha (Lodd). 7-8
18 (90%), . 2009 . 13. , 2009 ,
(-18).
2010 . .
:
1.
2,6-.
. 2,6-
( ) , , - [7, 8].
2. (II) (I) .
( ) .
, Cu2+,
[2, 9].
3. ( ) (
267
______________________________________________________________
-102)
- [1].
4. -102. 0,1 [4].
5. .
105 ( 28561-90) [10].
Crataegus 2009 . ,
72,052,65 %, : C. deksiflora, C. crus-galli. : C. maximowiczii (64,931,75 %) C. arnoldiana
(65,931,78 %). C.
macracantha (80,962,19 %).
, 2010 ,
,
, C. arnoldiana 3,5
(231,477,35 %). C. flabellate
(233,167,4 %), C. crus-galli (217,46,9 %), C. submollis (231,797,36 %).
(. 1).
2009 ,
, , .
4,651,15 %. , : C. flabellate, C. deksiflora,
C. canadensis, C. sanguinea. : C. rotundifolia (2,810,46 %), C. ellwangeriana (2,370,39 %). C. chlorosarea 10,491,73%. [3, 6].
Crataegus
C. arnoldiana
C. flabellata
C. deksiflora
C. chlorosarea
C. canadensis
C. rotundifolia
C. macracantha
C. maximowiczii
C. monogyna
C. sanguinea
C. turkestanica
C. crus-galli
C. ellwangeriana
C. pentagyna
C. persimilis
C. submollis
C. holmesiana
C. pringlei
-* -
, %
2009
2010
65,93
1,78
231,47
7,35
70,42
1,90
233,16
7,40
72,27
1,95
107,97
3,63
70,59
1,91
116,00
3,90
73,32
1,98
105,92
3,56
-*
113,39
3,81
80,96
2,19
112,41
3,78
64,93
1,75
101,87
3,42
70,29
1,90
110,29
3,71
119,95
4,03
75,58
2,04
91,26
3,07
72,86
1,97
217,40
6,90
75,35
2,04
111,91
3,76
116,00
3,90
112,06
3,77
231,79
7,36
108,78
3,66
113,64
3,82
268
_______________________________________________________________
. 447 , . 4,981,75 /100. : C. sanguinea, C. ellwangeriana, . maximowiczii, C. rotundifolia, (2,13 2,89 /100). C. monogyna 13,433,40 /100 (. 2).
2
Crataegus
C. arnoldiana
C. flabellata
C. deksiflora
C. chlorosarea
C. canadensis
C. rotundifolia
C. macracantha
C. maximowiczii
C. monogyna
C. sanguinea
C. turkestanica
C. crus-galli
C. ellwangeriana
, %
, /100
3,39 0,56
4,40
1,11
4,44 0,73
3,57
0,90
4,24 0,70
5,10
1,29
10,49 1,73
6,31
1,60
4,42 0,73
5,49
1,39
2,81
0,46
2,89
0,73
5,22 0,86
3,07
0,78
3,17
0,52
2,50
0,63
6,29 1,04
13,43
3,40
4,39 0,73
2,13
0,54
6,09 1,01
9,39
2,38
3,13
0,52
3,99
1,01
2,37 0,39
2,47
0,62
,
: . 276,767,83
163,336,93 /100, .
39,55%.
C. pringlei 30,53%,
C. sanguinea 49,60%.
(. 3).
3
Crataegus
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
C. holmesiana
C. canadensis
C. flabellata
C. pentagyna
C. crus-galli
C. deksiflora
C. persimilis
C. rotundifolia
C. turkestanica
C. monogyna
C. maximowiczii
C. sanguinea
C. submollis
C. chlorosarea
C. arnoldiana
C. macracantha
C. pringlei
C. ellwangeriana
,%
37,68
46,19
35,65
44,17
48,10
35,84
45,82
36,06
37,39
42,31
32,86
49,60
36,34
45,00
33,43
43,39
30,53
31,62
Crataegus
:
269
______________________________________________________________
-
1. .. // / - (,
28 1 2009 ) : , 2009. . 126-140.
2. .., .., - .. . / 2- . . . ., 1972. 283 .
3. .. - . , 1986. 287 .
4. .. //
/ - (, 28
1 2009 ) : , 2009. . 114-126.
5. / .
. , 1983. 340 .
6. .. . ., 1987. 248 .
7. ,
/ . ... , 1973. - . 251-279.
8. 24556-89 . C.
9. 8756.13-87 . .
