Академический Документы
Профессиональный Документы
Культура Документы
9 (104) 2011
15/1
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77-21121 19 2005 .
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Humulus lupulus L. .., .. 5
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. .., .., ..,
.. 10
Rorippa Scop.
. .., .. 16
Epipactis palustris[L.] Crantz . ..,
.. 23
Eremotheciaceae.
.., .. 29
Helictotrichon krylovii (Pavl.) Henrard
(- ). .., .. 34
in situ. .., .. 38
(Viburnum opulus L.) .
.. 44
.., .. 50
(Bromopsis inermis (Leys.) Holub - . .. 56
Plectranthus amboinicus
(Lour.) Spreng. . ..,
.. 60
Dipteronia sinensis Oliv. (Ageraceae). .O. 64
Corylus L. subsect. Corylus (Betulaceae). .. 67
. .., .., .. 71
Forsythia
Oleaceae. .. 76
Vitaceae
. .. 80
Sorbus L. . .., .. 86
Comoranthus Schrebera (Oleaceae).
.., .. 88
-
. .., .. 92
Taraxacum ceratophorum (Ledeb.) Dc. . .., .. 99
. ( - ). .., .., .. 105
Pyrola incarnat (Dc.) Fleyn - ( ).
.., .. 112
. ..,
.., .., .. 117
. .., .. 124
. .., .., .. 131
Amarillidaceae Jaume St.-Hill.
. .., .., .. 136
Campanula L. . .., .. 140
Sorbus L. Aronia L. . .. 145
Chrysanthemum L. .
.., .., .., .. 150
(Oxytropis DC.) . .., .. 157
_______________________________________________________________
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E-mail: prisniy@bsu.edu.ru
21.07.2011
6084/8
Georgia, Impact
. . . 47,07
1000 .
156
:
81466,
39723
-
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.
.., .., .. 165
Iridaceae Juss. Asparagaceae Juss.
.. 171
.
.., .. 175
- . .. 180
. .., .. 186
Rhododendron L. . ..,
.. 192
Ambrosia artemisiifolia L.,
Iva xanthiifolia L., Xanthium albinum (Widd.) H. Scholz - . .., .. 199
(Fragaria x ananassa) . .., .. 204
( ). .., ..,
.., .., .. 210
. .., .. 217
Campanula latifolia L. ex situ. .., .. 223
Momordica charantia L.
(Cucurbitaceae), . . ,
.. 228
.
.., .. 232
Momordica charantia L. Momordica balsamina L. (Cucurbitaceae) . . , .. 237
,
. .., ..,
.. 242
Conyza canadensis (L.) Cronq . .., .. 247
Conyza canadensis (L.) Cronq. .., .. 252
. .. 257
Helianthtmum Mill .
.., .., .. 261
. .. 268
Hydrangea Dumort. . .. 272
Magnolia sieboldii K. Koch. . .., .. 277
. .., .. 281
. .. 290
Lonicera L. .
.., .., .. 296
Juglans -
. .., .., .. 302
. .., .. 308
Sambucus L. . .., .. 313
- - .
.., .., .., .. 317
.
.., .. 321
Iris pumila L. . .., .., .. 326
- upatorium cannabinum L., Lophanthus
anisatus Benth., Monarda citriodora Cerv., . .., .. 332
Rsa L. (Rosaceae Juss.).
.., .., . . 337
. .. 348
- . .., .. 357
. .., .., .., .., C.. 364
. .., .., .., ..,
.. 371
haenomeles japonica C. maulei. .., .., .., .., .. 379
386
390
, 2011
9 (104) 2011
Issue 15/1
SCIENTIFIC PEER-REVIEWED JOURNAL
Founded in 1995
The Journal is included into the list of the
leading peer-reviewed journals and publications coming out in the Russian Federation
that are recommended for publishing key results of the theses for Doktor and Kandidat
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Founder:
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Publisher:
Belgorod State University
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The journal is registered in Federal service
of control over law compliance in the sphere
of mass media and protection of cultural
heritage
Certificate of registration of mass media
77-21121 May 19, 2005.
CONTENTS
BIOLOGY
Biomorphological aspects of introduction and cultivation Humulus lupulus L. Degtereva
O.P., Savinykh N.P. 5
Specifically conservational natural territories of Kirov region: present status and development prospects. Savinykh N.P., Perestoronina .N., Kiselyova .., Shabalkina S.V. 10
The construction of bine systems of some species Rorippa Scop. from the perspective of
modular organisation. Shabalkina S.V., Savinykh N.P. 16
Biomorphology of Epipactis palustris[L.] Crantz from the point of view of the protection of
the species. Chuprakova E.I., Savinykh N.P. 23
Morphological research of some representatives of family Eremotheciaceae. Semenova E.F.,
Shpichka A.I. 29
Ontogenetic structure of Helictotrichon krylovii (Pavl.) Henrard coenopopulations under condition
of the upper Yana region (North-east Yakutia). Skryabina R.N., Andreyeva S.N. 34
Crop wild relatives of european Russia from the problem of their in situ concervation. Smekalova T.N., Chukhina I.G. 38
Introduction a Guelder-rose ordinary (Viburnum opulus L.) in the condition of the Tomsk
region. Suchkova S.A. 44
Age changes of common flax plants in onthogeny. Fadeeva T.M., Semenova E.F. 50
Editor-in-chief
L.J. Djatchenko
Rector of Belgorod State University, doctor
of sociological sciences, professor
Deputy editor-in-chief
A.P. Peresypkin
Vice-rector for scientific research of Belgorod
National Research University, candidate of
pedagogical sciences
Assistant Editor
Moskovkin V.M.
Doctor of geographical sciences, professor
of world economy department Belgorod National Research University
Krolevetskaya E.N.
Candidate of pedagogical sciences,
associate professor of Pedagogics
department of Belgorod National Research
University
A biomorphological features Plectranthus amboinicus (Lour.) Spreng. depending on different level of electric light and fertilizers. Tsitsilin A.N., Motina E.A. 60
Fruit anatomy and morphology of Dipteronia sinensis Oliv. (Ageraceae). Yatsenko I.O. 64
About fruit structure of Corylus L. subsect. Corylus (Betulaceae). Yatsenko O. V. 67
Creation of gene bank of rare plant seed in the Udmurt university Botanical Garden. Baranova O.G., Dedyukhina O.N., Yagovkina O.V. 71
Study of Forsythia fruits development and structure in connection with systematics and phylogeny. Filonenko A.V. 76
The peculiarities of ontogenesis in decorative vines of Vitaceae family in the conditions of the
monsoonal climate of southern Primorye. Vrzhosek E.V. 80
Phenological atypicity of introduced species of genus Sorbus L. in Ufa botanical garden. Abdullina R.G., Vafin R.V. 86
Comparative carpology of genera Comoranthus Schrebera (Oleaceae). Filonenko A.V.,
Efremov A.N. 88
Life strategyes of Chamaenerion angustifolium (L.) Scop. in Western Yakutia. Gavrilova
Zh.A., Cherosov M.M. 92
Evaluation of ecological condition and life strategy of Taraxacum ceratophorum (Ledeb.) Dc
in the Middle Lena valley. Everstova T.A., Cherosov M.M. 99
Hordeum jubatum L. in the viginities of Mirny town (experience of population-biological
analysis of coenopopulaiton). Gavrilova Zh.A., Cherosov M.M. 105
Influence of zoogene factorson ontogenetic parameters condition coenopopulaitons
Pyrola incarnat (Dc.) Fleyn in Lena-Amginski interfluves (the Central Yakutia). Nikiforova A.A., Cherosov M.M. 112
Use of GIS-technologis for correction of contours of small-scale map of vegetation. Ammosova E.V.,Cherosov M.M., Nikolin E.G., Troeva E.I. 117
Comparison of regional floras of the Verkhoyansk Range use the cluster analysis. Nikolin
E.G., Cherosov M.M. 124
L.J. Djatchenko
Rector of Belgorod National Research University, doctor of sociological sciences, professor
Terms of flowering of species and culyivars of some genera Amarillidaceae Jaume St.-Hill. in the
greenhouse of the Winter garden. Arhipova I.N., Sorokopudova O.A., Suchkova O.V. 136
Chief editor:
A.V. Prisnyi
Doctor of biological sciences, professor
(Belgorod National Research University)
Deputies of chief editor:
O.E. Lebedeva
Doctor of chemical sciences, professor
(Belgorod National Research University)
Hemeroby status of plants of Yakutia. Pestryakov B.N., Cherosov M.M., Ishbirdin A.R. 131
The ontogenesis of some Campanula L. genus species under introduction in Bashkir Predurale conditions. Allajarova I.N., Mironova L.N. 140
Seasonal rhythm of the species of Sorbus L. and Aronia L. with the introduction on Saratov.
Arestova E.A. 145
Phenological phases of species and varieties of Chrysanthemum L. in the conditions of the
south Central Russian upland. Stetsovich A. S., Sorokorudova O.A., Sergeeva E.V.,
Alehin A.A. 150
Comparative analysis of of South-Ural species of the Oxytropis DC. by morphometric parameters of chromosomes. Arslanova L.R., Kalashnik N.A. 157
Selection and estimation of parent material for strawberry selection in conditions of Yakutia.
Belevtcova V.I., Mironova A.A., Sorokopudov V.N. 165
Sleedprotein complex evolution in family Iridaceae Juss. and Asparagaceae Juss. Sokolova S.M. 171
_______________________________________________________________
A.G. Kornilov
Doctor of geographical sciences, professor
(Belgorod National Research University)
Drought resistance and heatproof new varieties and hybrids of strawberryes. Govorova G.F.,
Bulanov A.E. 175
Responsible secretary:
Yu.N. Kurkina
Candidate of agricultural sciences, associate
professor (Belgorod National Research University)
Short results of the introduction of daylilies in Bashkiria. Zaynetdinova G.S., Mironova L.N. 186
The peculiarities of artificial vegetative propagation of arcto-montane willows. Epanchintseva O.V. 180
Anatomic features of bud scales of generative buds of alien Rhododendron species in the
Southern Primorsky region. Koksheeva I.M., Tsarenko N.A. 192
Invasive species distribution peculiarities of Ambrosia artemisiifolia L., Iva xanthiifolia L., Xanthium albinum (Widd.) H. Scholz in the southwest of the Middle Russian Height. Tokhtar V.K.,
Volobuyeva J.E. 199
Variability of pollen fertility in agomospermic offsprings Fragaria x ananassa Duch. Baturin S.O., Ambros E.V. 204
Peculiarities of edafotop condition formation within the site with technogennously changed
relief (on the example of Botanical Garden BelSU). Goleusov P. V., Tokhtar V.K., Afanasev E.G., Jurieva M.N., Tolkacheva A.A. 210
Effect of planting schemes on rooting and morphological features green guttings of honeysuckle. Kondratyev A.V., Belosohov F.G. 217
Ontogenesis of Campanula latifolia L. ex situ. Komir Z.V., Alyokhin A.A. 223
The comparative analysis of antibacterial activity of Momordica charantia L. (Cucurbitaceae)
samples, grown up in Vietnamland under the conditions of the Belgorod region. Doan H.
Zhang, Tokhtar V.K. 228
Sowing qualities of seeds of herbs with antifungal properties. Kurkina Yu. N., Pshenichnaya O.G. 232
Momordica charantia L. and Momordica balsamina L. (Cucurbitaceae) cultivation under the
Belgorod region conditions. Doan H. Zhang, Tokhtar V.K. 237
Comparative analysis of alien florae within borders of Graivoronskiy and Rovenskiy districts
of the Belgorod region. Kurskoy A.J., Tokhtar V.K., Kolchanov A.F. 242
Morphological features study in Conyza canadensis (L.) Cronq. populations under the
southwest of Central Russian upland conditions. Tokhtar V.K., Mazur N.V. 247
Variabylity of correlation structures of morphological features in populations of Conyza canadensis (L.) Cronq. Tokhtar V.K., Mazur N.V. 252
Results of the introduction of honeysuckles in arboretum Gornotaezhnaja station FEB RAS.
Malysheva S.K. 257
Isolation populations of rare Helianthemum Mill species in the Urals. Minogina E. N.,
Zimnitskaya S. A., Semkina S. L. 261
Weedy plants in seeds parties of olive cultures. Mikhailova S.I. 268
The seasonal rhytm of growth and development of Hydrangea Dumort. representatives in
Bashkir Cis-Urals. Murzabulatova F.K. 272
Biological features Magnolia sieboldii K. Koch. at introduction in the south of Primorski
krai. Petuhova I. P., Kameneva L. A. 277
Recreational potential of forests in urban areas. Rysin S.L., Lepeshkin E.A. 281
Phytoindication for monitoring of forests in urban areas. Rysin S.L. 290
Collection of the Lonicera L. genus in the Botanical Garden of the Institute of Biology Komi Science
Centre Ural Division RAS. Skupchenko L.A., Ryabinina M.L., Skrockaja O.V. 296
Criteria of the estimation of level of drought resistance of kinds of sorts Juglans in the conditions of
the Belgorod region. Rezanova T.A., Sorokopudov V.N., Nazarov N.V. 302
Some approaches to the directed selection of species at stable agrophytocoenoses creation in
anthropogenously transformed ecotopes. Martynova N.A., Tokhtar V.K. 308
Drought resistance of some representatives of sort Sambucus L., in the conditions of the Belgorod region. Koltsov S.V., Sorokopudov V.N. 313
Peculiarities of natural-reserved fund florae formation in the southwest of the Central Russian upland in the conditions of moderate anthropogenous impact. Tokhtar V.K., Tretyakov M. Ju., Skorbach V.V., Koniaeva I.A. 317
Parameters of generative organs of Juniperus communis L. in the South Urals. Farukshina
G.G., Putenikhin V.P. 321
Populations monitoring of Iris pumila L. in Rostov-on-Don area. Fedyaeva V.V., Shmaraeva A.N., Shishlova J.N. 326
Anatomist-morphological particularity sheet Eupatorium cannabinum L., Lophanthus anisatus Benth., Monarda citriodora Cerv. as one of the factors to adaptation type toecological
condition of the south Tomsk area. Kharina T.G., Kirsanova N.V. 332
Features of the anatomic structure of sheet of sort Rsa L. (Rosaceae Juss.). Rezanova .., Sorokopudov V.N., Svinarev . N. 337
CHEMISTRY
Passed for printing 21.07.2011
Format 6084/8
Typeface Georgia, Impact
Printers sheets 47,07
Circulation 1000 copies
Order 156
Subscription reference in Rospechat
agency catalogue 81466,
In joint catalogue Pressa Rossii 39723
Dummy layout is replicated
at Belgorod National Research University Publishing House
Address: 85, Pobedy str., Belgorod, Russia, 308015
______________________________________________________________
581.4
HUMULUS LUPULUS L.
..
..
610007, ., . , 198
e-mail: botany@vshu.kirov.ru
Humulus lupulus L..
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(Humulus lupulus L.). (Cannabaceae) [1]; [2].
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Humulus lupulus
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Humulus lupulus 2008-2011 . , , .
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1. .., .. Mangoliaceae Limoniaceae //
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3. .., .., (Humulus lupulus L.) //
______________________________________________________________
: ( 3 ). 3.
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4. .., .. (Humulus lupulus L.) // - .
VIII. .2. : , 2010 . - .77-81.
5. .., .., .., .. :
. . . 2-, . . ., 2005. 256 ., .
6. .., . .: , 1984. - 287 .
7. .., . - .: , 1993. - 286 .
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1977. 158 .
10. .., .., . .6:
. .: , 1975. 1232 .
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17. .. // . . . . . - . .. . 1954. .37, .2. .3-20.
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// . 17.10.2007. 192 193 (4080-4081). . 9.
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16. . ., . . // . .7. . , 1997. . 223-232.
17. : , , / . .
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18. . . ., 2003. 39 .
In the article historical aspects of becoming of network of specifically conservational natural territories (SCNT) in Kirov region, present
status and development prospects of system of SCNT are considered,
perspective territory (PT) in accordance with the modern conceptualization of SCNT is defined categories of SCNT and examples of PT on result of work are given.
Keywords: biodiversity, specifically conservational natural territories (SCNT), categories of SCNT, reserve, reservation, reserve territories,
perspective territory.
16
________________________________________________________________
581.4
RORIPPA SCOP.
..
..
, ,
610007, . ,
. , 198
e-mail: botany@vshu.kirov.ru
Rorippa amphibia (L.) Bess., R. palustris (L.) Bess., R. anceps (Wahlenb.) Reichenb.,
: , , .
, .
: , , , , Rorippa, .
.
. ,
[1, 2, 3, 4].
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Rorippa.
, (.): Rorippa amphibia (L.) Bess. ,
R. palustris (L.) Bess. , R. anceps (Wahlenb.) Reichenb .
2007-2010 .. . (SYKO), . . . (LE), (MOSP) . . .
(IBIW), (YOLA, . -).
.
R. amphibia -- [9, 10],
[11].
[12].
R. amphibia [13]
, ,
10-40 . , [14], ,
, [15].
17
______________________________________________________________
. ( [16]):
Rorippa amphibia; R. palustris; R. anceps.
: Tm ; Kn ;
Om -; Cr ; Hd ; Tr ; Rc ; Nt
; fH ; Lc -.
R. palustris - [9],
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- . , - . .
( ,
), .
. - ;
() ,
- . , ,
-
.
, () R. amphibia
, () .
R. palustris 1) , ; 2)
, ; 3) , . , .
9 (.). 8-14 . . ,
.
, , ,
, .
19
______________________________________________________________
2
(),
Rorippa x anceps
Rorippa palustris
Rorippa amphibia
, -
, 1
, -
(),
, -
10
11
1 ; 2 ; 3 ;
4 ; 5 ; 6 - ; 7
; 8 ; 9 ; 10 - ; 11 .
, ,
. . . [24] , , ; , , ,
.
, [25], .
20
________________________________________________________________
1
10
11
,
2
12
, ,
13
, , -
14
, ,
15
, , - ( 1)
( )
16
, ,
-
17
, -
18
, 2
19
,
( ) ( )
20
, -
R. anceps 1) -
1
21
______________________________________________________________
, ;
2) - , , .
, ()
, .
9 .
16-30 , ,
.
.
, 4 (.). R. amphibia ,
, , .
( ),
( R. amphibia R. anceps); ( R. amphibia R. palustris),
( R. amphibia), ( R. anceps).
, , :
, , .
Rorippa :
( ) ( ); , ( );
( ).
1. . ., . .
// . . . 1999. . 60, 1. . 49-59.
2. . . // . . .
1999. . 60, 1. . 6-17.
3. . . : .
. - . . ., 2000. 32 .
4. . . // . . V. -, 2007. . 15-34.
5. . . - - : . .. . . , 2006. 21 .
6. . . Caltha palustris L. // . 2008. . 9, 25 (85). . 134-138.
7. . . :
. . . . . , 2009. 21 .
8. . . - : . .
. . . , 2008. 20 .
9. . . - (, .). .: - , 2000. 781 .
10. . . 587. Roripa Scop. / . .-.: -
, 1939. . 8. . 135-141.
11. . . (Cruciferae Juss.) / Turczaninowia,
2002. . 5, . 3. . 97-100.
12. . . . . : .
, 2007. . 1. 440 .
22
________________________________________________________________
13. . . // . 1985. 6.
. 8-13.
14. . . . ., 1964. 880 .
15. . ., . ., . . . . ., 2009. 219 .
16. . . . .: , 1983. 197 .
17. . . Rorippa (Brassicaceae) // . . 1998. .
83, 8. . 98-106.
18. / . . . . . .: , 1979. 355 .
19. . . :
. : ,2001. 200 .
20. . . // . . (-, 14-18/ 2002 .). , 2002. . 95-96.
21. . . // . . . . 1974. . 79, . 1. . 100-113.
22. . . Veronica officinalis L. //
. . . . 1978. . 83, . 43. . 123-133.
23. Raunkiaer C. Types biologiques pour la geographie botanique // Forhandbinger Kongelige
Danske Videnskabernes Selskabs. 1905. T.5. P. 347-437.
24. . . . .: .
, 1952. 391 .
25. Troll W. Die Infloreszenzen. Jena: Fischer Verlag, 1964. I Band. 615 s.
23
______________________________________________________________
581.4 : 582.594 : 58:502.75
..
. .
,
610007, . , . , 198
e-mail: botany@vshu.kirov.ru
,
- (), Epipactis palustris.
8 , .
: , , - , , , - , Epipactis palustris.
. .
, ,
, , - .
Orchidaceae . . ,
[1, 24].
[1, 2, 4, 7, 12, 25]. -
; Cypripedium calceolus L.
C. guttatum Sw. [26], Gymnadenia conopsea
(L.) R. Br. [5] - [14] .
, , .
Epipactis alustris (L.) Crantz
(. Orchidaceae) III- ,
[9], [10] , [11], .
E. alustris
.
E. alustris, ( )
[17]. ( ) --
Bistorta major S. F. Gry, . ( 100 1 2).
24
________________________________________________________________
. , , .
20092010 . ,
: IBIW, LE, MOSP
.
-
[18, 19 .]. 10 . ,
.
: , , , , ,
, .
, ,
,
. .
[8]. . . [21] [16].
- Troll [28]
[3]. [15].
E. alustris ; ; () . [27].
, .
E. alustris
- (), . . [25].
,
. E. alustris . . [20].
() () .
() , ,
. : 0,7 ,
(27 ) ( 1 ), . ,
;
( 7 ). . : 2 (13 )
36
(715 ), ; ( 2 1
). ,
(0,2 ) , , ( 12 )
2 . . . [25] . -
25
______________________________________________________________
, ,
.
-
, . ( 34 )
( 12 ) ,
.
.
. . [20] .
. 2 , : 34 .
, , ;
1 .
( 2 ) . . [25]. , 815 (30) , 1,52,5 (4) .
12 3
. , ( 69
0,51,2 ), (13).
, 615 (22) , 912 (6)
. , ,
. , [6].
, , - :
, 2
(. 1; 1);
(. 1; 2);
2
2 89
(. 1; 3);
(. 1; 4).
[15] E. alustris
: , .
, , ( ).
8: , ; , ; , 2 , , ; , 2 ; , ; ,
; , () .
, 2
n+1 .
26
________________________________________________________________
. 1. - E. alustris
- (1 4) : ;
; 1 ; 2
; 1 ; 2 ;
3 ; ; ; .
, ,
. E. alustris
.
, , . . . [13]
.
9 . - .
, -.
- , ; .
( ) ,
. , , , .
,
. . [23], .
E. alustris . 12 -
27
______________________________________________________________
:
.
, .
, - , ,
.
. ,
.
, . ,
- , 9 . ( ) . , , ,
. . [22], 12 .
. , , , -, , . ,
. , . .
, . , (, ),
.
. . -
. . .
1. .. , : . . . . . ., 1998. 16 .
2. .. -
: . . . . . .,
2010. 44 .
3. .., .. - // . 1990. .75, 10. .1420-1425.
4. ..
: . . ... . . . ., 1992. 20 .
5. .. Gymnadenia conopsea (L.) R. Br. (Orchidaceae)
: . . . . . , 2009. 18 .
6. .., .., .., .., ..
// . . 1997. .13. .51-62.
7. .. (Orchidaceae
Juss.) : . . ... . . . ., 1999. 24 .
8. .., .., .., .. :
: . . . 2-, . . ., 2005. 256 ., .
9. : , , / . .
.. , .. . : - . -, 2001. 288 .: .
28
________________________________________________________________
10. ( ): . , 1996. 279 .
11. . , 1980. 281 .
12. ..
Orchidaceae () : . .
. . . , 2008. 16 .
13. .. . . 1. . .: , 1997. 630 .
14. .. -
(Orchidaceae) - : . . ... .
. . , 2006. 18 .
15. .. // II . ., 2002. .95-96.
16. .. // : ( 3 ). . 2. . :
, 2009. .16-21.
17. .., .., .., .., ..,
.., .. - . , 2009. 303 .
18. .. . .1. // . .
- . . . 1948. .53, .2. .49-66.
19. .. . .2. // . .
- . . . 1949. .54, .2. .47-62.
20. .. . ., 1952. 391 .
21. .. // . .: , 1964. .3. .146-208.
22. .. . .:
, 1971. 360 .
23. .. -
// : , . ., 1981. .161-179.
24. .. : , , . .;
1996. 207 .
25. .. (Orchidaceae Juss.) : .
. . . ., 2007. 429 .
26. .., .., .., ..
. , 2003. 180 .
27. Raunkier C. The life forms of plants and statistical plant geography. Oxford, 1934. 632 p.
28. Troll W. Die Infloreszenzen. Jena: Fischer Verlag, 1964. B.1. 615 p.
Features of the structure, structural-functional zones of the shootroot complex (SRC), forthputting model and seasonal development of
Epipactis palustris are described. A universal module and 8 types of the
elementary modules composing the body of a plant are allocated.
Key words: biomorphology, structural organisation, structurally
functional zone, forthputting model, seasonal development, shoot-root
complex, Epipactis palustris.
29
______________________________________________________________
581.4:582.28
EREMOTHECIACEAE
..
..
, 440026, . ,
. , 40
e-mail: sef1957@mail.ru
- 2
Eremotheciaceae.
.
: Eremothecium ashbyi Guilliermond, Eremothecium gossypii Kurtzman, - ,
in vitro.
, Eremotheciaceae , , .
Eremothecium - , . E. ashbyi E. gossypii 2 [1]. , , Eremothecium
, (69,584,9 %) (12,727,7 %), - , , ,
. 180 / ,
500600 [2-5].
- Eremotheciaceae.
Eremothecium ashbyi Guilliermond
1935 BKM F-124, F-3009 (, BKM
F-124) Eremothecium gossypii (S.F.Ashby et W.Nowell 1926) Kurtzman 1995 (
Ashbya gossypii (S.F.Ashby et W.Nowell 1926) Guilliermond 1928) BKM F-1398, BKM F3276 (, BKM F-1398), BKM F-3296.
Fungi, Ascomycota, Endomycetes, Saccharomycetales, Eremotheciaceae [6,7].
, (4%) (1%) [5], -, , , - , , [1]. -3 (
10, 40, 100) , . Panasonic DMC-FX100 c Lumix 12 mega pixels.
. E. ashbyi ( ) , . , 812 ( - 3 .
271). -, -
30
________________________________________________________________
- , , .
, E. shbyi F-124
, , (, .).
E. gossypii , -, -
, 3
271 6 , , . , , , ,
. 10 -, , . () . , -, - , , .
. E. ashbyi , , . , ,
.
2,516,5 . . . , , ,
, 65-9014-20 , , . , , ,
. : 20,226,7 , 2,52,8 .
E. gossypii , , , , . . . ( , ) , , , 100-20010-20 , .
. 2 2 6, ; 4 32 . - 25-372-5 ; , .
. ( ) . 3648 . , (4860 ): ,
() - . ,
.
Eremothecium ashbyi, Eremothecium gossypii (Ashbya gossypii, Nematospora gossypii),
Kurtzman ,
, Eremotheciaceae [6,7].
31
______________________________________________________________
. 1. : ;
; ;
( 4100, )
. 2. (: 440, 4100; )
. 3. (),
;
( 4100, )
32
________________________________________________________________
. 4.
( 4100, )
. 5. ;
;
( 4100, )
,
: , , , ;
; ;
4-32 , ,
, , [2-4, 6,7].
1. Eremothecium ashbyi
( 1,5-2 ) E. gossypii ,
3 .
2. Eremothecium
ashbyi Eremothecium gossypii ; : , , ,
;
; ,
.
33
______________________________________________________________
3. - ,
Eremothecium ashbyi, Eremothecium
gossypii (syn. Ashbya gossypii, Nematospora gossypii),
Kurtzman
Eremotheciaceae.
242 .
1. .. . : , 1973.
34
________________________________________________________________
581.16 : 633.28.(282.256.74)
HELICTOTRICHON KRYLOVII (PAVL.) HENRARD
(- )
.. 1
.. 2
1)
, 678500,
. , . 3,
()
e-mail: raika60@rambler.ru
2)
,
677980, . , . 41
e-mail: Sandren_1601@mail.ru
Helictotrichon krylovii (Pavl.) Henrard. . . . krylovii
, -- - .
H. krylovii 3 9 . 3 : , .
H. krylovii , , .
: , ,
, .
35
______________________________________________________________
. krylovii
(40 60 %). . krylovii 1 3 %. H. krylovii (50 55 ); - (12,4 12,6 ) [3].
H. krylovii
[4, 5, 6].
[7, 8, 9, 10, 11, 12].
, . 10-20 .
, , . H. krylovii : , , ,
, ; - , .
Helictotrichon krylovii .1.
H. krylovii
3 9 .
2- , . . 1-4 , 1,
().
.
9 . , .
2-3 I III . 2 1 14 , . , , , .
. 1. H. krylovii
( )
36
________________________________________________________________
, .
. 19 , 1
. , , .
,
1-2 .
, . , 3 4
.
. 30 . , .
2-3 . 2 5. . 60 . ,
.
. ,
1. .
2-3 ,
. . . , , .
. ,
.
II , -.
.
3
: , , .
, ,
57,14 % . - (v).
(g2). (g3).
. , .
H. krylovii , , (.2).
1
0,8
0,6
0,4
0,2
0
j
im
g1
g2
g3
ss
min
max
. 2. Helictotrichon krylovii
37
______________________________________________________________
[2] , . H. krylovii
- ( I=1,10 I=3,07; I=0,75 I=2,85 ). , (=0,27 0,32, =0,43 0,59). , , 1,9 4,6 0,5 2.
H. krylovii
. 3 9 .
3 : , .
H. krylovii , , .
1. .. :
. , 2003. 64 .
2. .., .. ( ). : , 1998. 413 .
3. .., .., .. . . , 2005. 108 .
4. . .2. : , 1990. 361 .
5. . : , . ,1974. 544 .
6. (). .1: . : , 2000. 256.
7. .. / ..
// . . . 3. . .; .: - , 1950. . 6. . 7-204.
8. .. // , 1975. 2. . 7-34.
9. ( )./ .., .., .. .. ., , 1976. 217 .
10. .., .., ..
// . .: , 1991. . 5-22.
11. .. , // , 2001. 1. . 3-7.
12. .. . -: ,
1995. 224 .
e-mail: raika60@rambler.ru
Institute for Biological Problems
of Cryolithozone SB RAS, 41 Lenin
Ave., Yakutsk, 677980
2)
e-mail: Sandren_1601@mail.ru
38
________________________________________________________________
: 581.9 (470)
IN SITU
..
..
. .. ,
190000, -,
. , 42-44
e-mail: t.smekalova@vir.nw.ru
i.chukhina@vir.nw.ru
() in situ
, .
674 47 .
Poaceae (167 ), Fabaceae (112), Rosaceae (81).
.
:
(), in situ , , .
,
- ,
, ().
( ), ,
.
()
, , ,
. - ,
, , - .
. ,
,
. . , .
. . , 30- -
, - , .
[2, 3,4, 10,12 .]. ..,
, , .
, . , , ;
.
.., .., : ,
, , -
39
______________________________________________________________
, ,
[9, . 4]. , ,
( ).
613 , 130 36 .
.. () ,
, ,
, [5],
.
..
, , : , - , , ,
,
[6]. 183 . 763, 211 53 ,
405 , , (), 145 ,
, .
1990- (),
,
, .
,
. , in situ ( )
, .
. , -
, .
, .
, , , [16]. , , ( ), (GP1),
(GP2), (GP3). ,
.
,
,
.
,
4 , [17] 5 [11,13,14]. , , , , ;
1 2, 1
4 [18, p.27; ..]; , ,
40
________________________________________________________________
( - ), , ,
. [15, .].
, ,
; () ( ), - .
, -
, -
[1].
,
, .
,
, , . in situ , , (
, , ),
.
,
,
.
1644 115
47 .
674
167 33 .
Poaceae (167 ), Fabaceae (112), Rosaceae (81), ,
, , .
, 76 . 152 , 18 (Agropyron tanaiticum Nevski, Agrostis
korczaginii Senjan.-Korcz., Avena aemulans Nevski, Rosa microdenia Mironova, Lotus
zhegulensis Klokov .) 13 (Agropyron lavrenkoanum Prokudin,
Agrostis salsa Korsh., Elymus uralensis (Nevski) Tzvel., Trifolium borysthenicum Gruner,
Rosa grossheimii Chrshan. .).
(. 1),
-, -, , - -.
401
(59,5%). :
(189), - (187), (179),
(171), (170). Elytrigia repens
(L.) Nevski 38, Vicia cracca L. 37, Rubus saxatilis L. 36, Alopecurus pratensis L.,
Poa pratensis L., Amoria repens (L.) C.Presl 35 46
.
41
______________________________________________________________
1
-
-
-
-
-
-
- -; . ; .
; .
. ;
. ; .;
.
, , .
.
, , , ,
.; . (
); . - . .
, , .;
; ; . . ; . . ; .
, , , , , , , , .;
; ; .
( ); ,
, . , . .
. . ; .; . , -;
, .
, . . ;
. . ;
; .
. . ; .
105
169
215
232
241
264
244
277
366
354
283
240
in situ, , ,
(2008). 12 (. 2). 7
() [8, 9].
( , - ) 232 (34,4%).
3 , ,
, . :
;
, , ;
, , ;
, , ;
, , .
42
________________________________________________________________
2
, ,
Papaver lapponicum
(Tolm.)Nordh.
(),
(), (), ()
(2V), (R), (R),
.
(3R)
(R), .
(2V), (I), (V),
.,
, .
(3R), , , .
(2V), .,
.
(3R), , ,
.
(2V), (R), (R), ,
, .
(0Ex), .
(3R), .
(3R), (V), ,
, ,
.
(3R), (E), (R), , , , , , .
(3R), ,
, , , ,
.
(V), (V), .,
, .
(3R), (I), ., ,
43
______________________________________________________________
4. .. . 3- . .: , 1971. - 752 .
5. .. ( ). ., 1985, 14 .
6. .. . ( ). ., 1986. - 126 .
7. . . .: , 1988. 592 .
8. ( ) /
; ; ;
; . . . ; . .: . .
.; . . . . .: , 2008. 855. .: .
9. IUCN Red List of Threatened Plants. Cambridge, 1997.
10. .., () ..
(). ., 1975. - 70 .
11. .., .., ..
(). .-, 2006. - 46 .
12. .. ( ). .:
, 1969. 480 .
13. .., ..
.// I
, 2002. - . 265-272.
14. .., .. . , . 76, , 2005, 54 .
15. Conserving Plant Genetic Diversity in Protected Areas: Population Management of Crop
Wild Relatives (eds.: Iriondo, J., Dulloo, M.E.,and Maxted, N. 2008, CAB International Publishing,
Wallingford, p. 65-87.
16. Harlan, J.R. and de Wet, J.M.J. Towards the rational classification of cultivated plants. Taxon, 20, 1971, p.509-517.
17. Maxted, N.&Kell, S.P., 2009,Establishment of a global network for the in situ conservation
of crop wild relatives: status and needs. 266 .
18. Maxted, N., Ford-Lloid, B., V., Jury, S., L., Kell, S.P., Scholten, M., A. Towards a definition
of a Crop Wild Relative. Biodiversity and Conservation, 15 (8), 2006. P.2673-2685.
44
________________________________________________________________
631.581 : 634.745
..
,
, 634050,
. , ., 36
e-mail:suchkova.s.a@mail.ru
: , , , , , , ,
, .
, .
20 %,
11 %, 3,3 %, 130 %, 900 %,
5 %, 2,1 %, .
- ( 1500 %).
( )
1989 .
, .
,
.
.
.
, , , , . .
.
( 1000 /) ,
. .
, [1].
1998 2010 .
, (. ).
- , ,
, ,
, , .
:
7 . .. , 24 4
. 100
.
:
,
[2,3], ,
[4]. .. [5].
45
______________________________________________________________
,
.
.
III I ,
II , II III .
III I . III
II . 4 9 .
179 185 .
- .
, , , ,
.
.
4 :
1/04.
. .
3 . (0,52 ), , ,
-. 100 52,4 . 1000 23,1 .
(2,5 ). 52,8 % ().
2/04.
. .
. (0,76 ), , , .
100 76,4 . 1000 41,0 . (2,6
). 52,8 60,0 %.
4/04.
. . .
(0,64 ), -, . 100 63,8 .
1000 35,8 . 3,1 . 32,7
48,3 % .
5/04.
. .
, . (0,74 ), , ,
. 100 74,4 . 1000 37,8 .
4,6 . 45,8 %.
.
.
, ,
.
,
( ).
.
79,5 %. 69,0 % () 85,3 % ().
, ,
2.
11,2 21,1 %.
8,9 17,8 %.
2. 5,4 14,0%.
(. 1).
.
46
________________________________________________________________
57,2 %, 18,2 %
. 2.
49,4 26, 3 %, 12,6 4,7 % .
1
2000
05
2001
05
05
.
%
29,0
100
41,7
143,8
39,4
135,9
34,3
118,3
4,2
21,3
100
33,0
154,9
38,0
178,4
28,6
134,3
3,4
25,1
100
37,4
149,4
38,7
157,2
31,5
126,3
4,8
%
16,9
100
18,7
110,7
21,0
124,3
17,7
104,7
1,8
17,3
100
19,8
114,5
19,4
112,1
18,1
104,6
1,3
17,1
100
19,3
112,6
20,2
118,2
17,9
104,7
3,3
.
59,8 %.
, .
,
24,1%, 2
23,9 15,7 % .
,
,
, .
, .
:
( ).
.
92,9%.
23,2% () 28,0% ().
.
23% .
, ,
, .
59,2 %, .
68,2%. - 9,3 %.
9,1 %, 16,1 %.
,
(.). ,
III I .
.
37,4 %. 2 12,2 %,
6,1 % [6].
47
______________________________________________________________
40
35
24.05
30
,,
07.06
25
22.06
20
06.07
15
24.07
06.08
10
24.08
5
0
.
: (24.05; 07.06; 22.06; 06.07; 24.07; 06.08; 24.08).
.
, ,
. , ,
. ,
15 .
(. 2). 8 (61,0
90,0 %). 15 76,2 81,7%.
20
. 20,0 56,5%.
20 (76,0%).
.
(1-2 ) ,
.
, .
,
, ,
Tubercularia vulgaris, Cytospora sp. .
.
.
( ).
,
.
, ,
48
________________________________________________________________
. .
1. .
2. .
3. .
4. .
5. .
6. .
8
15
20
8
15
20
8
15
20
8
15
20
8
15
20
8
15
20
, %
90,0
81,7
47,0
87,0
81,6
76,0
61,0
81,1
56,5
88,0
78,9
20,0
83,0
76,2
24,0
71,0
77,7
29,0
,
.
32,4 2,7
27,52,0
16,7 2,9
31,0 2,6
28,82,1
26,8 3,0
18,7 2,0
15,8 1,9
16,9 4,3
24,8 2,0
16,8 1,4
2,5 0,6
15,4 1,7
12,9 1,1
11,4 3,9
13,3 1,2
9,3 0,8
2,0 0,4
15,5 0,4
13,50,1
11,9 0,8
14,8 0,8
12,9 0,9
10,0 0,8
11,9 0,7
10,9 0,6
10,2 1,1
13,4 1,0
11,2 0,9
5,7 1,9
7,7 1,4
6,9 0,8
5,7 1,7
8,7 0,9
6,9 0,5
1,9 0,7
,
( Cercospora opuli), (Botrytis cinerea) (Microsphaera hedwigii). ,
,
Microsphaera sparsa Cercospora viburnicola, .
, , ,
. , ,
.
, [7].
,
, .
: , , , , ,
. 1/04, 2/04, 4/04,
5/04.
. , ,
2.
5,4 21,1%, (18,8 51,6 %)
.
(26,3 57,2 %), (4,7 18,2 %).
(47,2 50,2 %). 15
, 20 .
49
______________________________________________________________
1. .., .., .., ..
// . 1987. .2. . 178 -189.
2. , . /
. . .. , . .., 1973. 496 .
3. , . /
, . .., .. : , 1999. 608 .
4. , /
, . .. : , 1995. 502 .
5. .. . .: , 1985. 351 .
6. ..
/ .. . . .-. . ,
2006.18 .
7. .., .. (Viburnum opulus L.)
// / . VIII .
-. ., 1. : - , 2008. . 147-149.
S.A. Suchkova
Tomsk state university,
the Siberian botanical garden,
634050, Tomsk,
Lenin's avenue, 36
e-mail:suchkova.s.a@mail.ru
Results of researches on introduced a guelder-rose ordinary (Viburnum opulus L.). Analytical selection on creation of a local genofund is
conducted. Perfect forms are allocated. To the accelerated reception of a
landing material of a guelder-rose it is applied green cuttings. Dependence of rooting of shanks and quality of received saplings from the
high-quality accessory, applied regulators of growth and ways cultivations is established.
Keywords: a guelder-rose ordinary, introduction, selection,
grades, perfect forms, reproduction, growth factors, saplings, illnesses.
50
________________________________________________________________
581.4+581.1
.. 1
.. 2
1)
440046,
. , . , 40
e-mail: coko@sura.ru
2)
, 440026,
. , . , 40
. ,
.
: , , , , .
e-mail: sef1957@mail.ru
[20], .
. ,
, : ( ), () , , (). ,
[15].
. [8]. ,
, . ,
, .. ,
[14]. .
.
(Linum usitatissimum L.) - , , 2- 5 .
, - .
558718 , 10
22082910. ()
51
______________________________________________________________
0,8 1,9. ( = 0,8), ( = 1,2; 1,1) ( = 1,9;
1,4). , . [12].
, [7, 19]
( ), ( ), ( ) ( ) (. 1, . 1).
()
, .
30 .
5
1-3
5-7
6-8
14-25
10-13
9-12
6-14
5-7
4-8
6-12
10-15
15-25
10-20
. 1. ,
52
________________________________________________________________
Linum usitatissimum L. , 30 , . 5 . 1000
4,217,04 . , 2,7 13 [16]. , 3,4-4,2 ,
1,8-2,4 , 0,5-1,1 , , , , . , , -, . , , - [6, 11].
: , . . 12% [9, 10].
, ,
. , , ( ).
, , [4].
36 56
. :
, , .
. 1-3 . () , , .
7-8, , 2-5
, , ,
. 6-8 .
. 6-7
(). , .
,
.
- 8
23 . . ( ) 0 6,15.
, 6-10 . 1-2
.
8-10 5-6 .
,
. . ,
, . . . 2,1 12 2,9 29 . 70 120 . , , -
53
______________________________________________________________
-, -, , ,
, .
.
40 68 , , . 9-12 .
, . , , . , , , . . , ,
. 9-12
, , , .. 4560- .
,
. , .
, , -,
3 , -, 5-6
, 2-4 . .
, , , , , 12-15
, 6-8 , -, ,
, . ,
( ), , , , -. - [1].
( 5-6 ) ,
. 9-10 ( ). ,
15-25 . . , ,
, .
. [3],
20-22. . , , .
. , . , .
,
. , . .
10 . . .
, , .. .
25 42 . [2, 5, 13,
54
________________________________________________________________
17]. : , ,
. , . ( ) - , (,
) ( , ). () ()
.
, , , (6 5
) (11 8,5 ). .
, 10 , . . .
,
, (. 2).
85 93 ,
73 . 107 . : ,
. , .
. 2.
25-45 .
( ,
). , . , 6-12%.
, ,
55
______________________________________________________________
,
, Linum usitatissimum L.
, .
1. / . .. . ., 1963. 504 .
2. .. . . . .-. , 1962. 157 .
3. .., ., .. . .: .
1971. 110 .
4. .. / . .
. .: . 1954. . 147-164.
5. .. ,
,
: . . .-. :
05.01.79. , 1978. 25 .
6. .., .., .. . . .: , 2005 344 .
7. .. .
. .: , 1977. 288 .
8. .. . .: , 1984. 240 .
9. . .: , 1988. . 40-41.
10. . .: , 1992. . 59-60.
11. / . . .. .: . 1967. 583 .
12. : (Linum usitatissimum L.): / .:
.. , .. . ., 1988. 29 .
13. .. . .: . - .-. . 1957. 179 .
14. .., .., .., ..
/ . 1968. . 37, . 4. .12-25.
15. .. . .: - , 1992. 352 .
16. / . .
.. . .: . 1967. 351 .
17. .., .. /
. 1992. 4-5-6. . 25-27.
18. .. /
. . ., 1937. . 5-26.
19. ., .., .. . - : . . . / .: . ., 1990. 191 .
20. .., .., .. .
. .: , 1982. 93 .
e-mail: coko@sura.ru
2)The
e-mail: sef1957@mail.ru
56
________________________________________________________________
574.32:633. 262 (282.256.65)
(Bromopsis
inermis (Leys.) Holub.) - .
- , .
: , ,
, IVC.
.
, ,
.
,
[1].
.
: ,
; ,
- ; - [1].
: Bromopsis inermis (Leyss.) Holub - .
(Bromopsis inermis (Leys.) Holub.) , , , .
, , [2].
. 7
, , , .
.. . (. 1).
.
: , b c, .
Q:
1. Q = ( + b) > ;
2. Q = ( + b) = ;
3. Q = ( + b) < [1].
IQ = ( + b) / 2. > 1 -
57
______________________________________________________________
, < 1 , 1,
, .
.. , .. [3]
(IVC),
. :
N
X X
i
IVC
i 1
(1)
- i- ,
, N - .
- i 1
Bromopsis inermis (Leys.) Holub
1.
( )
2.
( )
3. ( )
4.
( )
5 ( .
)
6.
( )
7.
( )
Vicia crassa L., Trifolium
pretense L., Galium verum
L.
Elytrigia repens (L.) Nevski, Hordeum brevisubulatum (Tin) Link, Poa pratensis L.,
Alopecurus arundinaceus
Poir.
Elytrigia repens (L.) Nevski,
Poa pratensis L., Artemisia
commutate Bess., Thalictrum simplex L.
Trifolium pretense L.,
Chamaenerion angustifolium (L.) Scop, Rosa acicularis Lindl., Thalictrum
simplex L.
Elytrigia repens (L.),
Hordeum revisubulatum
(Tin) Link, Alopecurus
arundinaceus Poir.
Geranium ptatense L.,
Trifolium pretense L.,
Veronica longifolia (L.)
Equisetum pretense (L.),
Trifolium pretense (L.),
Elytrigia repens (L.),
Galium verum (L.)
(
)
(
)
61,75
13,09
62,54
12,83
52,34
10,47
54,9
12
60,5
12,76
61,33
11,95
61,64
2,52
:
, , , 1,2 3 , 1, 2, 3 , ,
58
________________________________________________________________
, , ,.
30 .
, ,
.
Bromopsis inermis
,
.
, -
Bromopsis inermis , : 1,
2, 4, 5, 6, 7, 3 (.2).
2
Bromopsis inermis (Leyss.) Holub.
( )
1
2
3
4
5
6
7
(Q/)
3,25
3,25
0,65
7,5
7
7
1
(IVC)
0,96
0,99
0,87
1,04
1,11
0,99
0,99
1, 2, 4, 5. ,
, IVC Q.
6 7 b .
, ,
.
3,
, , .
, 5 4, (1,11 1,04 ). 3, - ,
(0,87) Q 0,65.
, 2, 4, 5, 6 ,
3 , 1 7 .
, Bromopsis inermis, 2, 4, 5, 6.
, 1 7,
( , )
3.
Bromopsis inermis (60 65 ) (11 13 ). -
59
______________________________________________________________
- ,
, [4].
80
74
70
66
63
58
57
60
50
43
42
40
35
31
27
30
20
13 13
b
c
25
13
7
10
40
0
1
. Bromopsis inermis
% (: , , b )
1. .. . , 1989. - 146 .
2. . . .
.; . - 1980. .1. - . 38 - 41.
3. .., .. //
. VII . . 16-21 2004.
, 2004. . 2. - . 113 - 120.
4. .., .., .., ., .., .. .
, 2005. - 108 .
60
________________________________________________________________
635.954:633.82
(PLECTRANTHUS AMBOINICUS (LOUR.) SPRENG. )
..
..
-
,
127216, ., . 7
e-mail:fitovit@gmail.com
Plectranthus amboinicus (Lour.) Spreng., - 100, 500, 2000 .
,
2000 . 500
55% 1 / .
: Plectranthus amboinicus, ,
, ,
61
______________________________________________________________
, , . ,
, , . -, , , , . 2-10 , 1.5-8 .
. : () - 2000 , 500 , 100 , (
) - - , 1- 1 /, 2 2 /, 3 3 /. 4 .
1 . : 1 , 1 1 . , (-400) (, ) 2000, 500 100
12 .
[7].
-
[8]. ( , ) [9].
15-20
. , . 3-4
, . , 2000 .
,
. , ( ), 46-49% , ( 1).
1
P. Amboinicus
,
,
,
2000
500
100
2000
500
100
2000
500
100
2000
500
100
23,85,2
13,86,1
0,0
6,80,3
3,50,6
0,0
8,20,3
4,20,7
0,0
5,80,2
3,10,5
0,0
1
28,54,2
17,83,0
0,0
6,50,5
4,50,5
0,0
9,50,2
6,50,3
0,0
6,90,2
5,00,2
0,0
2
25,43,0
13,23,5
0,0
6,80,6
3,50,6
0,0
8,60,2
5,60,2
0,0
6,20,1
4,00,2
0,0
3
14,14,0
10,92,4
0,0
5,50,6
4,00,4
0,0
5,40,4
4,70,2
0,0
4,00,3
3,40,1
0,0
25-40
100 . -
62
________________________________________________________________
100
1,5-2,5 , 500 - 3,5-5 .
1 2
3 .
.
.
2
P. amboinicus
, 2
2000
500
100
05 2,74
05 4,75
2000
500
100
1
2
3
21,4
23,4
22,5
14,2
9,7
14,4
9,9
9,5
1,0
1,0
1,0
1,0
10,7
12,9
11,1
8,2
469,1
138,2
0,0
05 2,38
20,4
10,9
1,0
10,8
1
2
3
549,6
511,8
215,2
214,0
103,4
91,3
0,0
0,0
0,0
05 , 05
, 05 .
,
. , 2000 , ,
( 239%), , ( 2). 1 / , 500
1,5 .
500 , . 1 /
,
( ) 29-61%.
55%.
2 3 / .
, .
1,42 % 2000 ,
:
500 - 1,03 % , 100 -0,96%.
63
______________________________________________________________
. 1 /
500 55% .
, 2000 . (1-2 ) 500 ,
.
1. .., .., .., ..
Plectranthus // .- 2008.-.280.-.251-255.
2. ..
// . . (15-16 2006).-.: - -, 2006..82-85
3. .. . - .: - .-, 1982.- 384 .
4. .., .. . (
).-. : -, 2000.- 112 .
5. Lukhoba C.W., Simmonds M.S.J., Paton A.J. Plectranthus: a review of etnobotanical uses //
Journal of Ethnofarmacology.- 2006.-V.103.- . 1-24.