10. 28561-90 . .
Article is devoted in the ascorbic acid, carotenoids, anthocyanins and sugars in the fruit of the genus Crataegus L., grown in the
Botanical Garden of Belgorod State University.
Key words: hawthorn, biologically active substances, vitamin C,
ascorbic acid, carotenoids, anthocyanins, sugar, dry matter.
270
_______________________________________________________________
543.54:547.973
-1
.. , ..
.. 1, .. 1
.. 1, .. 2
.. 2, .A. 3
.. 4
1
,
308015 . , . 85
e-mail: deineka@bsu.edu.ru
2 -
..,
127550 . , . , 49
. ..
, 197022, . -
. , 2,
-, 308015 . ,
. 85
Lonicera
caerulea L., . ,
,
: 3- 3-,
. ,
, () .
0.182 100 -
0.120. ,
-, - 0.445 0.455
100 .
: , ,
, , , ESI/MS
, ,
[1]. , ,
, [2, 3]. , [4]. , ( ) ,
.
, (
) [5 - 7], [8, 9]
.
( ) , , , .
[7, 8],
,
-. ,
.
1
-
2009-2013 ., 508 14.05.2010
.
271
______________________________________________________________
-
- .. 2011 ( )
. .. .
0.1 HCl ,
98%- .
-56
- .
. -3-,
, = 26900
/() [8].
Agilent 1200 Infinity
- . : 2504.6 Symmetry C18, 5 ; 40. : 8 .% 10 .%
, 1 /.
3- 3
, . 1.
[10 - 11] ,
( )
[12].
. 1.
1 Dp-3-Glu, 2 Dp-3-Rut, 3 Cy-3-Glu, 4 Cy-3-Rut. : 4.6250 Symmetry C18;
7 .% H3CN 10 .% HCOOH ; 1 /; 515 . :
, .
272
_______________________________________________________________
, , ,
. 3-
3-
~2 , .1. , -, - .1 M/z
(287)
(303), .2.
Dp-3-Rut
Dp +
Cy-3-Rut +
Cy +
. 2. ESI/MS 3- (Dp-3-Rut)
(Cy-3-Rut)
, ,
, . ,
Dp-3-Rut Cy-3-Glu,
Dp-3-Glu Dp-3-Rut. [13],
, . 3.
, ,
(0.7 % - 3- 5.9% - 3-).
, 3- 3-, . 1,
, .
(. ) ,
( ), . 2.
273
______________________________________________________________
Cy-3-Rut
lgk(i)
Cy-3-Glu
Dp-3-Rut
Dp-3-Glu
lgk(Cy-3-Glu)
. 3.
1
(, 20011 .)
*(.)
**, (-)/
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
0.175
0.188
0.192
0.258
0.152
0.250
0.119
0.259
0.204
0.112
0.114
0.146
0.136
0.152
0.116
0.0319
0.0243
0.0285
0.0454
0.0476
0.0347
0.0357
0.0347
0.0368
0.0275
0.0233
0.0240
0.0240
0.0191
0.0323
, % ( )
Dp-3-Glu Dp-3-Rut Cy-3-Glu Cy-3-Rut
12.7
54.3
3.4
26.7
8.3
53.5
2.6
33.1
5.8
55.3
1.9
35.0
20.6
39.5
8.1
29.7
11.5
48.6
4.0
34.0
8.1
50.9
2.8
36.2
12.3
46.6
5.2
34.5
15.9
41.7
5.3
27.0
14.3
41.6
7.3
36.0
15.1
40.3
8.4
33.5
13.4
40.6
5.6
34.9
10.8
43.0
4.8
38.3
21.5
30.3
10.2
36.0
6.2
31.8
4.8
52.1
12.0
21.9
18.5
46.0
* - Cy-3-Glu, /100 ; ** -
, 1 .
* -
, 100
0.156 0.038
0.177
0.221 0.031
0.164
0.173 0.046
0.152
0.248 0.062
0.259
0.114 0.052
0.114
0.173 0.046
0.152
0.182 0.120
0.173
0.182
274
_______________________________________________________________
, [9]. .2
, 250 100 [14].
[9]
274.9 499.1 /100 ,
(350 450 100 ). ,
, ,
, .
1. .., .., .., .., .. . // . . - 2006. - 3.
- . 16-20.