6. Gurib-Fakim A., Brendler T. Medicinal and Aromatic Plants of Indian Ocean Islands.Sttutgart: Medpharm, 2004.-568 p.
7. .. , // . 21 : , : . . . , 18-21 2009 .-.: "", 2009. .284285.
8. : .2. .
/ . - 11 ., . - .: , 1989. - 398 .
9. .. . .:
, 1984. - 424 .
64
________________________________________________________________
581.145.2 + 582.772.2
.O.
. .. ,
127276, ,
., 4
Dipteronia 2
Aceraceae.
Dipteronia
sinensis Oliv. Butia -. 2 ,
.
e-mail: i_o_yatzenko@mail.ru
: Dipteronia, Aceraceae, , ,
Aceraceae Juss.
,
- . Dipteronia Oliv. (2 ) Acer L. ( 150
, ) [4]. . Dipteronia 2 , , Acer . ,
, Dipteronia , B. A. Hall [6] , Acer
Acer Dipteronia,
Dipteronia Acer [7], [8], [9]. Aceraceae . Dipteronia ( Acer) , ,
.. .
.
Dipteronia sinensis Oliv.
Dipteronia sinensis
. [2].
( 1.)
, , ,
, 2(3) .
65
______________________________________________________________
22-26 48-55 . 2 . 24-27
22-24 . , ,
4 11 . 5 1,5-2 .
( 1.;.)
(18-21 ) -, .
, , . , . .
16-19 , 2 .
10-12 . , . 4-5 ,
. .
, , . .
Aceraceae
. Dipteronia , , Acer . Dipteronia Butia- . [3].
Acer [5]. Butia-
Acer, ( 1.;.):
(A. barbinerve Maxim., A. campestre L., A. glabrum Torr., A. laetum C.A.Mey., A. miyabei Maxim., A. mono Maxim., A. negundo L., A. palmatum Thumb., A. platanoides L., A. pseudoseboldianum (Pax.) Kom., A. spicatum Lam., A. tegmentosum Maxim., A. tschonoskii Maxim.)
1-3 , A. micranthum Siebold & Zucc., A. monspesslanum
66
________________________________________________________________
L., A. velutinum Boiss. [1], [5]. , Acer
, Olea- A. ginnala A.
semenovii, [5], Physena- A. ukurundense Trautv. & May. [1]. , ,
Dipteronia Acer, Butia- (A. micranthum, A. monspesslanum, A. velutinum). Dipteronia Acer - :
Dipteronia 1/8 1/10 , - 1/6 3/5. Acer Dipteronia , Dipteronia .
Dipteronia .
2 : ,
, . Dipteronia
Butia-.
Dipteronia Acer, , .
1. . ., . . // ..
- . -1951. .7, .2. .170-192.
2. . ., . ., . ., . ., . ., . . .: , 2000. - 127 .
3. . ., . ., . . Magnoliophyta. - .:
URSS, 2009. - 406 .
4. . . . : , 1966. 611 .
5. . .,
6. Hall B.A. The floral anatomy of Dipteronia//Amer. J. Bot. - 1951. - Vol.38 - P.:793-799.
7. Li J., Yue J., Shoup S. Phylogenetics of Acer (Aceroideae, Sapindaceae) based on nucleotide sequences of two chloroplast non-coding regions//Harvard Pap. Bot. - 2006. - Vol.11(1) P.101-115.
8. Pfosser MF, Guzy-Wrbelska J, Sun BY, et al. The origin of species of Acer (Sapindaceae)
endemic to Ullung Island, Korea//Syst. Bot. - 2002. - Vol.27 P.351367.
9. Tian X, Guo ZH, Li DZ. Phylogeny of Aceraceae based on ITS and trnL-F data sets//Acta
Bot. Sin. - 2002. - Vol.44 - P.714724.
Dipteronia is one of 2 genera from Aceraceae. Anatomical structure of Dipteronia pericarp have never been studied. Our studie reveled
morphogetetic fruit type of Dipteronia sinensis Oliv. as pirenarium Butia-type. Pericarp consists of two zones: outer arenchyma zone developed from ekzocarp and outer layers of mezocarp and inner drupel developped from inner layers of mezocarp and endocarp.
Key words: Dipteronia, Aceraceae, carpology, fruit type, pericarp
anatomy.
67
______________________________________________________________
581.145.2 + 582.632.1 + 575.86: 582.632.1
CORYLUS L. SUBSECT.
CORYLUS (BETULACEAE )
..
. .. , 127276,
, ., 4
e-mail:
olga.ystenko.msu@gmail.com
,
6 150 . -. : Corylus L., Carpinus L., Ostrya Scopoli, Ostryopsis Decne. Alnus Mill., Betula L.
[7] Betulaceae
: Coryloideae Betuloideae. Coryloideae Carpinus, Corylus, Ostrya Ostryopsis, Betuloideae Alnus Betula.
[8]. Kuprianova [9] Carpinaceae (Carpinus, Ostrya, Ostryopsis), Corylaceae (Corylus) Betulaceae (Alnus, Betula).
Abbe [6] Coryloideae Carpineae
(Carpinus Ostrya) Coryleae (Corylus Ostryopsis). Takhtajan [10] Corylaceae Betulaceae s.str. (Alnus Betula) ,
: Carpinoideae (Carpinus, Ostrya, Ostryopsis) Coryloideae (Corylus). Corylus : Acanthochlamys Corylus Corylus, Colurnae, Siphonochlamys.
, [12].
Corylus , ,
[2], .
. Corylus
, [3, 11]. Corylus Corylus Coryloideae.
2 ( 4) sect. Corylus subsect. Corylus:
C. avellana L., C. americana Marshall.
[1].
68
________________________________________________________________
Sect. Corylus
Subsect. Corylus
Corylus americana Marshall
(.1.) 2 5, , , ,
. , -, ,
10-15 .
. .
38-42 (.1.). -: 2/3 ,
, ,
.
() . 1-3
. ( 11 ) ,
. . .
Corylus avellana L.
1 5 (.1.). , 3/4 . ,
, 15-20 . 12-20 . .
C. avellana (.1.) C. americana,
30-35 .
, 2/3 ,
, C. mericana - , 3-4
.
C. americana.
. [2] , ,
- . Corylus . . , . orylus Corylus-.
Corylus Coryloideae [4, 5],
Carpinus, Ostryopsis (.1.) Ostrya (.1) ,
. ,
. Ostryopsis, Carpinus Ostrya
, .
Carpinus, Ostryopsis Ostrya Olea. Carpinus, Ostrya
Ostryopsis
Olea- Corylus-.
. , Corylus
, Corylus -
69
______________________________________________________________
. , [7] Corylus
. ,
Olea- Coryloideae,
Corylus .
Corylus Corylus - ,
.
Corylus. ,
Coryloideae Corylus .
1. . ., . ., . ., . ., . ., . . . - .: , 2000. - 127 .;
2. . ., . ., . .. Magnoliophyta. .:
URSS, 2009. 283 c.;
3. . .. Betulaceae // . . (.).
. . 3. .: , 1991. . 134140;
4. . ., ;
5. . ., . . , . . . Ostrya virginica Willd. (Betulaceae s. l.) // . . . . . . ., . 100-
. - .. . 2009, . 674-677;
70
________________________________________________________________
6. Abbe E. C. Flowers and inflorescences of the Amentiferae // Bot. Rev. 1974; - V.40.
P. 159261;
7. Chen Z.-D., Manchester S. R., Sun H. Y.. Phylogeny and evolution of the Betulaceae as inferred
from DNA sequences, morphology, and paleobotany // Amer. J. Bot. Vol. 86. 1999.
P. 11681181;
8. Hutchinson, J. The genera of flowering plants. Oxford: Claren.Pr. 1967. Vol. 2.
P.124126; 132134;
9. Kuprianova L. A. On a hitherto undescribed family belonging to the Amentiferae // Taxon.
Vol.12. 1963. P.1213;
10. Takhtajan A. Diversity and classification of flowering plants. New York: Columbia University Press. 1997;
11. Vaughan J. G. The structure and utilization of oil seeds. L.: Chap. & Hall LTD. 1970.
279 p.;
12. Whitcher I. N. and Wen J. Phylogeny and Biogeography of Corylus (Betulaceae): Inferences from ITS Sequences // Systematic Botany. 2001. Vol.26(2). P. 283-298.
71
______________________________________________________________
581(075.8) (471.51)
1
..
..
..
,
, .
, 426034,
, . ,
. , 1
.
: , , .
e-mail: ob@uni.udm.ru
olga_dedyukhina@mail.ru
yaloyagov@mail.ru
, [4, 7]. ex situ .
(IPGRI) :
1. ;
2. , in vitro ( ,
);
3. .
()
[1, 2, 5, 6, 14, 15, 16].
[17].
, . , [11].
, . 3 7 . [11].
.
,
( ) . , 56 .
(2007). . 1. ,
1
. 2.2.3.1. 3997
(20062008 ) 2.2.3.1/3578 (20092010),
-
(2005
2009 .) ( ).
72
________________________________________________________________
,
[9, 10]. ,
,
: -, , , .
, ,
12
[11].
, , , ,
[8, 12]. 5 [1, 5]. , 9 (Betula humilis Schrank, Betula nana L., Calipso bulbosa (L.) Oakes, Convallaria majalis
L., Cypripedium calceolus L., Larix sibirica Ledeb., Paeonia anomala L., Petasites frigidus
(L.) Fries, Populus alba L.) 56 ,
(13 ), [12].
, ,
, (Adonis vernalis, Digitalis grandiflora, Delphinium cuneatum, Hypericum elegans, Iris pseudacorus, Iris sibirica, Plantago maxima, Scleranthus perennis). . , , Senecio erucifolius, Gentiana pneumonanthe, Lythrum virgatum. (. 1).
1
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
Adenophora liliifolia (L.) A. DC.
Adonis vernalis L.
Anemone sylvestris L.
Althaea officinalis L.
Artemisia armeniaca Lam.
Artemisia dracunculus L.
Artemisia latifolia Ledeb.
Aster amellus L.
Astragalus falcatus Lam.
Allium schoenoprasum L.
Briza media L.
Carex bohemica Schreb.
Campanula sibirica L.
Carex flava L.
Centaurea sumensis Kalen.
Delphinium cuneatum Stev. ex DC.
Digitalis grandiflora Mill.
Eremogone longifolia (Bieb.) Fenzl
Eremogone saxatilis (L.) Ikonn.
Galeobdolon luteum Huds.
Gentiana pneumonanthe L.
(3)
(1)
(3)
(1)
(4)
(1)
(2)
(3)
(3)
(1)
(3)
(3)
(3)
(2)
(3)
(3)
(1)
(2)
(3)
(1)
(3)
++1
++
++
++
+
+
+
+
++
++
+
++
++
++
+
++
++
++
++
++
++
+/+/+
+/+/+/+/+
+/+/+
+/+/+
+/+
+/+/+
+/+
+/+/+/-
73
______________________________________________________________
. 1
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
38
39
40
41
42
43
44
45
46
47
Geranium palustre L.
Galium tinctorum (L.) Scop.
Herniaria glabra L.
Hypericum elegans Steph. ex Willd.
Iris pseudacorus L.
Iris sibirica L.
Laser trilobum (L.) Borkh.
Lilium martagon L.
Lychnis chalcedonica L.
Lythrum virgatum L.
Melica altissima L.
Plantago maxima Juss. ex Jasq.
Potentilla humifusa Willd. ex Schlect.
Pulsatilla patens (L.) Mill.
Pulsatilla uralensis Turczh
Salix lapponum L.
Salix myrtilloides L.
Salvia tesquicola Klok. et Pobed.
Scleranthus perennis L.
Senecio erucifolius L.
Stipa pennata L.
Terhroseris integrifolia (L.) Holub
Trifolium lupinaster L.
Thymus ovatus Mill.
Thymus marschallianus Willd.
Vaccinium uliginosum L.
(3)
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, , 1, ,
(66 %)
. , Galeobdolon luteum, Centaurea sumensis ,
, [12]. ,
in vitro, (Delectus).
, [17], , , Melica altissima, Anemone sylvestris; , (Pulsatilla patens, Pulsatilla uralensis, Adonis vernalis), , (Artemisia latifolia, Artemisia armeniaca, Artemisia
dracunculus, Herniaria glabra, Petasites frigidus) (. 1).
in vitro,
. , [1, 4, 7].
, ,
,
- [3, 8, 13]. , , ,
(g3) Pulsatilla uralensis -
74
________________________________________________________________
- , , (. 2).
2
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ex situ . . (1974) . . (2007)
g1
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g3
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, .
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(m), (CV, %).
, ,
. , ,
, , .
1. . ., . ., . . // . . . . 2010. . 2.
. 4855.
2. . ., . . Pulsatilla
flavescens // . 2008.
. 10 (24), 2. . 380388.
3. . . // . 1974. . 59, 6. . 826831.
4. . BGCI : Richmond, U.K., 2002. 16 .
5. . ., . . // Ii i :
i . . . / . . . . . , 2006. .4951.
6. . .
// : V .
. / . . . . , 2009. . 270273.
7. . ., . .
// : , : , 70- 70- .. (10-14 2009., -) /
. .. . -: , 2009. . 165167.
8. . . ( ). .:
, 1981. 96 .
9. //
. 1981. . 119. . 9496.
10. , , ,
// 31 5.03.2007. (-
75
______________________________________________________________
20.03.2007 N
RU18000200700037).
11. ..
// /
. , 1982. . 6067.
12. . . - . .: ,
1991. 216 .
13. . . // . : . . (58 2007 .) / . . . .
. ., 2007. . 630632.
14. . ., . . Pulsatilla
Mill. // , . . 2009. . 151, . 4. . 112123.
15. .. Pulsatilla flavescens (Zucc.) Juz.
// : .
. , 2007. . 188190.
16. .. Pulsatilla Mill. // : . . , 2010. . 170174.
17. www.bioversityinternational.org
76
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582.931.4 (581.47 + 581.8)
FORSYTHIA
OLEACEAE
..
, 129243,
. , . ,
.6, . 5,
e-mail:
avfilonenko@yandex.ru
Forsythia Vahl.
. , ,
. , Forsythia , . Forsythia
Forsythia-, . Oleaceae. .
: Forsythia, Oleaceae, ,
, Forsythia-.
77
______________________________________________________________
Forsythia ,
(. 1 ). , , 1 10;
F. europaea 2 ( 10). -,
, , 1,8 1,2
(. 1 ). Forsythia
, . -
. Forsythia -, , . , (. 1 ).
. 1. Forsythia europaea:
; (
); ( , , ); ;
( ).
Forsythia . Forsythia -, - ;
, .
( ) 68 , F. viridissima Lindl.
12 . , (, , ). ( - ),
, , .
, . ,
(
). -
78
________________________________________________________________
, .
, F. suspensa Vahl, . ,
48 (
F. giraldiana Lingelsh. 15), , . . , , ,
. , , . . -
,
, .
Forsythia , , - (. 1 , ).
, (F. giraldiana, F.
suspensa, F. viridissima) (F. europaea).
Forsythia . 412 ,
F. viridissima 15 . . .
(23 ), , (. 1 ). , , , , . ,
(F. europaea) (. 1 ,
).
.
, , Forsythia .
,
Forsythia Forsythia- [10]. ,
Forsythia
. ( 5 ) , Oleaceae,
. , Forsythia Forsythia-,
Oleaceae. , , . , , , Forsythia- Olea-. Olea-
79
______________________________________________________________
, . Fraxinus L., Abeliophyllum Fontanesia Labill. , , . , ( ,
) Galanthus-,
,
.
1. Hebb R. S. Thestory of Forsythia // Arnoldia. 1971. V. 31. P. 4163.
2. Kim K. J. Molecular phylogeny of Forsythia (Oleaceae) based on chloroplast DNA variation
// Plant Systematic and Evolution. 1999. V. 218. P. 113123.
3. Knoblauch E. Oleaceae // Die naturlichen Pflanzenfamilien / Ed. A. Engler. Leipzig,
1895. Vol. IV. P. 1-16.
4. Taylor H. Cyto-taxonomy and phylogeny of the Oleaceae // Brittonia. 1945. Vol. 5. P.
337367.
5. Johnson L. A. S. A review of the family Oleaceae // Contribution New South Wales National Herbarium. 1957. Vol. 2. P. 396418.
6. . . Oleaceae //
c . 2002. . 34. . 138150.
7. Wallander E., Albert V. A. phylogeny and classification of Oleaceae based on rps16 and
trnL-F sequence data // American Journal of Botany. 2000. Vol. 87, 12. P. 18271841.
8. OBrien T. P., McCully M. E. The study of plant structure: principles and selected methods.
Melbourne, 1981. 342 p.
9. . ., . ., . . . . . .: , 2004. 312 .
10. . ., . ., . . Magnoliophyta. .,
2009. 400 .
A.V. Filonenko
Moscow State Pedagogical
University, 129243, Moscow,
Kibal`chicha st., 6-5, Department
of Botany
e-mail: avfilonenko@yandex.ru
80
________________________________________________________________
: 582.783
VITACEAE
.
..
,
690024, . ,
. , 142
e-mail boteco-bsi@yandex.ru
2 , . Partenocissus 1
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4 .
, 2-3 .
: , Vitaceae.
81
______________________________________________________________
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, .
-, , , , . ,
[8].
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.
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. ,
.
82
________________________________________________________________
. c , P. quinquefolia, , , .
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%
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P.tricuspidata
A. aconitifolia
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7,0 (P.tricuspidata) 15,0 (P. quinquefolia)
, 1 4 , , (1,6-3,5 ) . , , - , 1,7-2 1,24-1,6 , P.tricuspidata , A. aconitifolia . - .
20-25 A. aconitifolia
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. .
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,
. ,
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83
______________________________________________________________
. , 120.
, . .
A. aconitifolia .
- , 6-7 - .
,
. . . . Partenocissus , . P.tricuspidata
-, P. quinquefolia -, 5, 3- . , , .
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, .
.
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, P.tricuspidata P. quinquefolia, , , .
, 15-40 0,2-0,5 , 1-5 , 10-50 . 1/3
, , 3-5 .
0,3-0,5 , 50-60% , 1-2 .
2 , . ,
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, 1/3 . ,
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. ,
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.
84
________________________________________________________________
, ,
[9, 10] P.tricuspidata .
4-5 , , . 30- A. aconitifolia 20- P.tricuspidata
.
, [11]. , 10 . P.tricuspidata 2 - -
.
,
. Partenocissus Ampelopsis 5
, .
:
3-4 . ,
, . , , ,
. , , .
, .
, , . A. aconitifolia 1
. , .
, . ,
,
.
1. : . . : , 2008. 688 .
2. Young J.A., Young C.G. Seeds of woody plants in North America. Portland, 1992. 407 p.
3. .. Vitis L. // . . .
2003. .60, 2. . 164-169.
4. Galet P. Etude morphologique et systematique des plantules de Vitacees // Naturalia monspeliensia. Ser. Bot. 1967. Fasc.18. .231-269.
5. Sadao Asano Seeds/Fruits and Seedlings of Plants in Japan. Tokyo: Zenkoku-NosonKyoiku-Kyokai, 2006 . 82-83.
6. .. // . . .3. . 1950. . 6. .7-204.
7. .., .., .., .. : . . . .: , 2005. 256 .
85
______________________________________________________________
8. Corner E.J.H. The seeds of dicotyledones. Vol.1. London, N-Y, Melbourne: Cambridge Univ. press., 1976. 311 p.
9. Millington W.F. The tendril of Parthenocissus inserta: determination and development. //
Amer. J. Bot. 1966. Vol. 53. P.74-81.
10. Schah J.J. and Dave Y.S. Morphohistogenic studies on tendrils of Vitaceae // Amer. J. Bot.
1970. Vol. 57 P. 363-373.
11. .. Vitaceae Juss. // : V . . (, 20-23 2009 .)
: , 2009 . 249-252.
86
________________________________________________________________
630.181.8:634.18(470.57-25)
SORBUS L.
..
..
,
,
e-mail: rimmaabdullina@yandex.ru
15
10 2002 2010 . ,
(
). ,
Sorbus, ,
.
: Sorbus, , .
,
,
,
. .
, , .. [1978, 1981, 1984].
15 2002 2010 10 : , , , , , , , , , .
, .. [1981],
. ( ) ( ) . , 7 (4 )
() ,
.
S. sibirica, S. decora S. aucuparia ,
Sorbus. S. x arnoldiana S. aucuparia. S. amurensis
S. commixta , S. turkestanica . S. aucuparia S. sibirica -1,
87
______________________________________________________________
, .
5,
(+1 0), .
.
S. sibirica Hedl.
S. aucuparia L.
S. commixta Hedl.
S. amurensis Koehne
S. x arnoldiana Rehd.
S. decora (Sarg.) Schneid.
S. turkestanica (Franch.) Hedl.
S. rufo-ferruginea (Schneid.) Schneid.
S. graeca (Spach) Hedl.
S. thuringiaca (Ilse) Fritsch
S. hybrida L.
S. mougeottii Soy.-Willem. et Godr.
S. intermedia (Ehrh.) Pers.
S. armeniaca Marsh.
S. intermedia (Ehrh.) Pers. var. arranensis (Hedl.) Rehd.
-0,849
-0,753
-0,697
-0,21
-0,21
-0,14
-0,129
0,173
0,317
0,415
0,468
0,486
0,527
0,614
0,625
4
4
4
4
4
4
4
5
5
5
5
5
5
5
5
( ). ,
Sorbus, , .
1. .. // . . . . 1974. . 94. . 3-10.
2. .. . .: , 1978. - 146 .
3. .. . - .: , 1981. 120 .
88
________________________________________________________________
582.931.4 (581.47 + 581.8)
,
129243, . ,
. , . 6, . 5,
e-mail: avfilonenko@yandex.ru
2)
,
644099, . , . 14,
e-mail: stratiotes@yandex.ru
Comoranthus Schrebera. , , . Comoranthus Schrebera Forsythia-. Comoranthus Schrebera,
Oleaceae.
: Comoranthus, Schrebera, Oleaceae, , , Forsythia-.
89
______________________________________________________________
. . . ( )
(LE), Missouri Botanical Garden (MO) (St. Louis, MO, USA), New York
Botanical Garden (NY) (Bronx, NY, USA), Kirstenbosh National Botanical Garden (Cape
Town, Western Cape, Republic of South Africa). (90 %
= 1 : 1 : 1) 1030 (
). 70 %- .
. 715 .
; [8, 9].
. 1. Comoranthus Schrebera:
C. minor; C. minor
; C. minor ; S. macrocarpa; S. macrocarpa ;
S. macrocarpa .
Comoranthus
4 . . , , , 26 16 . Comoranthus ,
. ,
, .
Comoranthus -,
, .
, . , .
90
________________________________________________________________
, , . Comoranthus . ( 25 )
( ),
. 30 ( Comoranthus obconicus 50) , ( ,
)
, .
, ,
, . ,
, ,
, , . Comoranthus , .
Schrebera 4 , 23 ( 4 ,
, , ). ,
, , 65 32 . Schrebera , . .
Schrebera - ;
, . . Schrebera , , . . Schrebera . ( 30 )
.
. 10 ( Schrebera macrocarpa 30)
( ), . , , ,
, . , . (Schrebera platyphylla, S. swietenioides)
, , .
[10], Comoranthus Schrebera Forsythia-. ;
.
Comoranthus Schrebera
, Comoranthus Schrebera
Jasminoideae, Oleoideae.
91
______________________________________________________________
1. Wallander E., Albert V. A. phylogeny and classification of Oleaceae based on rps16 and
trnL-F sequence data // American Journal of Botany. 2000. Vol. 87, 12. P. 18271841.
2. Green P. S. Oleaceae // The families and genera of vascular plants / Ed. K. Kubitzki.
Heidelberg, 2004. Vol. VII. P. 296306.
3. Bentham G. Oleaceae // Genera Plantarum / Ed. G Bentham, J. D. Hooker. London, 1876.
Vol. II. P. 672680.
4. Knoblauch E. Oleaceae // Die naturlichen Pflanzenfamilien / Ed. A. Engler. Leipzig,
1895. Vol. IV. P. 1-16.
5. Taylor H. Cyto-taxonomy and phylogeny of the Oleaceae // Brittonia. 1945. Vol. 5. P.
337367.
6. Johnson L. A. S. A review of the family Oleaceae // Contribution New South Wales National Herbarium. 1957. Vol. 2. P. 396418.
7. Weber G. 1928. Vergleichend-morphologische untersuchengen uber die oleaceenblute //
Planta Vol. 6. P. 591658.
8. OBrien T. P., McCully M. E. The study of plant structure: principles and selected methods.
Melbourne, 1981. 342 p.
9. . ., . ., . . . . . .: , 2004. 312 .
10. . ., . ., . . Magnoliophyta. .,
2009. 400 .
1)Moscow
State Pedagogical
University, 129243, Moscow,
Kibal`chicha st., 6-5, Department
of Botany
e-mail: avfilonenko@yandex.ru
2)Omsk
e-mail: stratiotes@yandex.ru
In this article we present the results of our study of Comoranthus and Schrebera fruits structure. It is with unified methodology that we first studied fruits structure and the pericarp anatomy of Comoranthus and Schrebera representatives. We demonstrated that mechanical tissues of the pericarp develop only from
the inner zone of the carpel mesophyll. Also, we showed that the
endocarp doesn`t undergo changes and remains monolayered
throughout its development. Based on our data we determined
morphogenetic type of fruits of Comoranthus and Schrebera as
Forsythia-type capsule, which is characterized by mechanical tissues localized only in the inner part of the mesocarp. We considered possible directions of fruits evolution in Oleaceae family and
analyzed phylogenetic relationships of taxa in Oleaceae family with
the help of our carpology data.
Keywords: Comoranthus, Schrebera, Oleaceae, pericarp,
morphogenesis, Forsythia-type capsule.
92
________________________________________________________________
574:581.52 (571.56)
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., 678170,
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e-mail: jeanna_mirny@mail.ru
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______________________________________________________________
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97
______________________________________________________________
, , ,
.
, Chamaenerion angustifolium
. (
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.
.
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. .
,
- ,
, Chamaenerion angustifolium. . , - ,
,
,
.
1. ..
// . : , 1987. . 9-19.
2. .. . -: ,
1995. 225 .
3. .., .. - // . . . .:
- , - "", 1995. . 11. . 166-191
4. .. . , 1989. 146 .
5. .., .. Epilobietea angustifolii Tx. et Prsg. in Tx.
1950 // . . 1 . :
- , 1996. . 62 - 71.
6. .., .. // . VII . . (16-21 2004). , 2004. .2. . 113-120.
7. .., .. Rhodiola iremelica //
: . . . VI .
. (2-6 2002). , 2002. . 76-78.
8. . . (
). : . . . . 2000. 147 .
9. .. //
. . . 3. . .; .: - , 1950. . 6. . 7-204.
10. .. : .., 2002. 308 .
11. .. // . . 1975. 2 . 7-33.
12. ( ) / .., .., .. .. ., , 1976. 217 .
13. ( ) / .., .., .. ...: , 1988.184 .
98
________________________________________________________________
14. .. . : - , 2005.
156 .
15. .., .., .., .., .., .. . : - ,
2005. 575 .
Technical College
in Mirny, 1 Lenin str., Mirny,
678170
e-mail: jenny_myrny@mail.ru
2)Institute
e-mail: cherosov@mail.ru
99
______________________________________________________________
574:581.52 (571.56)
TARAXACUM CERATOPHORUM (LEDEB.) DC
.. 1
.. 2
1)
. .. , 677000,
., . , 58
e-mail: tatyana_everstova@mail.ru
2)
, 677000,
. , . , 41
, , ,
, , - .
: , , ,
e-mail: cherosov@mail.ru
, ,
.
, , , ,
.
... ,
, (, 2009). Taraxacum ceratophorum (Ledeb.) DC. .
Taraxacum ceratophorum .
, . .
. () , . [1]. , [11].
- Taraxacum ceratophorum
, - ,
.
. . , ( 112 450-500.). , [10].
, 100 [2]. . ,
.. 2009-2010. 3-
:
100
________________________________________________________________
1) 10 - ;
2) ;
3) .
,
, .
9 ( ) Taraxacum ceratophorum 2009. 8 2010. 1,2,3 10 . ., 4,5,6,8,9 , 7 .
- (, 1950; , 1975; , 1976; , 1987, 1995;
, 1989; , , 2002, 2004; , 2002).
.. , .. , .. . (2005) (.1).
16
24
15
8
14
16
22
14
20
23
25
13
12 21
11
1
13
18 4
15
17
12
10
19
11
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55
57
59
61
63
65
. 1. ,
:
- 10 . .
- , Taraxacum ceratophorum
- ,
- .
- , Taraxacum ceratophorum
- ,
-
- , Taraxacum ceratophorum
- ,
- Taraxacum ceratophorum
, .
,
. ,
.
,
, .
:
, , , , -
101
______________________________________________________________
, , ,
, .
30
.
.. (1989).
, (, 1950;
, 1975, 1977). ..
(1969), .. (2001).
(, 1995).
.. , .. (2002, 2004) .
- EXCEL STATISTICA.
Taraxacum ceratophorum (.2), , 2010 . . , 2010 . , , () 70% (7). ,
Taraxacum ceratophorum. . ., 2010 . 2 2009., 3,5. 2010 . +36. 9 (53,3%).
2009.
2010.
100%
90%
80%
70%
60%
50%
40%
30%
a
b
c
20%
10%
0%
8
100%
90%
80%
70%
60%
50%
40%
30%
20%
10%
0%
a
b
c
102
________________________________________________________________
), ,
( ). , S-, ,
.
(IVC) .. ,
.. (2002, 2004) 8 (1,21). IVC
9(2009. 0,85;2010. - 0,86),
(54,4) (.1).
. 3. R RS Taraxacum ceratophorum
S- ( , )
2010.
60,00
20,00
7
8
10,00
0,00
60,0
30,00
70,0
40,00
80,0
50,00
90,0
50,0
40,0
30,0
20,0
10,0
0,0
pl
im
g1
g2
g3
ss
pl
im
g1
g2
g3
ss
. 4. Taraxacum ceratophorum
103
______________________________________________________________
5 (2009 . 490,7./; 2010 .
213,3./) 8 (2009.234,7./; 2010. 245,3./).
5 ,
2,24.
.
(I) (I) 2009
. 9(8,0), 2010. 7 (8,84). 9 (2009.33,0./; 2010.35,0./). 9 . , ,
.
(I) (I) 2009.
1 (0,44), 2010 . 2 (0,57).
(1 0,69; 2 0,64)
.
1
1
2
3
4
5
6
7
8
9
,
/
200
9
33,7
65,7
88,7
69,0
490,
7
229,
3
75,7
234,
7
33,0
, I
(g/
v+g),%
(g/p+j+im
+v+g),%
(p+j+im/v
+g), %
1,2
0,6
2,1
-
200
9
0,7
0,6
0,7
0,9
0,6
0,8
0,6
-
20
09
0,7
0,6
0,7
0,5
0,5
0,6
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-
200
9
0,1
0,1
0,1
0,7
201
0
0,4
0,2
0,4
-
(p+j+im/p
+j+im+g+s
s+s),%
200
201
9
0
0,1
0,3
0,1
0,2
0,1
0,4
0,4
-
41,7
224,0
107,7
-
20
09
0,4
0,8
0,5
0,9
201
0
1,2
0,6
2,1
-
200
9
0,4
0,8
0,5
0,9
213,3
6,3
11,3
5,7
11,3
0,5
0,3
0,1
0,1
2,2
0,7
0,7
0,7
101,3
4,6
1,5
4,6
1,5
0,7
0,7
0,2
0,4
2,8
0,4
0,7
0,4
62,3
4,5
8,8
4,5
8,8
0,4
0,3
0,2
0,1
1,1
0,7
0,5
0,7
245,3
2,1
2,4
2,1
2,4
0,5
0,7
0,3
0,3
0,7
0,5
0,4
0,5
35,0
8,0
5,2
8,0
5,2
0,3
0,4
0,1
0,2
1,8
0,5
0,7
0,5
2010
2010
2010
2010
104
________________________________________________________________
1. ..1.: - , 2003. .98-99
2. . : .. ,1989..39
3. .. . . . . . , 2009.40 .
4. .. // . : , 1987. 9-19.
5. . . . -: , 1995. 224 .
6. .. . , 1989. 146 .
7. .., .. - // .
VII . . (16-21 2004). , 2004, .2. . 113-120.
8. .., .. Rhodiola iremelica
// : . . . VI . . (2-6 2002). , 2002. - . 76-78.
9. .., . . , .. , .. , .. , .. .
. , 2005. 108 .
10. .. / .. // . . . 3. . .; .: - , 1950.
. 6. . 7-204.
11. .., .. . :.-,1991. 16-17.
12. : . / . . . . . .: , 1976. 225.
13. .. . : - ,
2005. 156 .
of Ecology
of the M.K. Ammosov North-Eastern
Federal University, 58 Belinsky Str.,
Yakutsk, 677000
e-mail: tatyana_everstova@mail.ru.
2)Institute
-mail: cherosov@mail.ru
The following parameters of coenopopulations of Taraxacum ceratophorum (Ledeb.) DC in plant communities of the
Middle Lena River valley under conditions of anthropogenic impact have been studied: density, vitality structure, trends of ontogenetic strategies, life strategy types, ontogenetic spectra, and
demographic parameters.
Key words: coenopopulaiton, life strategy, weed plants,
ecological condition
105
______________________________________________________________
574:581.52 (571.56)
.
( - )
.. 1
.. 2
.. 3
1)
. , 678170
. , . , 1
e-mail: jeanna_mirny@mail.ru
2)
7 . , 678170
. , . 12/1
3)
,
677000, . , . , 41
,
50- 20 , , .
: - , , .
e-mail: cherosov@mail.ru
,
, ,
, , ,
. ordeum jubatum L.
.
. .
. 30- 20
- .
40-50 ( 90) , , , .
, 0,3 . ,
37 , - , . 7
. 60 . . ,
, . ()
Hordeum jubatum.
106
________________________________________________________________
-,
(, 1950; , 1975; , 1976,
1988; , 1987, 1995; , 1989; , , 2002, 2004).
2009 . . 13 ( ), 2010 . 50 , .. .
:
1. ( , , )
2. ( , )
3. , (, , ..
).
30
( , - . ,
, , 1 , , , 2 ,
, , 3 , - ).
Excel, , .
: (IVC);
(r2), (cv, %).
. . (1950)
: (p), (j), (im), (v), (g1), (g2), (g3), (ss),
(s). (I, I, , ,
, .) . - Excel Statistica.
(IVC) (, , 2004) .
( .. . (2005)) . , 3 :
- Beckmannio-Hordeetum B-H. typicum, .
- Puccinellio-Hordeetum jubati,
, var.
Puccinellia hauptiana var. Hordeum jubatum () .
- Elytrigio-Artemisietum.
. var. Hordeum jubatum var. Chamaenerion angustifolium .
.. .
(2005) . 1.
107
______________________________________________________________
1
-
(-
1 .)
Elytrigio-Artemisietum
var. Hordeum jubatum
Beckmannio-Hordeetum
typicum
Puccinellio-Hordeetum jubati
Hordeum jubatum
Puccinellio-Hordeetum jubati
Puccinellia hauptiana
10
14,2
57,5
20,45
34
14,7
60,5
39,2
14,3
61,1
59,4
43
12,8
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4
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.
.
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.
.
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27,93
16
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22
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.
.
3,96
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.
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: : ( , , ); (. , , .
); (. , . , .
); ( , , ), ( , ,
).
* .. . (2005).
, , , , ,
108
________________________________________________________________
. , .
, .. , ,
. (IVC) . .
3
(.2) ,
. Hordeum jubatum , , .
, 3
,
, .
,
, .
2
\
( )
- . ( .)
- . ( .)
1 ( )
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. 1 ( )
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. 3 ( )
( )
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29,14515
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3,209091
4,699697
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4,761515
0,194636
5,002424
4,281045
0,193281
6,298871
7,150303
43,55455
27,86022
2,655914
3,201075
5,75086
0,206452
4,636129
4,713011
0,220108
4,744366
4,540516
0,216129
5,164473
6,623194
44,82151
29,07
2,679167
3,145833
4,315958
0,199167
4,240833
4,542917
0,26375
4,940833
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:
.
, , , , ,
.
. , . ,
, , ( , , ..).
,
, ,
,
109
______________________________________________________________
. .
:
.
(.1-3).
, , , , .. ,
(.1). .
. 1-3.
( -). ( , .2-3) -
(R-), . ,
, .
Scatterplot (Spreadsheet1 10v *50c)
Var2 = Distance W eighted Least Squares
48
0,18
0,16
36 11
0,14
16
0,12
4
0,10
R2
37
2
0,08
0,06
19
44
0,04
13 40
15
31
55047
28 32
22
49
43
30
6
9 45
24
27
41
46
8
42
33
710
2923
1439
34 18
17
38
26
13
12
21
20
35
25
0,02
0,00
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0,6
0,7
0,8
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1,1
1,2
1,3
IVC
. 1. Hordeum jubatum
.
, ,
,
.
( ) , .
, , .
110
________________________________________________________________
. 2.
. 3. ,
111
______________________________________________________________
1. .. 1987 // . : , . 9-19.
2. .. 1995 . -:
, 225 .
3. .. 1989 . ,
146 .
4. .., .. 2002 Rhodiola iremelica // : . . . VI
. . (2-6 2002). , . 76-78.
5. .., .. 2004 // . VII . . (16-21 2004). ,
2004. .2. . 113-120.
6. .., .., .. . 2005
. : - , 108 .
7. .. 1950
/ .. // . . . 3. . .; .: - , .
6. . 7-204.
8. .. 1975
// . , 2 . 7-33.
9. ( ) 1976/ .., .., .. .. ., , 217 .
10. ( ) 1988 /.. ,
.. , .. . .: , 184 .
11. .., .., .. ., 2005 . : - , 575 .
Technical College
in Mirny, 1 Lenin str., Mirny,
678170
e-mail: jenny_myrny@mail.ru
2)Institute
e-mail: cherosov@mail.ru
For the first time the results of complex study of coenopopulations of Hordeum jubatum L. are presented.
H. jubatum is an invasive species which has penetrated the territory of diamond mining activity in 1950s and has become one of
most active species of the vegetation cover of anthropogenically
impacted habitats. Its life strategy and morphological parameters
under various ecological conditions have been revealed.
Key words: Population-biological parameters, coenopopulaiton, invasive species.
112
________________________________________________________________
574.3:574.22 (571.56)
PYROLA INCARNAT (DC.) FREYN
- ( )
.. 1
.. 2
1)
- . .. , 677000,
. , . 48,
-mail: aanikif@yandex.ru
, 67700, . ,
. 41
2)
Pyrola incarnat . .
: Pyrola incarnat, ,
-mail: cherosov@mail.ru
(Pyrola incarnat) - .
(1 1 ),
3 , .
-, [6; 9; 10]. Pyrola incarnat - [2]. .. [6], .. [7].
, .. [1], .
(Pyrolaceae).
. , ,
[10]. . , .. 2 , , [1]. . Vaccinio-Laricion cajanderi Pestrjakov et al. 1992. Vaccinio-Laricetum cajanderi Gogl. in Pestrjakov et al.
1992, -
[3].
2009 2010 (). 12 (),
: 2009 5 (1-5), 2010 5 7 (6-12).
, - .
, , ,
113
______________________________________________________________
:
.
4
:1) (2,11) 2) 1,4,5,6,8 3) 3 4) 7,9,10,12
2 11 ,
, . , 2
(, , ..), 11 , 90-
.
6 . 4, 5 8
, - (4
2010 .; 5 8 2000 .).
3
, 3 2010
.
, . 10 , , 80- .
7 9 -
Vaccinium vitis-idaea, Pyrola incarnata, Linneae borealis
Arctous arctostofilos. 12 , .
, , ,
. 2, 12 11, 100 1 2. 8, 359 /2,
184 1 2.
0,87 2,50. ( 6, 7 9),
1. 1 (2,5) (.1).
.
, 6, 7, 9 11, ,
, .
, . 4 2010
, 2 11 .
, - 3 8 , 2.
, .
0,950,35 . 1-2 4,73 1,14.
2,54 0,52 , 2,210,56 . .
1,28 0,36 . 3-4 , 5,541,23 .
2,56,0,54 , 2,350,52 .
114
________________________________________________________________
1
I.
1
2
3
4
5
6
7
8
9
10
11
12
2,73
1,33
1,27
1,59
1,28
1,19
1,06
1,38
1,11
1,20
1,36
1,55
I.
2,50
1,26
1,24
1,59
1,20
0,87
0,90
1,29
0,94
1,19
1,21
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183,75
91,25
252,50
115,50
258,00
212,33
246,00
359,50
265,50
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79,50
49,60
8,70
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45,40
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149,70
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27,35
,
%
88,84
1,36
9,80
74,59
0,55
24,32
74,85
2,38
22,77
92,64
4,33
3,03
71,71
14,74
13,55
56,67
4,08
39,25
56,10
10,37
33,54
76,36
4,03
19,61
53,86
2,26
35,78
72,92
17,19
9,90
62,37
2,43
34,83
77,99
13,84
8,18
.
, 2,521,23 . 5-10 .
6,581,33 . 3,180,76 , 2,710,62 . . .
. 22,341,94 .
3-7 , 6,431,42 , 3,350,72 ,
2,410,98 . 5,721,73 , 6-11 . . .
. , 25,742,65 . 2 10 , 7,440,98 .
3,720,75 , 3,080,76 . 9,353,46 ,
- 7-16 .
, 19,453,26 .
, 1
5 , 6,251,64 . 3,360,75 ,
2,840,97 . 5,781,76 , 5-14 .
. .
.
3,40,86 . 1-5. , 5,340,38 , 2,680,53 , 2,340,47 . , , .
.
- 4,281,55 . 1-5.
: 4,721,76 ,
2,521,14 , 1,881,13 . , .
,
(g2). , . , ,
.
115
______________________________________________________________
2
Pyrola incarnat
()
(
Im
g1
g2
g3
Ss
0,95
0,35
1,280,36
2,521,23
22,341,94
24,742,65
19,453,26
3,40,86
4,281,55
1-2
3-4
5-10
3-7
2-8
1-5
1-5
1-5
2,19
0,64
2,880,71
3,40,67
3,080,7
3,620,98
2,890,89
2,660,38
2,20,62
2,54
0,52
2,560,54
3,280,76
3,250,72
3,720,75
3,360,75
2,680,53
2,521,14
2,21
0,56
2,350,52
2,610,62
2,410,98
3,080,76
2,84-0,97
2,340,47
1,881,13
5,721,73
9,353,46
5,781,76
6-11
7-16
5-14
12 Pyrola incarnat,
, , , .
, ,
2 , ,
3 , , . , 5
, . ,
. , ,
.
. , , ,
. , ,
, 8-10%. [5].
5 8, , , (9 11), . 3 4,
2010
, , 4,
. 2009
3 68%, - 7,08% - 25,34%, 2010 75%; 2,61% 23,26% , ..
. .
2009 4 78%, - 2,61%
- 19,58%, 2010 : 93%;, 1,33% 6,03,
.
198 /2 115,5 /2.
. , .
116
________________________________________________________________
, , . ,
. , . .
.
.
.
1. .. . -: , 1995. -224 .
2. .. _
//. . . . 1978. . 83. . 3. . 77-88.
3. ..
. . //. . -. : , . , 23-24 2006 . .
. . .66-71
4. .. Pyrola incarnat Fisch. - (
). // : . - XI . .-. . . 20-25 2010 .. . : , 2010, -231 . - . 212-213 .
5. .., .., .. . // . .
-. : , .
, 23-24 2006 .. .1. . .
: - , 2006. . 76-83.
6. .. // . .3. . ., 1950. . 7-204.
7. .. //. . 1975. 2. . 7-34.
8. , 11, Pyrolaceae - Lamiaceae (Labiatae), , 1997, 296.
9. ( ). .: , 1976. 214.
10. ( ) / .. ,
.. , .. , .. . - .: , 1988. - 236 .
-mail: aanikif@yandex.ru
Institute for Biological Problems of
Cryolithozone Siberian Branch of RAS.
677980, Yakutsk, 41, Lenina ave.
2)
-mail: cherosov@mail.ru
. 2011. 3 (98). 14
117
______________________________________________________________
528.9:581.9 (571.56)
.. 1
.. 2
.. 2
..2
1)-
, 677000,
. ,
. , 58,
E-mail: katammsva@mail.ru
2)
,
677000, . ,
. , 41
E-mail: cherosov@mail.ru
enikolin@yandex.ru
etroeva@mail.ru
, ,
Landsat 7
(1989) , ,
. , ..
,
, .
: , , , .
, , . (, , , , .) ,
, , .
,
.
, .
. 20%, 70% . . .. (1956), .. (1963 , ), .. (1963), .. , ..
(1980), .. , .. , .. (1991) (
. , 1989) . -- .
- , (Betula exilis) (Salix glauca, S. reptans) (Vaccinium vitis-idaea, V. uliginosum).
, ,
, .
, ,
-
118
________________________________________________________________
[3]. .
170 180 ,
374 , - . . . , -
. . . ,
. . ( ..). -- Alopecurus alpinus
Salix polaris [1].
1:5 000 000
(1989).
( ) ( ).
, .
ArcView 3.2,
Landsat 7/ETM+.
, ,
(1987). ( , ):
1.
1.1
1.1.1 -
1.1.2 -
1.1.3 (Alopecurus alpinus) (Ditrichum
flexicaule, Districhum capillaceum, Orthothecium chryseum) -
.
1.2
1.2.1 -
1.2.2 -
1.3
1.3.1 [2]
, -
. , , .
.
, -
. 2011. 3 (98). 14
119
______________________________________________________________
. (1989) . , , 1-2 . 1-2.
,
.
. , , ,
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. ,
- -
. .
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1
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1
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1
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2
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2
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2
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2
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3
58,8
3
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3
7,6
3
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250,00
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. 1.
( . 1, . 2):
. ; . ; . ; .
120
________________________________________________________________
.
,
(.3). ,
, , 10,6% 15,4%.
, , . 100 .
2
1
116,0
1
109,0
1
102,1
2
137,4
2
165,7
2
164,1
3
99,4
3
62,4
3
70,8
36,0
22,9
51,6
43,9
49,1
59,4
52,0
22,0
47,5
250
200
150
100
50
0
1
.2.
( . 2, . 3):
. ; . ;
. .
: - - , . , 4,7%,
.
. 2011. 3 (98). 14
121
______________________________________________________________
3
(1989) ( %)
/
-
-
(Alopecurus alpinus) (Ditrichum flexcaule, Districhum
capillaceum, Orthothecium chryseum) -
- (Alopecurus alpinus Salix polaris)
(Aulacomnium turgidum Hylocomium splendens var. alaskanum Orthothecium
chryseum)
- (Alopecurus alpinus Salix polaris)
(Aulacomnium turgidum Hylocomium splendens var. alaskanum Orthothecium
chryseum)
- (Alopecurus alpinus Saxifraga oppositifolia Salix
polaris) (Ditrichum flexicaule Distichium capillaceum) -
- (Rhizocarpon geographicum Haematomma ventosum
Umbilicaria)
(Deschampsia
brevifolia Poa alpigena)
18,0
0,0
0,5
17,7
24
32,8
48,7
14,3
8,7
1,1
1,9
16,1
16,2
,
.
, (- , , , , , , ,
).
. 3
. 3.
(, 1:5 000 000)
122
________________________________________________________________
(.4) (.4).
. 4.
(, 1:5 000 000)
:
-
-
- -
-
1. .., .., .., .. . : . 1987. 154 .
2. . 1989. .: , 115 .
3. .., .., ..
. : - , 1991.192 .
. 2011. 3 (98). 14
123
______________________________________________________________
USE OF GIS-TECHNOLOGIES FOR CORRECTION OF CONTOURS
OF SMALL-SCALE MAP OF VEGETATION
E.V. Ammosova1
M.M. Cherosov2
E.G.Nikolin2
E.I.Troeva2
1)M.K.
Ammosov North-Eastern
Federal University,
58 Belinsky Str., Yakutsk, 677000
e-mail: katammsva@mail.ru
2)Institute
for Biological
Problems of Cryolithozone SB RAS,
41 Lenin Ave., Yakutsk, 677000
e-mail: cherosov@mail.ru
enikolin@yandex.ru
etroeva@mail.ru
124
________________________________________________________________
581.9 (571.6)
..
..
,
677980, . , . , 41
e-mail: enikolin@yandex.ru
cherosov@mail.ru
, , .
, , - ,
.
:
, .
() ,
, - .
. ,
.. (1959, 1968), .. . (1966), .. (2006),
.. , .. (2010) .
(, 1991, 2009 , .).
, 900 ,
78 276 . 4 - : (), (),
() ().
-,
.
Statistica 6.0.
.
Statistica 6.0
(.1), (, )
( 21,5) ().
,
, (
14,5), .. .
. (690 , 77%
) (.1). , ,
. , , ,
. 2011. 3 (98). 14
125
______________________________________________________________
(542 , 60%
). - 254 (28 % ), 125 ,
. . , , 2.
. ,
, . - . ,
.
Tree Diagram for Variables
Ward`s method
Euclidean distances
22
21
20
Linkage Distance
19
18
17
16
15
14
13
. 1. -
1
( )
69(89)
63(81)
203(74)
171(62)
557(62)
452(50)
71(91)
238(86)
690(77)
61(78)
189(69)
542(60)
: 3, (%) .
2
-
( )
4
31
135
2
2
7
3
29
13
2
19
117
126
________________________________________________________________
:
, , . , . , , , .
.
, .
, ,
.
. , ,
, .
(),
.
. , .
,
. ,
, , , , , , .
,
.
, , ,
. ,
, .
.
- .
, 5
(), - (), (), () ().
() , , , , , , , - , , , .
- () , ,
, , , , - .
() , , , ,
, - .
- ()
Thuidium, Racomitrium,
. 2011. 3 (98). 14
127
______________________________________________________________
Schistidium ., . .
()
, .
, , , , ,
, , , , , , .
. 3 4 .
(.2). , 27, . 21,
: 1) 2)
.
Tree Diagram for Variables
Ward`s method
Euclidean distances
28
26
Linkage Distance
24
22
20
18
16
14
. 2.
3. , .
. , , .
, , -, - . , , , 159 (97
), 15 . ,
, 4. , .
, .
.
128
________________________________________________________________
3
72(92)
257(93)
749(83)
63(81)
49(63)
50(64)
186(67) 122(44) 147(53)
485(54) 247(27) 451(50)
13(17)
34(12)
53(6)
12
67
116
1
0
1
0
4
0
1
0
17
2
29
0
- .
5. 19 20
( ).
Tree Diagram for Variables
Ward`s method
Euclidean distances
45
40
35
Linkage Distance
30
25
20
15
10
5
0
. 3.