2. Wang H., Cao G., Prior R. Oxygen radical absorbing capacity of anthocyanins //
J. Agric. Food Chem. - 1997. V.45. P. 304309.
3. Einbond L.S., Reynertson K.A., Luo X.-D., Basile M.J., Kennelly E.J. Anthocyanin antioxidants from edible fruits // Food Chem. 2004. V.84. P. 2328.
4. Nakaishi H., Matsumoto H., Tominaga S., Hirayama M. Effects of Black Currant Anthocyanoside Intake on Dark Adaptation and VDT Work-induced Transient Refractive Alteration in
Healthy Humans // Altern. Med. Rev. 2000. V.5. P. 553.562.
5. Moyer R.A., Hummer K.E., Finn C.E., Frei B., Wrolstad R.E. Anthocyanins, Phenolics, and
Antioxidant Capacity in Diverse Small Fruits: Vaccinium, Rubus, and Ribes // J. Agric. Food Chem. 2002, - V.50. P. 519-525.
6. Hellstrom J., Hietaranta T., Karhu S., Mattila P., Tiikkala T., Vetelainen M. High variability in anthocyanin content between different blackcurrant varieties // NJF Report. 2010. V.6. P.
116-118.
7. .., .. , .., .. .. : // . .
. 2006. .49. .11. . 77-80.
8. .., .., .., ..
Grossulariaceae // . - 2003. - 4.
- . 324-325.
9. Rubinskiene M., Vikelis P., Stanys V., iknianas T., Sasnauskas A. Quality changes in
black currant berries during ripening // Sci. Works Lithuan. Inst. Hort. Lthuan. Univer. Agric. Sodin. Dar. - 2008. V.27. P. 235-243.
10. Slimestad R., Solheim H. Anthocyanins from Black Currants (Ribes nigrum L.) // J.
Agric. Food Chem. 2002. V.50. P. 3228-3231.
11. Nielsen I.L.F., Haren G.R., Magnussen E.L., Dragsted L.O., Rasmussen S.E. Quantification of anthocyanins in commercial black currant juices by simple high-performance liquid chromatography. Investigation of their pH stability and antioxidative potency // J. Agric. Food Chem.
2003. V.51. P. 5861-5866.
12. .., .., .., .., .., .. // .
. . 2009. .9, .4. . 529-536.
275
______________________________________________________________
13. .. // . . . - 2006. - T.80,
3. - . 511-516.
14. Sjka M., Guyot S., Koodziejczyk K., Krl B., Baron A. Composition and properties of purified phenolics preparations obtained from an extract of industrial blackcurrant (Ribes nigrum L.)
pomace // J. Horticult. Sci. Biotechnol. 2009. - Special Issue. P. 100106.
National Research
University, 308015 Belgorod Pobeda str.85
e-mail: deineka@bsu.edu.ru
2
Russian State Agricultural UniversityMTAA, 127550 Moscow, Timiriazeva str. 49.
3
. .. , 197022, . -
. , 2,
Flora-BAC Ltd. 308015 Belgorod Pobeda
str.85
276
_______________________________________________________________
615.322:543.544.5.068.7
..
..
..
..
e-mail: deineka@bsu.edu.ru
Calendula officinalis, .
-
-
, -, - -, , , .
, - . , ,
.
: Calendula officinalis, , ,
, .
, Calendula, , 10 20 [1]
, ,
.
(C. officinalis L.) [1, 2]
. , -, pot
marigold, : corn marigold (Glebionis
segetum), desert marigold (Baileya), marsh marigold (Caltha palustris) , ,
French marigold (Tagetes patula) African marigold (Tagetes erecta). ,
[3].
( ).
, ,
[4] . 672 ,
Calendula officinalis,
marigold,
, . , , .
[2, 5] [6],
:
1) (Flores Calendula officinalis) , ,
, . (10 200
) .
. ,
, .. .
277
______________________________________________________________
2) (Tinctura Calendulae) ( 70%- , 1:10)
, , ,
3) (Unguentum Calendula)
; , .
(Oleum Calendula), . , , , , , .
, ( )
, .
[7] , .
,
( 1),
: -, -, -, , , , ; , , , , , , , . ,
.
OH
HO
OH
HO
OH
O
HO
OH HO
HO
OH
OH
O
O
HO
OH
OH
HO
HO
HO
1.
OH
278
_______________________________________________________________
,
[10]
.
[11]
YMC Carotenoid (2504,6 , 5)
0/100, 12/100, 96/0 (/%), : --
(90:6:4 ), : -- (25:71:4
), ,
10%- NaCl.
.
(
E161a) (8R)-,
, .