- .
2 ,
(. ) ()
. 2011. 3 (98). 14
129
______________________________________________________________
3, 6 .
, 43,
, . .
, 4, .
34,
(, ). , ,
.
26,5 ,
16,5. , , .
22 3
. ,
.
18. , 4 , .
.
11. 11,5 , .
. -
( 10). (12),
.
, .
.
-
. - . ,
, ,
- ,
- .
5 .
5
-
523
441
315
439
396
197
233
298
182
247
262
73
179
143
0
369
22
27
7
30
130
________________________________________________________________
1. .. . .,
2006. 568 .
2. .., .. -
( ) // . ., 2010. - .95, 10. - . 1396-1421.
3. .. . . .
. . . , 1991. 14 .
4. .. - // . ., 2009. . 235-239.
5. .. // : . . . ., . .. . , 2009. . 250-255.
6. .., .., .. ( ) // . . 6. .-.: , 1966. . 7 39.
7. .. . // . . 1959. . 44. 8. . 1171-1189.
8. .. -. ., 1968. 236 .
For the first time the study of the Verkhoyansk Ranges regional floras has been conducted with the use of methods of mathematical statistics, particularly, the cluster analysis. The analysis of similarity
and difference of large regional floras has been made at the level of floristic regions, longitude-latitude regions, partial floras of altitudinal
belts and valley complexes.
Key words: Verkhoyansk Range, longitude-latitude regions,
partial flora.
. 2011. 3 (98). 14
131
______________________________________________________________
502.9:581.55 (571.56)
..1
..2
.. 3
2)
( )
W.Kunick (1982), S.Klotz (1984)
, .
.
e-mail: cherosov@mail.ru
: , , ,
1)-
, 677000, . ,
. , 58, ,
e-mail: pbnbot@mail.ru
, 677000,
. , . , 41
3)
,
450074, . , . , 32
e-mail: ishbirdin@mail.ru
, .
. , (
), , .. , , , , , , , .
, , .
, (
W. Kunick (1982), S.Klotz (1984) .), , .. (1988, 1994) , .. (1992) .
. .
H.Sukopp (, ,
) ,
( , ,
.). , .
. , , .
132
________________________________________________________________
.
.
, .
, ( ). , .
7
W.Kunick (1982), S.Klotz (1984). 7
(Frank, Klotz, 1990):
, ;
, , ..,
;
m , , , ;
b - , , , ;
- , ,
;
, ;
t .
. , .. .
(2005), .. . (2005) . 1835
1969 ( ., 2005)
, (Kunick, 1982; Klotz, 1984),
.
( ., 2005).
.. (1988, 1994) ,
.
- IBIS
( .. , )
(. 1). (4 , , -, ) , - . ,
(, , ). ( - , - - ),
, , .
,
(37,8 49,1%), (20,7 29,9%), 3 (15,3 16,3 %).
-, - ( 3,5-4,3;
2,5 4,0; 2,5 3,5%), .
. 2011. 3 (98). 14
133
______________________________________________________________
(37,3 42,9%), (16,8 25,3 %),
(15,6 17,6%). - (4,8 5,5; 4,9 7,8; 4,8 6,5 %).
.
:
16,8% () 49,1% ().
:
17,6% () 15,3% () .
, ,
, 42,9 % ()
20,7 % ().
- ,
(4,8 5,5 % 3,5 4,4 %
). - ( 7,8 % 2,5 % ).
6,5 2,5 %, 0,8 0,0%.
,
, ,
.
, , , . , , ,
.
1
( %)
a-
o-
m-
b--
--
p-
t-
1
2
3
4
5
6
7
8
16,8
17,5
42,9
5,3
6,7
5,7
0,8
4,3
25,3
17,6
37,3
4,8
4,9
4,8
0,4
4,9
17,1
15,6
42,8
5,5
7,8
6,5
0,7
4,0
37,8
15,8
29,3
3,6
4,0
3,5
0,5
5,5
39,5
16,3
29,9
4,4
3,2
2,5
0,1
4,1
41,9
15,3
29,3
4,0
3,4
2,7
0,0
3,4
49,1
15,4
20,7
3,5
2,5
2,5
0,3
6,0
29,8
17,0
33,5
3,6
5,0
4,4
0,7
6,0
134
________________________________________________________________
2 (
), .
(3- ), - -,
, , - .
, .
. 2.
2
(
, )
Scheuchzerio-Caricetea nigrae
Alectorio-Dryadetum octopetalae
Senecio-Poetum alpigenae
Saxifrago-Pinetum
Arctophiletea fulvae
Tripleurospermo-Poetum alpigenae
Ledo-Laricetum calanderii
Festuco-Laricetum cajanderii
Epilobietea
Calamagrostetea
Phragmiti-Magnocaricetea
Pulsatillion
Alopecurion
Festucion lenensis
Hordeion
Elytrigio-Glaucetum
Bidentetea
Potentillion anserinae
Artemisio-Caricion
Amorio-Artemisietum
Puccinellion
Beckmannio-Hordeetum jubati
Puccinellio-Salicornietum
Suaedo-Puccinellietum tenuiflorae
Stellarietea
Puccinellio-Hordetum jubati
Elytrigio-Artemisietum
Matricario-Polygonion
.. Chenopodium album
a
a
o
m
m
m
m
m
m
m
m
m
m
b
b
b
b
c
c
c
c
c
c
c
p
p
p
p
p
. 2011. 3 (98). 14
135
______________________________________________________________
,
.
, ,
,
.
1. .., .., .. . 2005
. : , 320 .
2. .. 1988 // : .
. . . , .25-36.
3. .. 1994 //
. , 1988 . 3- .
. , .269-276.
4. .. 1992 - 60-80 : . . . . . ,
16 .
5. .., .., .. 2003
// , .7, 7. .33-36.
6. .. .., .., .., .., .. 2005 . :
- , 575 .
7. Frank D., Klotz S. 1990 Biologisch-oekologische Daten zur Flora DDR. Martin-LutherUniversitaet, Halle Wittenberg,
8. Klotz S. 1984 Die ruderalgesselschaften eines neubaugebietes ihre verbeitung und kombination // Acta bot. Slov. Acad. Sci. Slovacae. Ser.A. .1. S. 111-125.
9. Kunick D. 1982 Zonietung des Stadtgebietes von Berlin (West). Ergebnisse Floristischer
Untersuchung // Gen. Schriftenr. d.Fachber. Landschaftsentwicklung u.Umweltforsch. 14. S. 1-164.
2)Institute
For the first time the results of previous studies have been
summarized on plant hemeroby (level of naturalness) according
to W.Kunick (1982) and S.Klotz (1984) both at the levels of syntaxa
and floristic regions. Characteristics and some common principles
within the natural zones of Yakutia are given.
e-mail: cherosov@mail.ru
1)North-Eastern
Federal University,
58 Belinsky Str., Yakutsk, 677000
e-mail: pbnbot@mail.ru
for Biological Problems
of Cryolithozone SB RAS,
41 Lenin Ave., Yakutsk, 677000
3)Faculty
e-mail: ishbirdin@mail.ru
136
________________________________________________________________
582.572.4:581.14:58.006
AMARYLLIDACEAE JAUME ST.-HIL.
..
..
..
, 308007,
. , . , 85
e-mail: Arkhipova@bsu.edu.ru
sorokopudova@bsu.edu.ru
Amarillidaceae
. 2 4 .
-
Clivia miniata, Crinum moorei Hymenocallis speciosa.
: , Amarillidaceae,
, .
. 2011. 3 (98). 14
137
______________________________________________________________
1:2:1. 2000- 2500 lx, 1500 lx, 25-30 ( 37) 0 15-17 ( 1017) 0 . , .
[7], .. [8].
.
. .. (. -) GRIN
Taxonomy for Plants [5-6, 9].
marillidaceae (. 1).
1
Amarillidaceae -
,
Clivia miniata
Crinum moorei
Haemanthus albiflos
Cyrtanthus elatus
Eucharis grandiflora
Hymenocallis speciosa
Zephyranthes candida
Zephyranthes carinata
White Giant
Ludwigs Goliath
Baby Star
Fire Dance
andy Cane
06.05.08
25.06.08
51
10.03.09
20.06.09
103
29.04.10
05.07.10
68
05.06.08
15.07.08
41
19.06.09
30.07.09
42
21.06.10
20.08.10
61
11.06.08
16.07.08
36
02.10.08
20.10.08
19
05.09.09
25.09.09
20
10.08.10
26.08.10
16
03.12.07
21.12.07
19
25.05.08
20.06.08
26
26.11.08
15.01.09
51
10.06.09
29.06.09
20
30.11.09
16.12.09
17
18.06.10
05.07.10
18
08.11.10
27.12.10
50
27.06.08
10.08.08
45
10.05.09
05.08.09
88
02.07.10
16.08.10
46
11.08.08
10.09.08
31
28.07.09
05.09.09
40
19.06.10
23.07.10
35
10.08.08
20.08.08
11
05.08.09
27.08.09
23
15.08.10
06.09.10
23
Hyppeastrum hybrida
16.12.06
20.01.07
36
02.01.08
03.02.08
33
25.01.09
04.02.09
11
15.01.10
22.01.10
8
22.01.08
10.02.08
20
26.01.09
28.02.09
34
08.02.10
10.03.10
31
24.03.08
10.04.08
18
09.03.09
20.03.09
12
14.02.10
05.03.10
20
05.04.07
22.04.07
18
26.03.08
14.04.08
19
9-11
8-17
3-4
3-9
30-40
3-4
3-4
5-6
4-6
4-26
4-5
4-6
2-4
4-5
2-4
138
________________________________________________________________
, . (40 ) Cyrtanthus elatus, Haemanthus albiflos, Zephyranthes candida,
Z. carinata Hyppeastrum.
Clivia miniata, Crinum moorei, Hymenocallis speciosa. Eucharis
grandiflora . Hyppeastrum
3 ,
. , (Crinum moorei, Cyrtanthus elatus,
Hymenocallis speciosa), 3-4 ,
4-6 .
Clivia miniata 9-11 .
2008-2010 . 2 :
- Eucharis grandiflora (I) (
)
(II). --, -, - -- (. 2).
2
Amarillidaceae
II
-
-
Clivia miniata
+
Crinum moorei
Cyrtanthus elatus
Haemanthus albiflos
Eucharis x grandiflora
+
+
Hymenocallis speciosa
Zephyranthes candida
Zephyranthes carinata
Hyppeastrum hybrida
+
I
+
+
+
-
+
+
-
+
+
-
: I , ; II ,
2 ; - , - ; - -
; ; - , - .
. 2011. 3 (98). 14
139
______________________________________________________________
,
marillidaceae .
. Eucharis grandiflora;
Hyppeastrum - . -- Clivia
miniata, Crinum moorei, Hymenocallis speciosa, Haemanthus albiflos, Cyrtanthus elatus,
Zephyranthes candida, Zephyranthes carinata.
20-40
Cyrtanthus elatus, Zephyranthes candida Eucharis grandiflora,
.
, ,
, .
40 - - Clivia miniata, Crinum
moorei Hymenocallis speciosa.
1. . . . - .: -, 2001. - 128 .
2. .., .. . : , 2001. 416 c.
3. . , . :
/ . . .. . .: , 2003. - 335 .
4. .. , . - : , 1995. - 304 .
5. .. . , .
.: , 1970. 179 .
6. .. . .: , 1983. 621 .
7. // . . .
. 1979. . 113. . 3-8.
8. .. . . . . . ,
2008. 16 .
9. Germplasm Resources Information Network (GRIN): Family: Amaryllidaceae.
: http://www.ars-grin.gov/cgi-bin/npgs/html/family.pl?2561.
Terms of flowering of some representatives of family Amarillidaceae in the conditions of a greenhouse of the Winter garden of the Belgorod state university are studied. 2 groups and 4 subgroups on seasonal
rhythms of flowering are established. Spring and summer-flowering species Clivia miniata, Crinum moorei and Hymenocallis speciosa are characterized by stable long-flowering in different years.
Keywords: rhythms of flowering, Amarillidaceae, introduction,
greenhouse.
140
________________________________________________________________
635.925
CAMPANULA L.
..
..
, 450080
, . , 195, . 3
e-mail: flowers-ufa@yandex.ru
11
Campanula L. . 3 : , , 7 . . 8 .
. , , , ,
, .
, . , [3, 8].
.. [8]
. .. [4]
.. [7].
..
. .. .. . ,
, - [1].
11 Campanula L. - .
( ).
. ,
,
, . , . 2009
, - 2008.
2 (C. thyrsoides L., C. sibirica L.) 9 .
. 2011. 3 (98). 14
141
______________________________________________________________
- C. alliariifolia Willd. (2003 ), C. punctata Lam. (2006); C. carpatica Jacg. (2000),
C. thyrsoides L. (2007). : C. glomerata L. (2006); C. latifolia L. (2006); C. persicifolia L.; C. rapunculoides L. (2006), C. sibirica L. (2007); C. rotundifolia L.
(2006); C. trachelium L. (2005) .
.. [4] .. [7]. .. [5, 6] .. .. [2].
(2008-2009 .)
: , (, , ) .
. . , , . .
(pl). 15-20- . . , , ,
; ,
. , .
9-12- .
(j).
2-4 .
III . C. carpatica C. rotundifolia .
(im)
, 5-8 , , ( C. latifolia, C. sibirica C. thyrsoides,
). C. carpatica C. rotundifolia .
IV , .
(v)
( C. latifolia, C. sibirica, C. thyrsoides) ,
; C. carpatica C. rotundifolia -
. C. alliariifolia, C. glomerata, C. sibirica C. thyrsoides
, - . ,
, .
68-74 (C. arpatica
C. rotundifolia) 414-442 .
. . sibirica . thyrsoides
III . (g1) . . sibirica 532, . thyrsoides - 261 .
142
________________________________________________________________
(g2) ,
(. thyrsoides) (. sibirica)
;
.
. sibirica 141, . thyrsoides
- 211 .
(g3) . thyrsoides , . sibirica .
, , .
362 .
. thyrsoides ; . sibiric . ,
. sibiric . thyrsoides . (. 1. ).
C. arpatica C. rotundifolia (100% ).
(g1). II .
(g2) .
, (
) . C. rotundifolia
, (. 1. ).
. . alliariifolia, . glomerata, C. latifolia, . persicifolia, . punctata,
. rapunculoides, . trachelium
(g1) (. 1. ).
. : 1. (C. sibirica, C. thyrsoides); 2. -- (C. persicifolia); 3.
--
(C. rapunculoides); 4. --
(C. trachelium, C. latifolia, . arpatica);
5. -
(C. alliariifolia); 6.
-
(C. punctata); 7.
-- (C. rotundifolia); 8. -
(C. glomerata).
. 2011. 3 (98). 14
143
______________________________________________________________
pl
im
g1
g2
g3
pl
im
g1
g2
pl
im
g1
.
,
- .
144
________________________________________________________________
2. , : ;
2-4 , ; , ;
, . C. arpatica C. rotundifolia,
.
1. .., .., .. . . . .: , - , 2001. 640 .
2. .., ..
. - : , 2006. - 296 .
3. : . / . .. -: , 2007. . V. 372 .
4. .. //
. 1950. .1. . 465-483.
5. .. // . ., .: , 1964. . 3. . 146-202.
6. .. . ., 1952.
390 .
7. .. // . . 1975. 2. . 7-34.
8. .. ( ) // . .: , 1967. . 3-8.
. 2011. 3 (98). 14
145
______________________________________________________________
630*17:582 (470.44)
SORBUS L. ARONIA L.
..
- , 410010,
. , . , 7
e-mail: arestova.elena@mai.ru
10 2 Sorbus 2 Aronia
.
: , Sorbus, Aronia, ,
,
,
[1].
, .
: , ,
.
.
[2, 4]. +50 ( ),
.
, . , ,
. . , . , ,
[1].
10 2 Sorbus L.
2 Aronia L. .
[3, 4] . 12 .
.
( ).
146
________________________________________________________________
, .
.
.
70 % .
.
.
, .
.
.
, , , .
,
, .
+50, . .
,
. (. 1).
,
, (. 2).
1
, %
2,0
98,0
4,3
95,7
8,7
91,3
54,9
55,1
, %
4
8
11
19
13,1 24,3
11,1 19,1
7,6 14,2
1,1 17,7
41
3,6 25,9
30,6
69,4
52
6
5
12
12
35
32
3,7 8,1
9,3 15,8
4,3 7,7
3,5 8,5
2,4 8,3
2,0 19,7
3,0 19,0
71,3
4,1
7,5
12,5
16,1
80,9
33,9
28,7
95,9
92,5
87,5
83,9
19,1
66,1
. 2011. 3 (98). 14
147
______________________________________________________________
2
25.4
147
2.5
249
15.6
971
16.8
2479
3.9
2721
3.5
257
17.5
457
25.5
594
21.4
106
26.4
157
3.5
260
15.6
970
24.8
2656
26.9
3060
9.5
356
19.5
484
25.5
594
S. aucuparia Aronia
melanocarpa
19.4
89
22.4
123
29.4
203
22.6
1364
19.8
2520
10.9
2827
5.5
291
15.5
423
23.5
554
S. mericana
20.4
98
23.4
131
1.5
214
14.6
976
10.9
2849
4.10
3141
4.5
269
20.5
498
4.6
776
S. amurensis
19.4
89
22.4
123
30.4
228
12.6
957
29.7
2491
3.9
2732
3.5
260
16.5
439
24.5
565
S. comixta
19.4
88
23.4
131
2.5
218
14.6
921
18.8
2186
7.9
2781
4.5
269
17.5
457
24.5
564
S. hibrida
23.4
121
26.4
164
4.5
269
11.6
935
30.8
2743
3.10
3133
7.5
312
17.5
460
24.5
565
S. discolor
19.4
88
23.4
126
1.5
222
14.6
964
19.8
2520
7.9
2790
3.5
257
18.5
474
24.5
565
S. domestica
25.4
147
30.4
228
10.5
362
13.6
968
7.9
2866
24.10
3272
7.5
312
16.5
439
24.5
565
S. intermedia
23.4
131
28.4
187
5.5
291
13.6
968
22.8
2602
24.9
3055
6.5
304
17.5
460
25.5
608
S. Mougeottii
23.4
125
29.4
198
7.5
312
13.6
954
19.8
2635
25.9
3040
4.5
269
20.5
489
26.5
613
S. sibirica
21.4
106
27.4
175
4.5
269
13.6
936
5.8
2299
13.9
2877
5.5
280
20.5
489
27.5
623
19.4
89
19.4
89
23.4
131
24.4
141
29.4
203
1.5
240
29.6
1309
30.6
1329
17.8
2495
3.8
2259
9.9
2813
7.9
2790
6.5
304
5.5
286
17.5
464
17.5
464
23.5
544
23.5
554
ronia prunifolia
A. melanocarpa
20.4
96
Sorbus aucuparia
Sorbus L. Aronia L. *
29.5
658
28.5
643
26.5
614
7.6
841
28.
5
643
27.5
628
31.5
703
28.5
643
28.5
643
30.5
672
30.5
671
30.5
674
26.5
615
26.5
615
9.7
1541
11.8
2309
13.7
1693
6.8
2190
4.8
2269
5.9
2765
14.8
2362
5.9
2765
20.7
1482
13.8
2331
1.8
1845
11.8
2294
19.7
1799
11.8
2312
1.8
2327
13.8
1542
13.7
1665
23.7
1898
22.7
1889
22.8
2402
30.8
2743
21.8
2681
30.8
2743
22.8
2602
5.9
2765
13.8
2395
13.8
2395
10.8
2373
* , .
2,0 80,9 %. ,
(2,0 7,5 %). 4 8 .
12,5 16,1 %. 12 .
, (80,9 71,3 %).
.
43 ( 132 174 )
(. 3). , .
148
________________________________________________________________
3
Sorbus Aronia
,
Sorbus ucuparia
S. aucuparia Crataegus sanguinea
S. aucuparia Aronia melanocarpa
S. americana
S. amurensis
S. comixta
S. discolor
S. domestica
S. intermedia
S. hibrida
S. Mougeottii
S. sibirica
ronia prunifolia
A. melanocarpa
131,5 9,81
2574 125
153,0 7,99
2904 64
140,5 6,59
2704 102
164,0 5,66
3010 36
134,5 9,92
2611 121
137,0 9,57
2650 142
136,8 9,07
2751 128
174,510,11
3044 57
149,5 8,03
2853 69
160,2 5,78
2970 64
149,5 8,54
2856 78
138,5 7,62
2702 142
138,5 7,35
2681 112
137,5 7,60
2642 117
8,0 2,27
6,5 1,50
7,8 1,75
15,5 2,22
7,5 1,85
6,8 1,31
6,0 1,47
8,0 1,24
8,0 1,96
6,5 1,85
6,5 2,40
7,5 1,04
5,8 1,25
6,2 1,25
134 16
110 12
121 13
278 16
129 13
116 12
102 12
136 14
136 14
105 15
111 16
134 12
96 13
106 15
. ,
. 13 6-8 , , ,
.
,
, . .
5 : , , , , .
. , , , , , , 90 100 %, , , 85 90 %, 65 70 %.
.
1. .
2. 2 :
:
2
7,5 %, 71,3 80,9 %;
19,1 98 %.
3. :
88 147;
423 498;
96 278;
2574 3044.
4. 3
, 2 :
. 2011. 3 (98). 14
149
______________________________________________________________
1 (131144 ) Sorbus ucuparia, S. amurensis, S. comixta,
S. discolor, S. aucuparia Aronia melanocarpa, S. sibirica, ronia prunifolia,
A. melanocarpa;
2 (145157 ) S. hibrida, S. intermedia, S. Mougeottii,
S. aucuparia Crataegus sanguinea;
3 (158174 ) S.mericana, S. domestica.
1. .. -
Sorbus L. Aronia L. ( ). . . . ... -: , 2000. 24
2. . . . .: , 1981. 118 .
3. . ., 1975. 28 .
4. ..
// .
., 1972. . 40 46.
150
________________________________________________________________
582.998.2:581.14
CHRYSANTHEMUM L.
.. 1
.. 1
.. 2
.. 2
1)
Chrysanthemum L.
. Chrysanthemum.
Chrysanthemum ,
.
, 308015, . ,
. , 85
e-mail: stetsovich.a@yandex.ru
. .. , 61022,
. , . , 52
2)
: , Chrysanthemum,
, .
e-mail: khbg@i.ua
Chrysanthemum zawadskii Herbich., .
(Chrysanthemum hortorum Bailey) .
- (. ), (. ),
- (. ), . .. , . .. (. ),
(. ), (. ),
(. ),
. .. (. ),
[1-7]. 2008 .
() (. ).
,
. 14,5 ,
- 13,5 ,
8-10 [8]. - , [1].
,
-
(- - )
.
2008-2010 . . .. . 80 , . -
. 2011. 3 (98). 14
151
______________________________________________________________
- 5035 c.., 3635 .., . - 4959 c.., 3613 .. -
. , . [9,10]. 2009
2010 . - - , . , 2010
., (.1).
1
( . )
,
2009 . 2010 .
8,3
9,8
7
14,4
17,5
15
20,9
22,3
18,8
21,4
25,3
20,3
18
25,5
19,3
16,1
14,7
13,5
8,5
4,8
6,5
3,4
6,9
-0,1
111
126,8
100,3
2009 .
5,7
26,6
17,2
58,6
12,2
16,3
46
50,6
233,2
,
2010 .
18,1
38
25,6
45
16,8
60
64,3
67
46,5
53
73,2
34
37,1
43
49,1
38
330,7
378
152
________________________________________________________________
13-14 , 12-13
[14].
. , ,
, . , . . , Chrysanthemum .
+15+18 [7,14,15]. +18
, +24+26
. 2010 . , (.
. 1). Chrysanthemum 4-6
.
, .
: ( - ), (
); ( );
( ); ( ); ( ) ( ).
(. 2).
2
Chrysanthemum L.
(2009-2010 .)
2009
2010
Ch. sibiricum
Ch. zawadskii
Ch. weyrichii
Ch. arcticum
23.08
15.09
6.10
-
Swemba Cars
8.09
14.09
5 *
Veria Dark
6.07
29.09
17.10
20.10
10.07,
20.10
2009
2010
7.10
10.09
15.10
10.11
1.12
26.11
1.12
10.11
**
2009
2010
2009
2010
46
31
56
-
67
43
40
22
7.10
26.08,10.11
30
48, 22
3.08, 8.10
1.12
26.08, 10.11
78
23, 34
15.09
26.08
20.10
29.10
36
65
16.09
22.09
24.09
30.09
1.12
1.12
1.12
1.12
25.10
26.11
1.12
1.12
76
70
68
62
31
42
42
21
20.10
10.09, 26.11
18
16, 16
1.12
1.12
10.11
1.12
58
58
21
42
3.10
Lipstick
Conaco Orange
4.10
4.10
25.09
20.10
20.10
10.11
26.08,
10.11
20.10
20.10
4.10
10.11
1.12
26.11
58
16
4.10
4.10
10.11
10.11
1.12
1.12
1.12
1.12
58
58
21
21
. 2011. 3 (98). 14
153
______________________________________________________________
. 2
Padre Liliack
- 3
- 2
Conaco Yellow
C
- 1
- 4
- 6
5.10
5.10
7.10
7.10
10.10
10.10
12.10
12.10
15.10
15.10
15.10
25.10
25.10
25.10
25.10
Spatsoomer
10.08
16.08
18.08
18.08
28.08
30.08
2.09
8.09
25.09
1.10
1.10
1.10
1.10
1.10
5.10
5.10
1.11
1.12
10.11
1.12
10.11
1.12
10.09
1.12
10.09
1.12
20.10
1.12
10.11
1.12
16.08, 29.09
1.12
3.09, 18.10
1.12
3.09
1.12
10.11
1.12
15.11
1.12
17.10
1.12
10.11
1.12
10.11
1.12
10.11
1.12
25.10
13.10
25.10
20.11
25.10
15.10
28.10
15.10
8.10
1.11
25.10
13.10
28.09
11.11
25.10
28.10
25.10
28.10
15.10
28.10
25.10
26.11
28.10
28.10
28.10
28.10
28.10
28.10
22.10
10.11
28.10
28.10
26.11
1.12
1.12
13.10
13.10
1.12
1.12
20.10, 15.11
13.10, 26.11
20.10
1.12
1.12
26.11
1.12
1.12
1.12
57
57
55
55
52
52
50
50
47
47
47
38
37
37
37
27
21
21
34
34
42
21
66, 48
41, 40
36
21
16
41
21
21
21
,
,
29.11
20.11
29.11
29.11
29.11
29.11
26.11
20.11
29.11
29.11
29.11
29.11
29.11
29.11
20.11
29.11
65
97
59
59
66
45
71
51
34
28
57
28
28
28
37
23
36
27
36
33
53
36
63
27
36
46
36
33
33
33
30
33
* , -
.
** ; ; ; ; ; ; .
, (-
)
2010 . . - 2010 . 1-2 1-1,5 , , .
2009-2010 . Ch.
zawadskii - 2 2010 .
9-14 2010 . Ch. weyrichii, , , , , , , Conaco Orange, Conaco Yellow
Lipstick. 1 2 , , , , , , , -4,
-5, -6, , Veria Dark (,
).
, , , , Swemba
Cars. 2010. Ch. sibiricum, -1, -2, -3.
, 2010 -
. ( Chrysanthemum sibiricum, ,
154
________________________________________________________________
). ,
,
( +18 2010 . ), .
. , ,
.
2 , 1,5 . ,
. ,
. Swemba Cars
, .
.
. , , .
2009-2010 . 41,7 . , 20 , ( 40 ) , , . 2009 . 78 97
.
2010 . 1,5
2009 . , 2009 49,4 , 2010
34 . Ch. sibiricum, Ch.
zawadskii, , , , , ,
, , Swemba Cars. 40
- Ch. sibiricum, Ch. weyrichii, Ch. zawadskii ,
, , , , -1, -5, , ,
, , , , ,
Conaco Orange, Conaco Yellow, Spatsoomer, Swemba Cars, Veria Dark. , .
, .
( , , , , , ).
. 2-5
, . , .
. 2011. 3 (98). 14
155
______________________________________________________________
, , , Swemba Cars
1-2 , - 1 2 .
- .
50 97 ,
.
( 40
) Ch. sibiricum, Ch. weyrichii, Ch. zawadskii Ch. sibiricum,
Ch. weyrichii, Ch. zawadskii, , , , , , -1, -5, , , , ,
, , , Conaco Orange, Conaco Yellow,
Spatsoomer, Swemba Cars, Veria Dark.
1. .., .. - //
: . . . . - --, 1988. . 164.
2. , /
- . - . . ., 2008. . 1: . - 276 .
3. .. . . - .: : , 2005. - 191 .
4. .. Chrysanthemum x hortorum Bailey // :
, 75-
. .2. - , 2007. - . 39-41.
5. .. // : (20-22 2001 .). - , 2001. : http://www.ict.nsc.ru/ws/show_abstract.dhtml?ru+18+1412.
6. .. hrysanthemum L. ( , , , ): .
. . / .. . , 2010. 42 .
7. ..
// : , 75-
. .2. - , 2007. - . 58-60.
8. .. (, ). :
, 1982. 167 .
9. . .: , 1972. 91 .
10.
:
http://www.kharkov.ua/region/nature.htm.
11. , .
() hrysanthemum spec. / . . . , 1995. 15 .
12. // . . . . .: - , 1979. . 113. . 3-8.
13. - .. ( ): . / .. -. : - , 1972. 480 .
14. .. . .: , 1990. 110 .
15. Okada M. Classification Of Chrysanthemum Varieties In View Of Their Environmental
Responses To Flowering // J. Hort. Assoc. Jpn. - 1957. - V. 26. P.5972.
156
________________________________________________________________
PHENOLOGICAL PHASES OF SPECIES AND VARIETYS OF CHRYSANTHEMUM L.
IN THE CONDITIONS IN THE CONDITIONS OF THE SOUTH OF CENTRAL RUSSIAN UPLAND
A. S. Stetsovich1
O.A. Sorokorudova1
E.V. Sergeeva2
A.A. Alehin2
1)
Belgorod National
Research University,
Pobedy St., 85, Belgorod,
308015, Russia
e-mail: stetsovich.a@yandex.ru
Botanical garden of the Kharkov
national university of V.N. Karazin,
Klochkovsky St., 52, Kharkov,
61022, Ukraine
2)
e-mail: khbg@i.ua
. 2011. 3 (98). 14
157
______________________________________________________________
576.316.7
(OXYTROPIS DC.)
..
..
, 450080,
. , . ,195,
.3.
e-mail: cyto.ufa@mail.ru
Oxytropis DC.
. O. ambigua, O.
gmelinii O. hippolyti O. uralensis.
: Oxytropis DC., , , , .
Oxytropis DC. . , ,
[3, 5, 7]. Oxytropis DC., ,
, [1, 2]. Oxytropis DC.: . uralensis, . ambigua, O. gmelinii, O. hippolyti
.
, : ( , . ), . ambigua ( ,
-; , -), O. gmelinii ( , -; , . ; , ; , . ), O. hippolyti ( , . ; , . - . ).
[6]. , -13
( 100, 7, 1,6).
(L),
(Lr) (Ic), .
20 - 25 . .. [4]. Oxytropis DC.
(
), Statistica 6.0.
,
.
158
________________________________________________________________
Oxytropis uralensis. .
(. 1). (p>0.05).
1
Oxytropis uralensis
1
2
3
4
5
6
7
8
L
0.54
0.71
0.44
0.53
0.95
0.79
0.74
0.51
.
Lr
0.58
0.49
0.07
0.30
0.65
0.76
0.59
0.34
Ic
0.63
0.89
0.39
0.19
0.95
0.82
0.19
0.85
Oxytropis ambigua.
-
, - (. 2).
- 7, 9
14- , - - 1 3- , - 13
14- .
- 5- , -
3 5- .
- , - (p<0.05), - (p>0.05).
Oxytropis gmelinii. .
(. . 15, 17 18- , -
. 14 17- , . 7, 13 16, 18 19- , . - . 15 17- ) (. 3).
: - . 3 8- , .
. 11, 15, 17 24- , - . 3, 4, 8, 10,
11, 15 18, 23 24- , - 7 18- , - .
- 4- , . 24- , . - .
1, 10, 11, 15 18, 23 24- , . - 16 17 (. 4). : . . 3 13- , - . 4, 7, 13 15- ,
- 6, 19 20- , . 3 7, 17 18-
, . - . 3, 11, 13, 14- , . -
13, 14, 18, 21 22- (. 5).
: - . , . (p<0.05).
. 2011. 3 (98). 14
159
______________________________________________________________
2
Oxytropis ambigua
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
-
L
Lr
0.000028* 0.07
0.00002*
0.09
0.00003*
0.12
0.0001*
0.38
0.00009*
0.39
0.00001*
0.11
0.000002* 0.02*
0.000003* 0.08
0.000001* 0.04*
0.000003*
0.12
0.0002*
0.96
0.0002*
0.78
0.00002*
0.13
0.000006* 0.02*
0.00009*
0.25
0.0006*
1.00
Ic
0.71
0.19
0.09
0.23
0.05*
0.39
0.80
0.58
0.76
0.43
0.91
0.88
0.60
0.77
0.83
0.38
-
-
L
Lr
Ic
0.27 0.01* 0.11
0.52 0.65 0.89
0.18 0.03* 0.15
0.39 0.25
0.41
0.51
0.46
0.47
0.83 0.86 0.26
0.87
0.35 0.24
0.74 0.40 0.68
0.61
0.24
0.47
0.66
0.31
0.79
0.70
0.77
0.50
0.62 0.68
0.13
0.38 0.36
0.73
0.70 0.86 0.34
0.96 0.95 0.94
0.72
0.56
0.57
-
L
Lr
0.0002*
0.53
0.000003*
0.04*
0.00*
0.0005*
0.00001*
0.05
0.00002*
0.13
0.000009*
0.16
0.000007*
0.15
0.00002*
0.36
0.00001*
0.35
0.00003*
0.59
0.0001*
0.75
0.00008*
0.50
0.000002*
0.02*
0.00001*
0.01*
0.0001*
0.29
0.0001*
0.58
Ic
0.25
0.26
0.004*
0.05
0.01*
0.06
0.17
0.36
0.33
0.32
0.45
0.19
0.41
0.54
0.79
0.76
. * O. ambigua 5%- .
Oxytropis hippolyti.
:
. - . - 19- , . - .
5, 9, 18 22, 24- , . - . 4 5-
(. 6). : . - . - 3 19- , . . 2 19- . : . - .
- 1- , . - . 9 19- , .
- . 1, 9, 10, 12- .
. - . (p<0.05).
160
________________________________________________________________
3
- -
- -
- -
Oxytropis gmelinii
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
0.18
0.06
0.02
0.04
0.40
0.55
0.89
0.74
1.00
0.66
0.39
0.58
0.49
0.47
0.47
0.29
0.31
0.18
0.36
0.41
0.92
0.56
0.72
0.34
0.56
0.42
0.41
0.31
0.50
0.34
0.34
0.54
0.40
0.88
0.83
0.66
0.49
0.08
0.01*
0.05
0.03*
0.05*
0.24
0.83
0.63
0.91
0.72
0.09
0.40
0.42
0.08
0.20
0.27
0.39
0.34
0.61
0.78
0.19
0.24
0.58
0.17
0.72
0.96
0.88
0.66
0.96
0.36
0.18
0.33
0.21
0.23
0.34
0.87
0.79
0.46
1.00
0.73
0.67
0.68
0.54
0.57
0.88
0.91
0.91
0.73
1.00
0.96
0.45
0.66
0.82
0.70
0.57
0.50
0.82
0.62
0.34
0.43
0.006
0.001
0.002
0.11
0.11
0.39
0.77
0.38
0.76
0.50
0.30
0.15
0.01*
0.0007*
0.002*
0.001*
0.0008*
0.03*
0.28
0.55
0.47
1.00
0.42
0.60
0.26
0.59
0.43
0.79
0.79
0.26
0.38
0.77
0.36
0.74
1.00
0.47
0.71
0.49
0.35
0.13
0.15
1.00
0.58
0.37
0.47
0.38
1.00
0.12
0.09
0.10
0.04
0.60
0.28
0.78
0.77
0.56
0.54
0.31
0.49
0.72
0.46
0.49
0.75
0.55
0.25
0.17
0.78
0.55
0.72
0.38
1.00
0.79
0.09
0.007
0.02
0.07
0.06
0.05*
0.24
0.24
0.23
0.31
0.30
0.03*
0.03*
0.006*
0.03*
0.07
0.05*
0.03*
0.10
0.13
0.15
0.38
0.42
0.43
0.26
0.10
0.29
0.29
0.59
0.57
1.00
0.77
0.76
0.74
0.73
0.28
0.06
0.006*
0.006*
0.007*
0.02*
0.42
0.41
0.23
0.72
0.38
0.42
0.29
0.57
0.28
0.19
0.43
0.18
0.16
0.24
0.38
0.13
0.18
0.49
0.28
0.71
1.00
0.53
0.37
0.77
0.18
0.41
0.76
0.28
0.08
1.00
. * - O. gmelinii 5%- .
. 2011. 3 (98). 14
161
______________________________________________________________
4
0.81
16
0.68
17
0.69
18
19
20
21
22
23
0.34
0.77
0.86
0.38
0.71
0.43
24
1.00
15
0.60
0.67
0.68
12
13
14
- -
0.96
- -
11
0.07
0.79
0.89
0.85
0.69
0.95
0.99
0.55
0.91
0.06
0.04
*
0.30
0.57
0.37
0.02
*
0.10
0.04
*
0.08
0.49
0.40
0.60
0.25
0.07
0.00
7*
- -
0.55
0.19
0.03*
0.06
0.97
0.67
0.09
0.04*
0.17
0.55
1
2
3
4
5
6
7
8
9
10
Oxytropis gmelinii
0.96
0.94
0.13
0.42
0.58
0.88
0.82
0.58
0.36
0.41
0.68
0.82
0.33
0.84
0.65
0.35
0.36
0.20
0.26
0.72
0.20
0.28
0.04*
0.03*
0.71
0.71
0.08
0.007*
0.12
0.01*
0.51
0.15
0.51
0.26
0.60
0.55
0.05*
0.13
0.62
0.14
0.30
0.26
0.22
0.03*
0.67
059
0.41
0.43
0.79
0.79
0.05
0.72
0.15
0.30
0.88
0.82
0.82
0.23
0.29
0.27
0.02*
0.96
0.38
1.00
0.96
0.36
0.34
0.05
0.19
0.02*
0.70
0.76
0.57
0.30
0.92
0.26
0.24
0.46
0.81
0.22
0.65
0.79
0.03*
0.60
0.81
0.09
0.02*
0.45
0.62
0.55
0.43
0.79
0.72
0.93
0.11
0.30
0.61
0.98
0.29
0.69
0.79
0.41
0.74
0.11
0.99
0.91
0.18
0.82
0.40
0.81
0.81
0.46
0.26
1.00
0.03*
0.35
0.80
0.21
0.02*
0.24
0.34
0.26
0.03*
0.15
0.46
0.48
0.005*
0.03*
0.07
0.54
0.01*
0.02
0.81
0.84
0.007*
0.01*
0.27
0.71
0.32
0.76
0.60
0.87
0.74
0.45
0.47
0.43
0.86
0.47
0.005*
0.31
0.48
0.15
0.11
0.007*
0.03*
0.93
0.39
0.22
0.35
0.33
0.51
0.28
0.57
0.08
0.57
0.95
0.47
0.27
0.87
0.82
0.51
0.08
0.03*
0.95
0.89
0.78
0.31
0.009*
0.14
0.24
0.77
0.62
0.72
0.36
0.98
0.42
0.005*
0.98
0.40
0.005*
0.05*
0.41
. * - O. gmelinii 5%- .
162
________________________________________________________________
5
- -
- -
- -
Oxytropis gmelinii
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
0.61
0.40
0.21
0.64
0.43
0.27
0.66
0.74
0.92
0.66
0.89
0.69
0.82
0.77
0.09
0.92
0.73
0.83
0.26
0.13
0.66
1.00
0.71
0.53
0.41
0.07
0.01*
0.07
0.17
0.09
0.13
0.62
0.76
0.70
0.55
0.93
0.02*
0.12
0.79
0.69
0.58
0.87
0.93
0.89
0.67
0.54
0.60
0.83
0.59
0.44
0.57
0.36
0.49
0.31
0.33
0.61
0.66
0.79
0.63
0.69
0.11
0.20
0.70
0.45
0.14
0.07
0.29
0.27
0.24
0.14
0.50
0.37
0.84
0.90
0.56
0.54
0.69
0.61
0.68
0.54
0.23
0.12
0.13
0.13
0.65
0.26
0.11
0.22
0.77
0.48
0.50
0.74
0.40
0.33
0.60
0.30
0.17
0.30
0.20
0.02*
0.54
0.53
0.04*
0.39
0.83
0.95
0.45
0.61
0.008*
0.06
0.04*
0.76
0.36
0.69
0.28
0.15
0.99
0.52
0.88
0.39
0.28
0.94
0.06
0.64
0.12
0.03*
0.58
0.85
0.72
0.86
0.72
1.00
0.06
0.10
0.18
0.37
0.24
0.10
0.02*
0.007*
0.10
0.13
0.76
0.11
0.75
0.45
0.48
0.26
0.22
0.53
0.38
0.33
0.25
0.24
0.15
0.24
0.82
0.38
0.95
0.17
0.96
0.34
0.63
0.22
0.67
0.31
0.35
0.08
0.76
0.27
0.002*
0.005*
0.03*
0.006*
0.01*
0.30
0.88
0.90
0.26
0.61
0.43
0.78
0.49
0.23
0.04*
0.04*
0.24
0.20
0.10
0.37
0.88
0.47
0.29
0.08
0.05*
0.20
0.07
0.21
0.25
0.91
0.34
0.22
0.03*
0.10
0.004*
0.006*
0.05
0.09
0.39
0.58
0.54
0.84
0.66
0.10
0.28
0.39
0.44
0.50
0.24
0.11
0.76
0.12
0.15
0.25
0.42
0.17
0.28
0.24
0.03*
0.01*
0.20
0.62
0.22
0.01*
0.07
0.14
0.04*
0.01*
0.21
0.88
. * O. gmelinii 5%- .
. 2011. 3 (98). 14
163
______________________________________________________________
6
Oxytropis hippolyti
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
. -
. -
L
Lr
Ic
0.75
0.69 0.008*
0.79
0.35
0.29
0.18
0.02*
0.24
0.45
0.09
0.61
0.91
0.40
0.59
0.79
0.80
0.79
0.77
0.26
0.77
0.95
0.57
0.51
0.65
0.95
0.27
0.42
0.49
0.38
0.59
0.79
0.26
0.16
0.14
0.20
0.44
0.59
0.17
0.29
0.30
0.25
0.49
0.61
0.45
0.93
0.67
0.53
0.67
0.89
0.06
0.17
0.11
0.35
0.04* 0.01*
0.29
0.49
0.69
0.21
0.22
0.33
0.91
0.45
0.82
0.99
0.93
0.47
0.09
0.94
0.69
0.51
. -
.
L
Lr
Ic,%
0.10
0.74
0.74
0.79
0.05*
0.43
0.50
0.17
0.85
0.09
0.93
0.07
0.04*
0.47
0.03
0.13
0.80
0.13
0.32
0.27
0.32
0.16
0.60
0.21
0.03*
0.58
0.05*
0.07
0.94
0.06
0.11
0.79
0.45
0.08
0.82
0.41
0.24
0.23
0.09
0.15
0.60
0.14
0.17
0.65
0.06
0.27
0.46
0.32
0.05
0.40
0.06
0.02*
0.17
0.02
0.001* 0.005* 0.007*
0.02*
0.24
0.08
0.007*
0.18
0.11
0.03*
0.91
0.14
0.18
0.46
0.17
0.05*
0.63
0.18
. -
.
L
Lr
Ic
0.20
0.94 0.004*
0.61
0.30
0.07
0.05
0.33
0.33
0.02* 0.08
0.20
0.04*
0.13
0.11
0.21
1.00
0.21
0.21
0.99
0.21
0.19
0.97
0.06
0.09
0.64 0.003*
0.31
0.55
0.007*
0.31
0.60
0.07
0.71
0.10
0.04*
0.68
0.09
0.76
0.72
0.13
0.75
0.50
0.35
0.26
0.32
0.76
0.72
0.13
0.49
0.99
0.36
0.83
0.15
0.21
0.76
0.10
0.10
0.44
0.61
0.13
0.71
0.15
0.16
0.91
0.16
0.16
1.00
0.73
0.06
0.39
0.06
. * - O. hippolyti 5%- .
164
________________________________________________________________
2. .. (Oxytropis DC.) // II . . .-. . : . , 2010. . 122-124.
3. .. - Oxytropis DC. //
. ., 1987. . 6. . 169.
4. .. . . .: , 1973. 256 .
5. .., .., ..
( ) // . . 1990. . 75. 11. . 1622-1624.
6. .. . .: , 1980. 304 .
7. .., .., .. Oxytropis (Fabaceae) // . . 1998. . 83, 6. .138-139.
Comparative analysis of morphometric parameters of chromosomes of four South-Ural species of the Oxytropis DC. from thirteen habitat allowed to reveal some similarities and differences between populations of studied species. The obtained results showed higher degree of
differentiation of O. ambigua, O. gmelinii and O.hippolyti and a lower
degree of differentiation of O. uralensis.
Key words: Oxytropis DC., morphometric parameteres of chromosomes, populations, variability, South Ural.
. 2011. 3 (98). 14
165
______________________________________________________________
634.75 631.526 (571.56)
.. 1
.. 1
.. 2
1)
677001, . ,
. -, 23/1
,
308015, . , . , 85
. , ,
, , .
: , ,
, , ,
, , , .
e-mail: sorokopudov@bsu.edu.ru
Fragaria
, , , , .
, , .
2500 ./ .
, .
, , , , .
. .. , .. , .. , .. . , ,
[4;6].
, ,
10% [4], .
,
; , , ,
, [2; 9].
- , ( ,
166
________________________________________________________________
). 18-19 ( 2294 .). -
86% .
93,6 /2, 15,6 /2 ,
.
0 150-165 :
.
10
1100-1600, [1].
.
-
, .
. , , , ,
[10].
0. I .
30 , 65-120 .
, ,
- .
,
90- .
.
,
- , (1995) (1999).
, ,
.
, ,
, , , . - ( 7,4 8,3). 0,20
2,5-3,0%.
, 100 . , (
7-10 ) 20%.
. 5964,9 (.) 2000-2001 . (-). - 2000-2001,
2001-2002, 2003-2004 . -55 -
. 2011. 3 (98). 14
167
______________________________________________________________
2006 . ( 20 ) 2002-2003 . -27,6; 2009-2010 . -25,6. 1999-2000 .
(1 ), 2004-2005 . (28 ), 2006-2007 . (10 ).
1271,1 (2004 .)
1938,7 (2008 .). , , 2006 ., 28 10 (105 ),
2000 ., 23 18 (56 ).
, Fragaria orientalis Los., , - .
- .
- .
, . , 1999 ., , , .
165,8 , 54,7 ., 324,8 ., 112,5 .
, Fragaria ananassa Dush., 90- .
- . ,
,
-
,
( ,
, ; , 1,5-2 .
). , , : , , ,
, , , . .. , , , 66; :
, , , .
, Fragaria veska ssp. veska f.
semperflorens Duch., 1999 .
. . : , , , , , , , . .
,
,
. 100% . ,
, , , .
1,5 3,5%. : , , , .
,
, , , : , , , .
168
________________________________________________________________
. (16-03, 33-07) (6-03, 7-03, 6-07). ,
, ,
(24-08, 28-08). , , , , , . , , , ,
. , : , , , , , , .
,
(32-03, 35-03, 37-03), F. orientalis Los.
. , . , , F. orientalis Los. 30-35 , , , , - , , , .
, , .
, .
, 67,5 . , . , . ,
, 4 ,
. I . I .
F. orientalis Los. . 4 .
.
, , :
1)
;
2) , , ;
3) , , ,
1. , .. / .. , .. . : , 1977. 192 .
2. , .. / .. . : . -,
1973. .120.
3. , .. : /
.. , .. , .. . .: /,
2001. . 117.
4. , .. : / .. .-, 1990.
5. .. / .. , .. // 1
2007. . 51-53.
. 2011. 3 (98). 14
169
______________________________________________________________
6. , .. / .. / . . .-. . .. . .: , 1986. . 91-94.
7. , . (
, .-. .. ) : - - , 1995. . 387-416.
8. , .
; - - , 1999. . 417-443.
9. .. / .. , .. // . 1953
10. .18-20.
10. , ..
/ .. . , 1936. .26-33.
e-mail: agronii@mail.ru
2)Botanic
e-mail: sorokopudov@bsu.edu.ru
170
________________________________________________________________
, .
,
.
,
.
.
20 ,
2003
2004
2004
2005
2005
2006
2006
2007
2007
2008
2008
2009
2009
2010
2002
2003
2001
2002
2000
2001
1999
2000
5145,0
5964,9
5035,2
4897,1
5377,2
-5319,5
5273,9
4959,8
4630,3
-5188,1
-5294,6
+1853,7
+1895,4
+1829,1
+1848,1
1702,3
+1932,6
+1934,7
+1907,6
+2097,9
+2052,0
+2008,6
3291,3
4069,5
3206,1
3049,0
3674,9
3386,9
3339,2
3052,2
2532,4
-3136,1
3286,0
1999
2000
2001
2002
2003
2004
2005
2006
2007
2008
,
-33,3
-37,2
-40,1
-37,8
-40,0
-41,1
-39,7
-45,0
-33,0
-37,1
+18,9
+18,5
+21,2
+19,3
+20,3
+18,0
+17,9
+17,9
+15,9
+19,7
,
2009
-35,7
+20,1
33
13
39
12
24
39
47
48
21
20
23
-49,8
-52,3
-52,5
-51,3
-52,6
-53,9
-55,0
-50,1
-49,5
-52,2
-52,0
-15,6
-22,5
-13,2
-27,6
-20,9
-16,6
-18,5
-15,5
-15,5
-19,6
-25,6
. 2011. 3 (98). 14
171
______________________________________________________________
577: 112.592.542
IRIDACRAE JUSS.
ASPARGACEAE JUSS
..
. .. ,
., 4,
127276
e-mail: gbs@ahc.ru
Iridaceae ( Iris L, Sysirinchium L.) Asparagaceae ( Asparagus L.).
. , -
.
Iridaceae Liliales .
: , , .
172
________________________________________________________________
(Iris), (Sysirinchium L.) (Asparagus L.).
.
.. . , , 80%- . : 10% NaCl , 70% (), 0,2 2,0%-
, , .
1030.
.. [13] (.., .., .., ..,
.. .) .
, Ae)
, . ,
. .