Pintea . [79]
Double Estrel Orange, 0,276 / ,
- (16,7%), (14,4%),
(14,1%), (14,0%), - (12,2%), (9,2%), (8,9%),
(4,5%) . Bonbon Abricot 0,048 / (42,1%);
(19,0%), (12,3%)
(11,3%) . , 18-
Nucleosil ODS, 5 (2504,6 )
( 9:1 0,25% ) ( 0,25% ).
, ; , , , .
,
,
,
- , , .
() .
,
( 3 5 ). .
. 0,4 .
,
- 10%- .
279
______________________________________________________________
(, ).
Agilent 1200 Infinity . : 2504 Reprosil-Pur C18-AQ, 5 .
, .1,
, , , 422 , .
, , max . .
, , ,
, . 2. - .
( -) ( 0,1:10) -, . 2,
. , ( 2:10) -,
, . 2.
. 1.
. -
, . 3.
,
(50 .% ) . 4 (60 .% ) . 5.
,
.
, , . :
, .5, max 399, 422 448 , , , [11] (
).
280
_______________________________________________________________
)
. 2. (SiO2)
1 2 ; 3
. : /- - ) 0,2:10; ) 2:8.
. 3. ()
()
-
1 -, 2 , 3 -; 2504
Reprosil-Pur 5C18-AQ, 33 .% ; 1 /.
2 ,
(446 474 ). 3 16 17
,
-, . , 3 (. 5) , 2,
,- - .
281
______________________________________________________________
. 4.
2504 Reprosil-Pur 5C18-AQ, 50 .% ; 1 /.
HO
2.
. 5.
2504 Reprosil-Pur 5C18-AQ, 60 .% ; 1 /.
.4, ,
.
, . .
. 1 (. 4) 463
472 , .
2 (. 4)
458 462 -,
max max - 453,5 (. 3 . 4), . 6.
282
_______________________________________________________________
- ( 3, . 4),
.
3 3 2 1
. 6. . 4
- , , , :
- (
) :
() = lgk(-cryptoxanthin)-lgk(rubixanthin) = 0,271-0,153 = 0,118.
- :
2 () = lgk(-carotene)-lgk(lycopene) = 0,575-0,340 = 0,235,
0,235/2
= 0,1175.
-:
lgk(-carotene) = lgk(lycopene)+(lgk(-carotene) lgk(lycopene))/2 = 0,458,
.
6,75 7,25 , .7: 1 .5,
3 , , [12]. () , . 5,6-
-, . , 8,0 8,5
.
, , , ,
- (0,111). ,
- , .
. : 0,911 / 1,015 /
( ), -
283
______________________________________________________________
. ,
450 , 9,4%, 4,5%, 8,7%,
21,2%, 25,2%, - 13,6%,
- 1,6% - 9,2%. -
( 17,3%), ( 4,4 3,5%, ).
1
. 7. (1)
(2)
, , .
2009 2013 , -174.
1. .., .. //
. 2007. 1. . 65-68.
2. . . , . . , . . , . . , . .
(Calendula officinalis L.) // . 2001. .5.
http://www.provisor.com.ua/archive/2001/N5/art_29.php
3. .., .., .., .. Tagetes sp. // .-. . 2007. - .41, 10. . 30-32.
4. High-Performance Liquid Chromatography in Phytochemical Analisis. Chromatographic
science series. V.102. Ed. M. Waksmundzka, J. Sherma. CRC Press, Taylor & Francis Group. 2011.
995 p.
5. Muley B.P., Khadabadi S.S., Banarase N.B. Phytochemical Constituents and Pharmacological Activities of Calendula officinalis Linn (Asteraceae): A Review // Tropical J. Pharm. Res.
2009. V.8. P. 455-465.
6. .. . 15- ., ., . .
.: , 2005. 1200 .
7. .., .. . : , 2000. 102 .
8. Goodwin T.W. Studies in carotenogenesis. 13. The carotenoids of the flower petals of Calendula officinalis // Biochem. J. 1954. V.58. P. 9094.
9. Bako E., Deli J., Toth G. HPLC study on the carotenoid composition of Calendula products // J. Biochem. Biophys. Methods. 2002. V.53. P. 241-250.
284
_______________________________________________________________
10. .. //
. - 2010. - 6. - . 93-93.
11. Pintea A., Bele C., Andrei S., Socaciu C. HPLC analysis of carotenoids in four varieties of
Calendula officinalis L. flowers // Acta Biol. Szegediensis. 2003. V.47. P. 37-40.