.
Iris, Sysirinchium Asparagus. Iris .. [4]. Limnaris , Apogon (29,13-58,10 % )
(24,3-30,7%) (31,52-45,02%).
.
1,7% 7,04%. (Ae) 0,49
1,03 (. 1).
1
( % )
Iris L.
I. gracilipes A.Gray.
I. kaempferi Sieb.
I. clarkei Baker.
I. sibirica L.
I. laevigata Fisch.
I. setosa Pall.
I. orientalis Thunb.
I. pseudocorus L.
I. tenax Dough.
I. wilsonii C.M.Weight.
6,9
5,6
6,3
19,9
13,8
15,7
10,2
10,7
14,2
6,1
I. paradoxa Steven.
I. atrofusca Baker.
I. susiana L.
15,7
16,0
15,4
0,2%
2,0%
Limnaris, Apogon
24,3
4,1
9,6
1,9
26,3
4,1
9,5
2,0
25.4
5,4
8,3
2,3
27,7
7,6
12,6
2,8
26,1
12,0
9,6
2,2
28,1
5,8
15,1
2,0
27,1
9,2
12,2
3,4
27,6
14,5
24,6
7,0
30,7
2,3
8,0
7,0
25,4
4,2
4,4
1,7
Iris, Onocyclus
35,8
7,8
13,1
2,0
38,2
6,0
8,0
2,5
39,5
6,5
7,5
2,0
Ae
0,49
0,61
0,60
0,92
0,84
0,73
0,69
0,70
1,03
0,50
1,15
1,30
1,37
. 2011. 3 (98). 14
173
______________________________________________________________
, Onocyclus
(51,50-55,90%), , (35,8-39,5%).
(2,0-2,5%). Ae , Apogon (1,15-1,37).
Apogon Onocyclus.
(
) 41,7-62,9% 36,6-41,2% .
.. .
( Apogon)
.
Sysirinchium . Sysirinchium
(28,66-51,98%) (15,6-30,7%).
(22,3-43,3%). ,
1,8-6,7% ( S. donglesii, S. convolutum, S. idahoense 8,1-13,3%), Ae 0,62 1,01. 35,5-58,6%.
, . Iris
, ,
, Ae. (5, 9)
. ,
, , , Aspargales .
Iridaceae Aspargales, .
.
(. 2.).
2
( % )
28
5,6-18,6
15,6-39,5
8,6-26,6
1,8-10,8
15
1,8,-3,8
19,3-22,4
45,2-55,1
2,0-11,2
Ae
0,501,37
0,370,50
35,5-69,9
62,2-72,1
,
. Ae ,
, .
, , , .
1. . . .: . 1982. . 6. . 180-194.
2. Dies L. Iris //Die naturishe Pflancenfamilien. Leipzig. Ed. Engler E. Prentl. 1930.
Bd. 2. 5. S. 200-250.
174
________________________________________________________________
3. Hatchinson J. The Familie of flowering plants monocotyledons //Oxford: Claredon press.
1973. 652 p.
4. .. . .: . 1961. 215 .
5. Takhtajan A.L. Diversity and classification of Flowering Plants //New York. Columbia University Press. 1996. 1. 250 p.
6. .. Iridaceae Liliaceae //. . 1983. . 68. 1. . 49-55.
7. Thorne R. Classification and Geography of the Flowering plants //Bot. Rev. 1992. V. 58.
P. 1-348.
8. Huber H. Die Samenmerkmele and Verwend-sohafsverhltnisse der Lilifloren //Mil. Bot.
Staatsamel. 1969. V. 8. S.219-538.
9. Dahlgren R. A commentary on diagrammatic presentation of the Angiosperms in relation to
the distribution of characters //Bot. notis. 1975. - V. 128. P. 119-146.
10. Hegnauer r. Chemotaxonomie der Pflanzen. Monocotyledonene Basel und Stuttgard.
Berkhnser Verlag. 1963. B. 2. 540 S.
11. Carles J. Climisme et classification Cher les Iris //Rev. gen. bot. 1935. V. 47. 1.
363 p.
12. . .
-: . 2006. 845 .
13. .. . .: .
1966. 327 .
14. .. Liliales Aspargales //. . 1994.
. 79. 3. . 1-12.
__________________________________________________________________________
634.75:631.524.86:632.112
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..
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. , . , 49
e-mail: galina.govorova@mail.ru
,
2010 .
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176
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-21.0 (1879)
-7.5 (1885)
-2.3 (1916)
1.3 (1886)
-1.2 (1885)
-8.5 (1881)
2.1
7.4
12.0
13.8
12.0
7.0
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12.8
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18.4
16.4
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10.8
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22.1
23.2
21.3
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.
28.0 (1950)
33.2 (2007)
34.7 (1901)
38.2 (2010)
37.3 (2010)
32.3 (1890)
2
1951- 2010 .
38
52
84
90
80
67
8 (1960)
7 (1986)
4 (1951)
6 (1997)
20 (1955)
12 (2005)
98 (1986)
120 (1976)
162 (1991)
180 (2008)
163 (1973)
131 (1996)
30 (1965)
39 (1976)
63 (1970)
62 (1981)
59 (2003)
49 (2004)
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2. .. . .: , 1968.- 26 .
3. .. -
// -. - 1983.- 1, 15-24 .
4. .., .. . ,1966.-348 .
5. .., .. (, , ), ., 2004.-348 .
6. ..
. - .: - , 1979.- 11 .
7. .M. , .: . ., 1964.- 667 .
8. .M. // . - ., 1971.- 507-513 .
9. .. , // .- , : , 1972.- 217-223 .
10.
Key words: strawberry varieties, hybrids, drought resistance, heat resistance,disease, pests, immunity, nutrition, and
ecology.
e-mail: galina.govorova@mail.ru
180
________________________________________________________________
581.16:582.681.81
-
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620144, .,
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e-mail: olgae06@mail.ru
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181
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S. serphyllifolia
S. glauca
S. saxatilis
S. lanata
S. finnmarchica
S. simulatrix
S. nakamurana
S. myrsinites
S. fuscescens
S. calcicola
S. recurvigemmis
S.x Brekkavier
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90
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60
62
83
77
43
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182
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2
,
S. retusa
S. finnmarschika
S.repens
S.x cottetii
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S.recurvigemmis
S.glaucosericea
S.glauca
S. x Brekkavier
S. repens Green carpet
n
M+m
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60
92
54
20
62
17
10
13
50
53
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46
40
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100
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, ,
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183
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3
S.nummularia
S.serphillifolia
S.saxatilis
S.breviserrata
S.repens
S.repens
S.repens GreenCarpet
S. x ausserdorferi
S.nakamurana
S. retusa
S. glaucosericea
S.reptans
S.fuscescens
S.fuscescens
S.glauca
S.lapponum
S.lapponum
S.glauca
S.Brekkavier
S.lanata
S.lanata
S.arbuscula
S.helvetica
1491
3597
2691
15597
11997
19297
898
18497
8997
16697
5797
4491
2791
2791
12097
1091-11
998
862
4897
14697
598
8898
5997
20
29
26
100
100
67
93
25
86
82
60
85
79
13
50
60
40
3
24
29
7
38
71
18
0
37
100
100
50
71
10
75
50
40
86
87
22
60
100
60
20
36
40
2
50
98
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.
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184
________________________________________________________________
. .
, , . ,
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. . 4.
4
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S. X finnmarchica
S. X simulatrix
S. nakamurana
S. hastata x S. reticulata
S. lanata
S. breviserrata
S. saxatilis
S. arctica v. crassijulis
S. repens Green Carpet
2597
9197
8997
8498
598
15597
1997
14197
898
30 - 6
95
65
72
36
17
47
25
27
29
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8,56
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7,18
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4,53
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87
83
79
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, , , , , .
,
, .
, , . ,
( ) , , , . .
.
, .
, ,
, - ,
, , .
(70-100%)
(40-70%) ,
.
(70-100%): S.foetida Schleich ex Lam. Et DC.,
S.breviserrata Flod., S.ivigtuttiana Lundstr., S.caucasica Anderss., S.calcicola Fern. et
Wieg., S.phlebophylla Anderss.,
S.arctophila Cock. ex Heller., S.apoda Trautv.,
S.nakamurana Koidz., S.helvetica Vill., S.lanata L., S.reinii, S.serphyllifolia, S.retusa, S. bicolor Ehrh. ex Willd., S.lapponum L., S.arbuscula L.
(40-70%) : S.nummularia Anderss., S.rectijulis Ledeb. ex Trautv., S.pyrenaica Gouan, S.saxatilis Turcz. ex Ledeb.,
S.saposhnikovii A.Skvorts, S.turczaninovii Laksch., S.herbaceae L.,
S.helvetica Vill., S.bicolor, S.lapponum L., S.arbuscula, S.hastata L.
185
___________________________________________________________________________
(<30%) , , , . , ,
, : S.arctica, S.reticulata,
S.polaris, S.recurvigemmis, S.herbaceae . ,
. , , , , .
1. - , .
(), .
2. ,
.
3. - , . , ,
.
4.
. , .
1. .. . / . . // . : . IV . -, 1986. . 553-554.
2. ..
. : -, 1982.-288.
3. , .. . .: . -, 1965. 98.
4. , .. , . ..: , 1950. 167 .
5. .. . - .: , 1938. - 232.
6. .., .., .., .. ( ). .: . , 1968.
The peculiarities of artificial vegetative propagation of arctomontane willows, depending on the type of cutting, timing of cutting,
the substrate and the type of stem were identified.
Keywords: willow, vegetative propagation, cuttings.
186
________________________________________________________________
635.92
..
..
-
, 450080
, . , 195/3
e-mail: zainetdinova80@mail.ru
Hemerocallis L. - .
,
.
.
: , , , , .
187
___________________________________________________________________________
.
. .
1
H. citrina
H. dumortieri
H. fulva
H. lilio-asphodelus
,
,
,
,
, ,
H. middendorfii
- ,
H. minor
, ,
, 1999 .
-, 1999 .
, 1952 .
, 1945 .
, 1945 .;
, 1962 .
, 2000 .
, , 3-5. , . 2008
(11.04),
2.9 .
H. middendorfii - 41 . H. citrin (100 ).
: H.lilio-asphodelus, H. dumortieri 42, H. minor 58, H. fulva
68 .
. . , .
H. middendorfii, H. dumortieri H. lilio-asphodelus (III ).
H. minor (II ). H. fulva (III ), H. citrin
(II ) (.).
( 20
73 ). (
2 4 ): H. dumortieri, H. middendorfii, H. lilio-asphodelus, H. minor; ( 4 8 ): H. citrina, H. fulva.
.
H. minor 7-8 . 10
, , .
0.3 , ( 75%). - 11 .
1 3 .
188
________________________________________________________________
I
II III
I
II III
I II III
I II III
H. citrina
H. dumortieri
H. fulva
H. lilio-asphodelus
H. middendorfii
H. minor
H. middendorfii, ,
6-7 . 11 .
0.6 . .
( 90%). 9 . 1 .
H. dumortieri , , 8
. 9 .
- 0.6 . , (
90%). 10 .
- 1 .
H. lilio-asphodelus 7-9 . 10 . 0.6 .
, ( 90%). 11 . , , 2
.
H. citrina ( 22 ) 10 . , .
.
8 . 9 . . 0.7 . 11 . - 1 .
H. fulva, , 8
; - 9 . ,
(50%). - 0.9 . 11-12 . 1 .
,
-. 3- (H. minor, H. middendorfii, H. dumortieri) . H. dumortieri. 120 . , - 28%. H. middendorfii 32 ., - 13%.
H. minor (.2).
, ( ). , - , - .
( ).
189
___________________________________________________________________________
2
,
./.
,
./.
, ./.
H. dumortieri
H. middendorfii
H. minor
152.8
81.2
171.1
1205.4
322.5
342.6
302.1
302.3
382.8
,
./.
81.6
40.9
20.5
, %
28
13
5
,
. , , : , , , ,
.
(.3).
3
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
H. citrina
H. dumortieri
H. fulva
H. lilio-asphodelus
H. middendorfii
H. minor
7-
+
+
+
+
+
+
5
6
5
5
6
6
190
________________________________________________________________
0.5-1 (H. minor), - 1-2 (H. dumortieri, H. middendorfii, H. lilio-asphodelus), - 2-3 (H. citrina, H. fulva).
:
, (H. dumortieri, H. middendorfii, H. minor); (H. lilio-asphodelus); (H. citrina, H. fulva).
, . (H. dumortieri, H. middendorfii)
(H. lilio-asphodelus, H. citrina, H. fulva, H. minor). H. citrina, H. fulva, H. dumortieri, H. lilio-asphodelus;
H. middendorfii, H. minor.
, H. citrina, H. lilio-asphodelus . H. dumortieri, H. middendorfii, H. minor -
. H. fulva .
3 (H. middendorfii) 54 (H. citrina) . 7 (H. dumortieri) 14 .
(H. minor). ( ) H. citrina (702 ); H. middendorfii (48), H. dumortieri (56).
H. fulva (200), H. lilio-asphodelus (100), H. minor (84).
.
, , (H. minor, H. lilioasphodelus), 3-5 (
), ( ) . , ,
9 30 .
H. fulva, H. minor. , (
, ) - H. fulva.
.
. .
- (. 4) 5 , 130 (H. citrina, H. dumortieri, H. fulva, H. middendorfii, H. minor).
.
(127) H. lilio-asphodelus.
.
. (H. fulva), (H. fulva, H. middendorfii),
(H. minor, H. dumortieri), , ,
, (H. fulva, H. citrina), , (H. minor,
H. lilio-asphodelus).
H. citrina
H. dumortieri
H. fulva
H. lilio-asphodelus
H. middendorfii
H. minor
100
97
92
94
88
91
90
50
44
40
48
39
40
42
()
150
141
132
142
127
131
132
191
___________________________________________________________________________
6 . :
- H. minor, H. middendorfii, H. dumortieri;
- H. middendorfii, H. dumortieri;
- H. citrina, H. fulva;
- - H. citrina, H. fulva, H. minor;
- , - H. fulva, H. minor,
H. middendorfii, H. dumortieri.
.
1. .., ..
. - : , 2006. - 296 .
2. .. // .
. - 1974. - . 59, - 6. - . 826-831.
3. . . - : , 1987. 248 .
4. . / . .. . .: , 1972. - 135 .
5. .., .., .. . - .: , 2006. - . 1. - 211 .
6. .. // . . 52. .: , 1964. . 79-87.
7. .. . - .: . ., 1962. - 378 .
8. . . . - : , 1973. - 89 .
9. Hemerocallis minor Mill., [ ]. URL:
http://bogard.isu.ru/herb/herb.files/hemerocallis_minir.html ( : 9.04.2009).
192
________________________________________________________________
581.1
RHODODENDRON L.
.. 1
.. 1,2
1)
-
, 690024 . , . , 142.
e-mail: koksheeva@yandex.ru
, 690010
. , . , 27
14 Rhododendron L., .
.
.
2)
: Rhododendron, , , .
e-mail: ntsarenko@mail.ru
193
___________________________________________________________________________
14 Rhododendron L. 4 (. 1).
2009 3-5 .
2
[9].
-2 (60, 150, 600). 1430 .
Nicon E 880.
Excel
2007.
Rhododendron
Rhododendron
Pentanthera
Hymenantes
Tsutsusi
Rh. dauricum
Rh. mucronulatum
Rh. micranthum
Rh. sichotense
Rh. japonicum
Rh. luteum
Rh. schlippenbachii
Rh. vasey
Rh. ponticum
Rh. smirnowii
Rh. catawbiense
Rh. kaempferi
Rh. obtusum
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poukhanense
, ,
,
, ,
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.
.
.
.
.
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.
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Rhododendron (.1).
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194
________________________________________________________________
, , Pentanthera Hymenanthes (. 2).
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2
Rhododendron L.
,
____
min-max
____
min-max
..,
____
min-max
Rhododendron
Rh. dauricum
Rh. mucronulatum
Rh. sishotense
Rh. micranthum
138,6
112,5-167,5
126,2
111,3-161,3
124,6
88,8-168,8
173,6
117,5-220
7,5
6,9
6,2
5,3
6,6
5,4
5,4
6,9
3,8
6,2
4,8
6,6
31
25-37,5
36,2
25-47,5
32,3
25-42,5
39,7
25-57,5
2-3
1-3
2-3
1-3
73
62,5-87,5
60,5
52,5-80
58,5
42,4-87,5
102,7
72,5-137,5
3-4
3-4
3-4
3-4
Pentanthera
Rh. japonicum
Rh. luteum
Rh. vaseyi
Rh. shlippenbachii
175
156,3-193,8
347,3
210-402,5
233
180-280
175,3
110-236,3
4,6
8,1
5,3
7,6
5,9
9,5
7,3
9,9
6,3
7,9
10,3
5,1
9,5
7,5
28,3
22,5-40
146
87,5-172,5
41,8
27,5-45
35,3
22,5-52,5
2-3
3-4
2-3
2-3
102
87,5-122,5
151
62,5-187,5
146,8
105-177,5
93,5
47,5-112,5
4-5
4-5
6-8
4-5
Tsutsusi
Rh. kaempferi
Rh. yedoense f.
poukhanense
Rh. obtusum
110,4
101,3-121,3
143
117,5-141,3
107,8
82,5-131,3
5,3
5,5
8,1
6,5
6,1
7,4
5,3
5,5
8,1
18
15-22,5
30,3
20-37,5
26,8
22,5-35
1-3
1-3
1-3
59,5
47,5-70
67,5
50-92,5
45,3
30-62,5
4-5
6-8
4-5
Hymenantes
Rh. catawbiense
Rh. ponticum
Rh. smirnovii
230,3
192,5-250
329
262,5-382,5
308,3
282,5-345
5,5
9,8
8,5
11
5,8
8,3
10
8,6
9,5
12
43,3
37,5-52,5
60,5
30-100
36,8
30-45
1-2
2-3
2-3
133,6
100-175
221,3
162,5-262,5
215,8
201,3-250
6-8
5-6
6-8
195
___________________________________________________________________________
, ,
Rh. schlippenbachii.
.2.
Rh. dauricum
. 3.
Rh. luteum: ;
; ; ;
;
. 4. Rh. obtusum
. 5.
Rh. yedoense f. Poukhanense:
; ;
; ;
;
196
________________________________________________________________
. 6.
Rh. Catawbiense: ;
. 7. Rh.
Catawbiense: ;
; ;
;
,
..
, Rhododendron.
[10] ,
Rhododendron Rhodorastrum [11] Rhododendron.
Rhododendron .
Rhododendron, : ;
. ,
47-59%. ..
Rh. schlippenbachii
Pentanthera ,
, , .
53%.
.
- Rh. japonicum
Rh. vaseyi. . ,
, ,
. 58-63% .
Rh. luteum, [12],
.. ,
, , , 43%. , , ..
Tsutsusi
. , Rh. yedoense f. poukhanense. , , -
197
___________________________________________________________________________
, 54%.
Rh. kaempferi, .
,
.
47%. Rh. obtusum ,
...
, , , 42%.
Rh. catawbiense, Rh.
ponticum Hymenanthes,
, , . .. (58% 67% ). .. , , . , , Rh.
smirnowii. (70% )
.., .
.. Rhododendron L.
:
, ,
,
, ..
:
Rh. dauricum, Rh. micranthum, Rh. mucronulatum, Rh. sichotense, Rh. shlippenbachii,
Rh. japonicum, Rh. yedoense f. poukhanense; Rh. vasey; Rh. smirnowii. Rh. catawbiense, Rh. obtusum, Rh. kaempferi, Rh. ponticum
.
1. . . . .: +, 2001. 191 .
2. . . . : ,
1981. 95 .
3. .. . : , 2006. 175 .
4. . . . : , 1981. 230 .
5. .. //
. 1980. . 76-80.
6. .. . : 2006. 131.
7. ..
// . 1979. . 52-59.
8. . . .: , 2001.
528 .
9. , .. . .: , 1974. 288 .
10. Chamberlain, D., R. Hyam,, G. Argent, G. Fairweather & K.S. Walter. The genus Rhododendron. Its classification and synonymy. //Edinburgh: Royal Botanic Garden. 1996. 181 p.
198
________________________________________________________________
11. Yamazuki T. A. Revision of the Genus Rhododendron in Japan, Taiwan, Korea and Sakhalin.
Tokyo: Tsumura Laboratory, 1996. 179 p.
12. .., .., .., ..
. .: , 2003. . 3. 214 .
1)
e-mail: koksheeva@yandex.ru
Eastern National University,
Oktyabrskaya St. 27, Vladovostok,
690010, Russia
The anatomy of bud scales of generative buds of 14 Rhododendron species introduced in the south of Primorsky region was
studied. It was identified the diagnostic and adaptive features. The
dependence of the generative buds winter hardiness from the bud
scales structure was ascertained.
2)Far
e-mail: ntsarenko@mail.ru
scales.
199
___________________________________________________________________________
581.9
AMBROSIA ARTEMISIIFOLIA L., IVA XANTHIIFOLIA L., XANTHIUM ALBINUM (WIDD.)
H. SCHOLZ - 1
..
..
, 308015, . , 85
e-mail: tokhtar@bsu.edu.ru
- . -
. .
. Xanthium albinum (Widd.) H. Scholz - . Iva xanthiifolia L.
, Ambrosia artemisiifolia
L. .
.
- .
: Ambrosia artemisiifolia L., Iva xanthiifolia L.,
Xanthium albinum (Widd.) H. Scholz, , .
, , ,
, ,
, .
.
.
, (--, , 1992).
, . , , , , .
. Ambrosia L.,
Iva L., Xanthium (Widd.) H. Scholz ( Asterea, Asteraceae) .
, 16.740.11.0053 01.09.2010
200
________________________________________________________________
Ambrosia artemisiifoli, Iva xanthiifolia, Xanthium albinum. ,
,
, Asterea (Asteraceae)
. - , ,
- ,
.
,
.
, .
: 0 - , 1 ,
2 , 3 , 4 , 5 .
: - , , , , , , ; , ,
. 10 .
, , , , . Ambrosia artemisiifolia
,
, ,
( ) (Fumanal, 2007).
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(Chauvel, 2010). , ,
, (Otto, 2008). . ,
, , , (Bohren, 2007).
: , , , , , ,
, .
. .
.
201
___________________________________________________________________________
/
/
, ,
Ambrosia artemisiifolia
4
4
2
4
3
3
0
Iva xanthiifolia
5
5
3
2
4
5
3
Xanthium albinum
4
5
4
3
2
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5
Ambrosia artemisiifolia
Iva xanthiifolia
Xanthium
albinum
.
..
...
6
5
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2
1
0
. , . X. lbinum , , , ,
.
. I. xanthiifolia ,
. , . A. rtemisiifolia
, .
. ,
202
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- , , , . ,
.
. , .
X. albinum 1922 .
.. . I. xanthiifolia
1958 . A. rtemisiifolia , 2006 . , . ,
, , ,
. , .. 1967 . (, 2010). ,
X. lbinum .
1922 , .
,
, , . , , ,
, . , . Ambrosia
. , 50 .. A. rtemisiifolia
, , 55 ..,
,
(Meusel, 1992). , , ,
60 ..
(, 2001). .
203
___________________________________________________________________________
3. .. . .: , 2001.
278 .
4. Boris Fumanal, Bruno Chauvel, Franois Bretagnolle. Estimation of Pollen and seed production of common Ragweed in France // Ann Agric Environ Med . - 2007. - 14. P. 233-236.
5. Chauvel B., Dessaint F., Cardinal-Legrand C. and Bretagnolle F. (2006) The historical
spread of Ambrosia artemisiifolia L. in France from herbarium records. // Journal of Biogeography.
2006. - 33 (4). P. 665673.
6. Christian Bohren. Ambrosia artemisiifolia L. in Switzerland: concerted action to prevent
further spreading. Station fdrale de recherches en production vgtale, Agroscope (ACW), Nyon
Switzerland,
2007.
[Electronic
resource].
Mode
of
access:
http://www.ewrs.org/IW/doc/AMBEL_in_CH_concerted_action_Bohren.pdf.
7. Christelle Otto, Beate Alberternst, Frank Klingenstein und Stefan Nawrath. Verbreitung der
Beifublttrigen Ambrosie in Deutschland Problematik und Handlungsoptionen aus Naturschutzsicht
BfN Skripten, Bonn - Bad Godesberg, 2008. - 44 s.
8. Meusel H., Jger E.J. Vergleichende Chorologie der Zentraleuropischen Flora. Text. Band
III. Jena etc.- 1992a. 333 s.
V.K. Tokhtar
J.E. Volobuyeva
Belgorod National
Research University,
308015, Pobeda-str., 85
e-mail: tokhtar@bsu.edu.ru
204
________________________________________________________________
575.22 : 581.163
(FRAGARIA X ANANASSA DUCH.)
.. 1
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1)
,
630090, ,
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e-mail: SO_baturin@mail.ru
,
630090, ,
. , 101
2)
e-mail: ambros@ngs.ru
, . , ,
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205
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2-5
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0,0% ,
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, (V,%).
( %):
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65 84; 85 104; 105
[13].
n1n2
n1 n2
= 2arcsin p
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,
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n2 , ,
, % [14].
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.) 17,6 % [15].
.
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.
206
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1
(1998 2003 .)
()
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96/5-13-2
96/5-13-5
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3.91.7*
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5.81.8*
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88.025.4
104.030.0
113.632.8
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.
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,
.
,
[12].
207
___________________________________________________________________________
. 2.
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208
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I .
, , . , .
-
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,
[12, 17, 18].