12. Pilar Cano M., Begona de Ancos Carotenoid and carotenoid ester composition in mango
fruit as influenced by processing method / J. Agric. Food Chem. 1994. V.72. P.2737-2742.
V.I. Deineka
I.. Gostyshchev
.Yu. tetjakov
I.V. Indina
Belgorod National Research
University
e-mail: deineka@bsu.edu.ru
The paper is devoted to the investigation of Calendula officinalis flower petals carotenoids harvested in Belgorod. By means of RP
HPLC with diode-array detector and TLC of unsaponified aceton extracts by retention parameters as well as spectral analysis the main
components were identified as lycopene isomers, isomers of -, - and
-carotene, unesterified flavoxanthin, lutein rubixanthin, monoesters
of flavoxanthin and luteoxanthin. The difference of the complexes of
yellow and orange colours has been found to be mainly due to the lower fraction of lycopene, -carotene and rubixanthin in the former case.
The attention has been paid to the problem of Tagetes Calendula
confusion in commercial and scientific information.
Key words: Calendula officinalis, carotenoids, HPLC, TLC,
spectrophotometry.
285
______________________________________________________________
..
..
..
..
..-.
..
. .
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
-
-
,
, .
,
, ,
, ,
, ,
,
,
, ,
,
,
,
,
,
,
, ,
, ,
, , - ,
,
,
286
_______________________________________________________________
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
. .
..
..
. .
..
..
..
..
..
..
..
..
..
,
, ,
. .. , , ,
.
,
, ,
, ,
, .
,
, , .
,
,
,
,
,
, .
,
,
, . .. , , , .
,
.
. .. , -
, ,
, ,
,
,
287
______________________________________________________________
..
..
..
..
..
..
..
..
..
..
. .. , -
,
.
,
, , . ..
, ,
,
, ,
,
288
_______________________________________________________________
:
, ,
() ,
. .
(1 .)
Microsoft Word Windows. ( -4): 3 , 2 . Times New Roman, 12 pt, , () 1,25 ., . . 12 ,
, .
( ).
, (12 pt) .
(12 pt) . , e-mail ()
(10 pt) .
()
(10 pt) . (57).
, ,
,
, .
,
.
. , :
( ), ( ),
, (), .
, , (. ).
, , : , , , , , , , ,
, , e-mail.
, , , ( ).
.
.
. ( ) 4- . ( ). , .
http://www.bsu.edu.ru:8806/dok_fakt.htm
289
______________________________________________________________
51-72:530.145
.. 1, .. 2, .. 3
-.
. ,
,
, , ,
. , .
: , -, , , , .
[1, 2]. ,
[3],
, [4].
.
1. ., . . .: , 1984. 528 .
2. .. //
. 1987. .32. . 1316-1321.
3. ..
// . 1978. .75, .4. . 1261-1268.
4. Gutzwiller M.C. Chaos in Classical and Quantum Mechanics. New York: Springer, 1990. 432 p.
5. Henon M. Integrals of the Toda lattice // Phys. Rev. 1974. Vol. 9, 4. P. 1921-1923.
Belgorod State University, Studencheskaja Str., 14, Belgorod, 308007, Russia, e-mail: Chekanov@bsu.edu.ru
2
Civil defence Academy of Ukraine, Chernyshevsky Str. 94, Kharkov, 61023, Ukraine
3
National Scientific Center, Kharkov Institute for Physics and Technology,
Akademicheskaj Str. 1, Kharkov, 61108, Ukraine
The receiving procedure of classical Hamiltonian to the Birkhoff-Gustavson normal form is described. With help of the Weyl correspondence rule for the classical normal forms of some integrable and nonintegrable systems their quantum counterparts are constructed and approximated energy spectra and wave functions are found. It is shown that thus obtained energy spectra are represented good exact ones in
an energy domain where the classical motion is regular but agreement is worsen strongly at the energy domain where the classical regular
motion is going into chaotic one. It is established that .
Key words: classical Hamiltonian, Birkhoff-Gustavson normal form, Weyl correspondence rule, energy spectra, wave function, method of
quantization.
290
_______________________________________________________________
291
______________________________________________________________
.
. 1
81466
( )
. -.
-
.
( )
2011
1
2
3
4
5
6
7
8
9
10 11 12
( ) ()
81466
( )
( )
. .
( )
2011
4
5
6
7
8
-
.
.
.
(, . .)
10
11
12