, . ,
. Niemirowicz-Szczytt [19] Fragaria ananassa Regina Redgauntlet
() 88,8% 43,4%, . ,
Redgauntlet,
,
, 1-2 . ,
10%, [19].
,
.
Fragaria ananassa Duch. R. Bauer [20].
Mieze Schindler, ,
.
. .. [2] K.Niemirowicz-Szczytt [19]
. ,
,
[4, 5, 21]. ,
- [21, 22].
(
, , ,
.)
, [12].
, ,
,
.
, .
,
, ,
, [22].
1. .., .., .., ..
. . :
, 2004. - 106 .
2. - .. // . -
. - ., 1962. - .26. - . 68-84.
209
___________________________________________________________________________
3. .. . .:
. : , 1976. - . 152-164.
4. .. (Fragaria
x ananassa Duch.) // . . . . . - , 1997. - 16 .
5. .., .., ..
(Fragaria ananassa Duch.) // . - 1994. - .30. -
2. - . 237-243.
6. Galletta G.J., Maas J.L. Strawberry genetics // HortScience. - 1990. - Vol. 25. - 8. - P.
871-879.
7. Scott D.H., Lawrence F.J. Strawberries // Advances in fruit breeding / Ed. J.Janick,
J.N.Moore. Purdue University Press, West Lafayette, Ind., 1975. - P. 71-97.
8. Valleau W.D. Inheritance of flower types and fertility in the strawberry // Americ. J. of Botany. - 1923. - 10. P. 259-274.
9. .. /
. .: , 1975. - . 312-343.
10. .. . - .: , 1988. 272 .
11. .., .. . - .: , 1979. - 192 .
12. Darrow G.M. The strawberry - history, breeding and physiology / Holt, Rinehart and Winston, New York, 1966. - 447 p.
13. .. . - .: , 1990. - 296 .
14. .. //
( , ). .:
, 1964. - . 166-171.
15. .. Fragaria
ananassa Duch. // . . XXI :
. , 2004. . I. . 143.
16. .. . .: - . -, 1975. . 43-49.
17. Erlanson E.W. Sterility in wild rose in some species hybrids // Genetics. 1931. V. 16.
P. 75-96.
18. .., .. Cerasus vulgaris Cerasus tomentosa (ROSACEAE) // . 2006. - . 91. - 2. . 206-218.
19. Niemirowicz-Szczytt K. Strawberry (Fragaria ananassa Duch.) haploids and their generative
progeny. Induction and characteristics. Warszawa. Warsaw Agricultural University Press, 1987. - 71 p.
20. Bauer R. Grundlagen und Metoden der zuchtung bei gartenerdbeere (Fragaria
ananassa Duch.) // Z. Pflanzenzuchtg. - 1960. - Bd 44. - 1. - S. 73-100.
21. .. -
// . - 2001. - 1. - . 39-43.
22. .., ..
(Fragaria x ananassa Duch.) // .
. .-. . - 2010. - 9. - . 36-42.
e-mail: SO_baturin@mail.ru
2)Central
e-mail: ambros@ngs.ru
210
________________________________________________________________
504.54
( )1
..
..
..
..
..
308015, . , . , 85
e-mail: Goleusov@bsu.edu.ru,
tokhtar@bsu.edu.ru
. .
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- 2009-2013 .
211
___________________________________________________________________________
. 1.
. 2.
212
________________________________________________________________
1
10 (), %
, %
, %
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15,33
23,84
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3,04
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56,58
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41,92
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61,46
115,95
46,43
1,45
35,76
46,16
43,43
40,93
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1,58
0,03
0,53
1,04
3,40
1,35
0,05
0,43
0,06
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1,50
1,39
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213
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. 4. 10 20
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214
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. 5. :
(), (),
,
. ,
- .
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, : - (Androsace koso-poljanskii Ovcz.), (Hyssopus cretaceus Dubjan.), (Hedysarum
grandiflorum Pall.), (Matthiola fragrans Bunge) .
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215
___________________________________________________________________________
. 6. :
(),
, -
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1. , ..
/ .. // .
2009. 1. . 37-41.
2. .. ..
// i : i i i (. , 7-9
2008 .). , 2008. . 135-138.
216
________________________________________________________________
3. ..
// .
. - 15 (86). - 2010. - . 12. - . 13-19.
PECULIARITIES OF EDAFOTOP CONDITIONS FORMATION WITHIN THE SITE WITH TEHNOGENNOUSLY CHANGED RELIEF (ON THE EXAMPLE OF BOTANICAL GARDEN BELSU)
P. V. Goleusov
V.K. Tokhtar
E.G. Afanasev
M.N. Jurieva
A.A. Tolkacheva
Belgorod National
Research University
308015, Belgorod, Pobeda-str., 85
e-mail: Goleusov@bsu.edu.ru,
tokhtar@bsu.edu.ru
217
___________________________________________________________________________
635.939.73
..
..
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e-mail: mgau@mich.ru
, . , 7 7 ,
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218
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.
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220
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,
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222
________________________________________________________________
1. , ..
/.. // . ,1992. . 84-88.
2. , ..
/.. , .. , .. , .. / , 6. 2007. - . 10.
3. .. ./ .. // - .: , 1959.-352.
It was detected the influence of planting schemes green cuttings of honeysuckle on the number of rooted cuttings, the total
length increment and root development. Optimal allocation scheme
is 7 x 7 cm, increasing the length of the growth and yield of rooted
cuttings (some varieties of 6 ... 16% more). For such varieties as Goluboe Vereteno and Berel recommended compacted scheme 7 x 5
cm. Increasing the area of supply up to 7 x 10 cm stimulates root
development of rooted cuttings of most studied varieties.
Keywords: honeysuckle, green cuttings, planting scheme,
rooting.
223
___________________________________________________________________________
58.087:581.14
..
..
. .. ,
, 61058, . ,
. , 52
e-mail: khbg@i.ua
Campanula latifolia L.: , (), . ,
: ,
, , .
(, ).
Campanula latifolia - , .
( ), ( , ,
), , .
: , , , , .
. . , , . . , .
, [1-4],
. , , . :
in situ [5,6 .];
ex situ [7 .];
[8 .];
[9 .];
[10 - 12 .].
[13, 14].
ex situ [15,16 .]
.. .
: , , , , ,
.
ex situ 87 , 400 ,
450 .
Campanula latifolia L. Campanulaceae Juss.
224
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, , , , . .
, :
.
.. [17],
.. , .. . [18],
[19 - 21], [22].
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.
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226
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2. : ; ,
; , , ;
, , .
3. Campanula latifolia , .
4. .
5. , , ,
, ex situ
.
6. , .
7. Campanula latifolia
,
.
1. . .1. // . .. . .: , 1985.
317 .
2. .. . .: ,
1967. 122 .
3. .. Campanula L.
. // , 1974, . 59, 3. . 439-447.
4. .. - . ,
1981. 212 .
5. .. . // .
., 1979. . 38-55.
6. .. . //
. , . ., . ZXXIII (4), 1968. . 63-71.
7. .., .. . ex situ . .: , 2001. 192 .
8. ..
Salvia L. ( Lamiaceae). // . , . ., . 89, . 3, 1984. . 100-114.
9. ..
. //
. ., 1974. . 119-131.
10. .. . .: - , 1977. 85 .
11. .. - // . , . ., T. ZXII, (6), 1957. . 35-58.
12. .., ..
. // , ., 1976. . 167-185.
13. .. : . 1. . .: , 1997. 630 .
14. .. : . 2. . .: , 2002. 859 .
15. ..
Gymnospermum odessanum (DC.) Takht. //
. ., 1993. . 171-172.
16. . Saussurea pozcii Degen // . - . . . . , 2007. . 24.
227
___________________________________________________________________________
17. ..
. .: 1983. 55 .
18. .., .., .. . ( ). .: , 1976. 216 .
19. .., .., .. . . .-.: - , 1956. 301 .
20. .., .. . . .: , 1979. 295 .
21. .. . . .,
1990. 204 .
22. .., .., .., .. : . .: 2002. 240 .
23. ... . . .3. 1964. . 146 215.
24. ... . .
1972, .4. . 5 94.
Z.V. Komir
A.A. Alyokhin
Botanic Garden of V.N. Karazin
Kharkiv National University,
Ukraine, 61058, Kharkov,
Klochkovskaya, 52
e-mail: khbg@i.ua
228
________________________________________________________________
581.151
MOMORDICA CHARANTIA L. (CUCURBITACEAE),
.
..
, 308007,
. , . 85
e-mail: tokhtar@bsu.edu.ru
M. harantia,
,
, ,
, . . ,
, .
, . M. charantia
, ,
.
Keywords: Momordica charantia, ,
229
___________________________________________________________________________
.
.
.
300 10 30 /. , [4].
:
Staphyllococcus aureus, Bacillus subtilis Escherichia coli,
Pseudomonas aeruginosa.
[2].
, Momordica charantia
[1], .
(. 1).
1
M. harantia ( %)
(%)
M. charantia M. charantia
26.42 0.49
26.69 0.56
2
M. charantia
Staphyllococcus aureus
Bacillus subtilis
Escherichia coli
Pseudomonas aeruginosa
()
M. charantia M. charantia
21.46 0.52
12.00 0.88
16.37 1.25
16.25 0.94
14.28 0.87
13.62 2.16
15.27 1.12
12.23 0.62
* :
( 2). , . M. charantia
Bacillus subtilis.
Escherichia coli Pseudomonas aeruginosa, Staphyllococcus aureus (.).
M. harantia, ,
, , , . . -
230
________________________________________________________________
, , . ,
. M. charantia ,
.
Momordica charantia
20
16,37 16,25
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15
13,18
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sa
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ru
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M. charantia - ""
M. charantia - ""
.
Momordica charantia
1. , .. , .. . Momordica
charantia Momordica balsamina (Cucurbitaceae) //
. " ". 2010. . 18. . 114- 117.
2. Kivanc M. and B. Kunduhoglu. Antimicrobial activity of fresh plant juice on the growth of
bacteria and yeast // Journal of Qafqaz University. 1997. V. 1. P. 26-35.
3. Nerurkar PV, Lee YK, Linden EH, et al. Lipid lowering effects of Momordica charantia (Bitter
Melon) in HIV-1-protease inhibitor-treated human hepatoma cells, HepG2. // Br. Jour. Pharmacol.
2006. 148(8) P. 1156-64.
4. Roopashree TS, Raman Dang, Shobha Rani RH1, Narendra C. Antibacterial activity of antipsoriatic herbs: Cassia tora, Momordica charantia and Calendula officinalis // Intern. Jour. Applied
Research in Natural Products. 2008. Vol. 1(3). P. 20-28.
231
___________________________________________________________________________
The comparative analysis of Momordica harantia plant samples, grown up in Vietnam and the plants received from seeds of the
same sample under the conditions of the Belgorod region, has shown
that formation and accumulation of biological active substances of the
samples had close values. However ratio of components was changed.
It led to declining of ntibacterial activity in comparison with the sample received from Vietnam. Successful introduction of plants M. charantia in the Belgorod region for the purpose of their use as perspective
sources of biologically active substances demands special, effective,
new agrotechnical ways of cultivation of these plants.
Keywords: Momordica charantia, introduction, antibacterial
activity.
232
________________________________________________________________
581.2; 581.16; 631.8
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. , 85
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e-mail: kurkina@bsu.edu.ru
, Hypericum pertoratum L., Tagetes patula L., Orignum vulgre L., Calendula officinalis L., Matricaria recutita L. Echinacea purpurea Moench. .
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233
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28
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13
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234
________________________________________________________________
, 80100 %
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48
38
52
20
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26
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,
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Hypericum. , , H. perforatum
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, .
236
________________________________________________________________
8. .., ..
Alternaria Nenses. // 2010:
. . (. , 25 2010 .). : . , 2010. . 176178.
9. .. // . 2011. 2. . 25.
237
___________________________________________________________________________
581.151
Momordica L. .
,
, ,
- .
Keywords:
.
Momordica,
Momordica L. (Cucurbitaceae) ,
. ,
, , , [1].
, [3]. , [4]. ,
.
,
Momordica , . -, . , , , . . , - . , , , . ,
(), . . (. 1,
2) [2]. Momordica , . ,
. . , , : 3
, 5 .
Momordica charantia L. Momordica balsamina L.
().
Momordica charantia , . Momordica balsamina, , Berlin-Dahlem (, . ).
238
________________________________________________________________
. 1. Momordica charantia L.
. 2. Momordica balsamina L.
239
___________________________________________________________________________
13 1-1,5
(22-25), .
. 17 ,
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8 .
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, ,
. [5].
( , )
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1
(2010 .)
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, .
,
4
9.8
11
58
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5
17.4
22
61
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6
22.3
28
52
20.3
7
25.3
30
54
41.4
8
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29
44
24.8
9
14.7
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10
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5
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35
30
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10
5
0
4
10
, .
. , .
. 3.
Momordica charantia L. Momordica balsamina L.
240
________________________________________________________________
,
15 . 6- . 10- Momordica balsamina
, Momordica charantia
13- .
1
Momordica L.
M. charantia
M. balsamina
6
6
13
56
69
10
50
84
81
91
. . . . . 28 .
Momordica
. , . , .
, .
Momordica charantia ( 10-12) . Momordica balsamina ( 3-4,5 ) . M. charantia M. balsamina
59.5 () 13.4 (). M. balsamina
26 , M. charantia 23 .
Momordica
. M. charantia M. balsamina 7.2 / 4.1 / .
, -
(0.40.5 /).
,
,
,
.
.
1. .. . .
-, 1989. 149 .
2. Abascal K., Yarnell E. (2008). Momordica charantia (Bitter Melon). Botanical Medicine, Integrative Medicine. 7(1): 21-24.
241
___________________________________________________________________________
3. Joseph J.K. Studies on ecogeography and genetic diversity of the genus Momordica L. in India. National Bureau of Plant Genetic Resourse. Thrissur, Kerala: 10 11. 2004.
4. Lee-Huang S, Huang PL, Chen HC, et al. Anti-HIV and anti-tumor activities of recombinant
MAP30 from bitter melon. Gene. 1995 Aug 19;161(2):151-6.
5. Roopashree TS, Raman Dang, Shobha Rani RH, Narendra C. Antibacterial activity of antipsoriatic herbs: Cassia tora, Momordica charantia and Calendula officinalis. International journal of applied research in natural products. Vol. 1(3), pp. 20-28, Sep/Oct 2008.
Peculiarities of the Momordica patterns cultivation are analyzed under the conditions of the Belgorod region in the paper. In
spite of the fact that the studied plants are adapted to warm and wet
conditions of tropical and subtropical climate, they were successfully grew in the botanical garden of the Belgorod State University that
allows to consider the plants as perspective untraditional species for
cultivation in the Middle Black Earth zone.
e-mail: tokhtar@bsu.edu.ru
zone.
242
________________________________________________________________
581.95(470.325)
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e-mail: tokhtar@bsu.edu.ru
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243
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Asteraceae
Poaceae
Apiaceae
Fabaceae
Rosaceae
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Chenopodiaceae
Grossulariaceae
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Cannabaceae
Moraceae
Polygonaceae
Amarantaceae
Potulacaceae
Papaveraceae
Brassicaceae
Aceraceae
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:
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4
4
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3
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244
________________________________________________________________
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(42,0 %), (22,0 %) (12,0 %).
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245
___________________________________________________________________________
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246
________________________________________________________________
, ,
, . , , ,
, , , . -, ,
[3].
1. .. . : , 1991. -168 .
2. .., .. : // .
2008. . 50-54.
3. .., .. . //
. . 2010. 21 (92). - . 13. . 20-23.
Study of alien floras formed within territorial borders of Graivoronskiy and Rovenskiy districts of the Belgorod region are presented.
Use of traditional methods of comparative floristics and a method of correspondence analysis of canonical correlations allowed to visualize interrelations among structures of florae, defining degree of their similarity
and distinction and the reasons of this similarity.
Keywords: flora, alien species, methods of multidimensional statistics.
247
___________________________________________________________________________
581.4:581:581.9
CONYZA CANADENSIS (L.) CRONQ.
- 1
..
..
, 308015,
. , . , 85
e-mail: tokhtar@bsu.edu.ru
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_______________________________________________________________
250
250
. 2011. 9 (104). 15/1
251
___________________________________________________________________________
, 23
Conyza canadensis (L.) Cronq.
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.
1. .I. .., .. ii i .: .., 1997.- 94 .
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3. .. Conyza Canadensis (L.) Cronq // . 2000. . 1. . 126 -129.
252
________________________________________________________________
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CONYZA CANADENSIS (L.) CRONQ.1
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e-mail: tokhtar@bsu.edu.ru
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253
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-0,27
-0,21
0,09
0,25
0,46
-0,33
0,00
0,14
0,13
0,01
1,00
-0,12
-0,02
0,07
-0,31
-0,42
0,16
0,08
-0,07
0,24
-0,47
-0,41
-0,42
0,29
-0,12
1,00
-0,04
-0,03
0,02
-0,24
-0,17
-0,16
-0,18
-0,13
0,23
0,18
0,13
0,03
-0,02
-0,04
1,00
-0,25
0,20
-0,04
0,29
-0,33
0,23
-0,31
-0,02
-0,07
0,41
-0,48
0,07
-0,03
-0,25
1,00
0,10
-0,16
-0,16
-0,27
-0,12
0,05
0,40
0,04
0,31
-0,27
-0,31
0,02
0,20
0,10
1,00
0,69
0,21
-0,12
-0,22
0,36
0,33
0,12
0,31
-0,21
-0,42
-0,24
-0,04
-0,16
0,69
1,00
0,46
-0,21
0,01
-0,19
0,04
-0,07
0,33
0,09
0,16
-0,17
0,29
-0,16
0,21
0,46
1,00
-0,32
0,04
-0,03
-0,02
-0,10
-0,46
0,25
0,08
-0,16
-0,33
-0,27
-0,12
-0,21
-0,32
1,00
0,09
0,02
-0,48
-0,16
-0,48
0,14
0,12
0,14
-0,16
0,03
-0,38
-0,28
0,04
0,08
0,22
-0,35
0,28
0,16
1,00
-0,42
0,13
-0,43
0,13
0,41
0,30
0,30
0,33
-0,46
0,13
-0,02
0,07
0,27
0,29
0,19
0,22
-0,14
0,30
-0,10
0,13
0,11
0,26
-0,12
P
Q
R
0,09
0,22
0,13
0,02
-0,35
-0,02
-0,48
0,28
0,07
-0,16
0,16
0,27
-0,48
1,00
0,29
0,14
-0,42
0,19
0,12
0,13
0,22
0,14
-0,43
-0,14
-0,16
0,13
0,30
0,03
0,41
-0,10
-0,38
0,30
0,13
-0,28
0,30
0,11
0,04
0,33
0,26
0,08
-0,46
-0,12
1,00
-0,48
-0,26
-0,48
1,00
0,29
-0,26
0,29
1,00
2.
Conyza canadensis
/
1
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
5
0,00
0,05
0,04
0,05
0,09
0,00
0,04
0,06
0,10
0,14
0,25
..
..
.
.. .
.. .
.. .
..
..
0,00
0,03
0,03
0,03
0,08
0,10
0,11
0,11
0,38
0,49
0,11
0,57
255
___________________________________________________________________________
:
23 17 9 11 7 2 14 5 10 8 19 22
21 15 3 20 18 1 12 6 13 4 16
, Conyza canadensis , .
, ,
(.2). , , , , , , . , -
. , -,
Conyza canadensis
, ,
(, 1999).
, , Conyza canadensis - , , .
, , Conyza canadensis , , (, 1999).
Factor Loadings, Factor 1 vs. Factor 2
Rotation: Unrotated
Extraction: Principal components
0,8
0,6
0,4
Factor 2
0,2
0,0
-0,2
-0,4
-0,6
-0,7
-0,6
-0,5
-0,4
-0,3
-0,2
-0,1
0,0
0,1
Factor 1
256
________________________________________________________________
1. . . . 1. Leucanthemum vulgare (Asteraceae) // 11.
1999 . .50-66.
V.K. Tokhtar
N.V. Mazur
Belgorod National
Research University, 308015,
Belgorod, Pobeda-str., 85
e-mail: tokhtar@bsu.edu.ru
257
___________________________________________________________________________
635.9.582.973
..
, 692533, ,
, . , 26
e-mail: malyshsveta@rambler.ru
(
). , , , .
: , , , , .
Lonicera L. , .
1935 40
. , , 1965 .
.
, ,
, , , , , .
.
, [1].
. L. alpigena L. :
.
1-2 . 11-
60 . , 5-9 34 . -, . -, . -, . . 1999 . 187
. . III. II.
.
. , , .
. L. caucasica Pall. : ;
, .
2 . , , 4-8
2-5 . -, , 12 .
, , . .
7-8 ; 18 . 1965 . 168 . . II. I-II.
. , , .
258
________________________________________________________________
. L. chrysantha Turcz. : ,
(-), - , .
2 . , ,
, 6-12 3-5 . -,
. , , .
, - . 1964 . 171 . . I. I.
, .
. L. edulis Turcz. ex Freyn. : ,
, , .
1-1.5 , . , , 1.5-5.5 1.5-2 . -. , . , , .
. 1967 . 176 . . I. I.
, , .
. .
. - L. glaucescens Rydb. : .
. .
, 1.5 . , , 4-8 3-5 ,
, -, . -
-, , 2 . . ;
. 2000 . 188 .
. I. I.
.
. L. Ledebourii Esch. : ().
1.5 , . 7 50 . , 6-10 , , -,
, 3-5 . , ,
- -, 1.5-2 . , . 1999 . 7- . 189 . . II. II-III.
.
. L. maackii xim. : , -; . ; - ; . . , , .
3-4 (5) , , . - .
, , 2-, , , ;
2 . , , -, 6-6.5 , 1-9 . - ; -. 1964 . 178 . . I. I.
- , : , - .
. L. maximowiczii (Rupr.) Regel. : , , .
3 . , -.
, , , 3-7
259
___________________________________________________________________________
2-4 . - , . -, . - , . 1962 . 179 . . I. I-II.
, .
. L. morrowii A. Gray. : : -
, .
2 , .
-, , , 2-4 1.5-3 . , , 1-1.5 . -. , -.
1999 . 198 . .
II. II.
, , .
. L. nigra L. : .
70 . ; -, , 4-6 2-3 .
-, -, 10 . -, , . 1999 . 4 . 190 . . I. I.
, , . .
. L. olgae Reg. et Schmalh. : , -. 2600 .
, 0.3-0.5 .
-, , , 1.5-3
0.8-1.5 . -, , .
. 2000 . 180
. . I. I.
, .
. - L. praeflorens Batal. : ( ), , .
2 , , , . , , 4-6
3-5 . . , , . , , - . 1962
. 190 . . I.
I.
.
.
. L. prolifera Rehd. :
.
1.5 . , , 5-8 3-5.5 , , -, . -, 2; 2.5-3 . , 8-9 . ; - . 1999 . 190 .
. I. I.
260
________________________________________________________________
, .
. L. ruprechtiana Regel. : ,
(-), - .
3-4 . - , , 6-8 3-4 . -
. , , , . , . 1964 . 187 . . I. I.
. .
. L. tatarica L. :
, , -
.
2-2.5 . , , 3-6 1.5-3 . -, , 2 . , . - ; . 1963 . 177 . . I. I.
, ; .
. L. vesicaria Kom. : .
1-2 . . , , , 5-10 3-4 . , , ;
2-3 . -, . , - . 2000 . 190 . . I. I-II.
(,
).
. L.xylosteum L. : .
, . , , .
1-3 . 7- 80 .
, , 3-6
2-4 . -, , 1-2
. -. , . 1999 .
190 . . III.
II.
. .
1. / . .. . .: ,
1973. 263 .
In article results of introduction of honeysuckles in the conditions of southern Primorski region (arboretum of GTS FEB RAS) are
resulted. Duration of the vegetative period, flowering and fructification terms, winter hardiness degree, perspectivity of introduction of
the investigated species of a honeysuckle are defined.
Keywords: introduction, honeysuckle, vegetation, winter hardiness, perspectivity.
261
___________________________________________________________________________
581.522.4 + 582.681.16 -13
HELIANTHTMUM MILL
.. 1)
.. 2)
.. 1)
1)
,
620144, . ,
8- 202
e-mail: minogina71@mail.ru
lidia.semkina@botgard.uran.ru
2)
, 620083,
. ,
. ,48
H.nummularium
H.baschkirorum .
. , .
.
: , , ,
, .
-mail: zimn@list.ru
- : , , , - , ,
. .
. , ; . ,
, . ,
, . [4].
. ,
,
. ,
, Helianthtmum nummularium (L) Mill
Helianthtmum basckirorum(Juz. ex Kupatadze) Tzvel ,
[5] [6].
, 20-30 , , ,
. , , 5-30 , -, ,
, -, - .
- - 5-7 , . 1,5-2,3
, 3-8 , ,
262
________________________________________________________________
, ,
, ( .helios anthos
). .
. - , , (, , ), , .
.
. [2]. 1 2
42 , , [10]. .
-9.
( LEICA DM 5000 B). , , ,
, .
, IKI [9].
[8].
.[1]
Helianthemum nummularium H. baschkirorum, . .
.
Helianthemum nummularium. ,
3- . : , .
. [11]. . ; . ,
3 . ( ); . ;
. . ( );
. ; ; , . , ;
. [.1].
elianthemum baschkirorum. , 2- . , . ,
. , . ; . ; .
( .); . [.2].
.
- , ;
, .
. 20-40 -
263
___________________________________________________________________________
. baschkirorum , 100 , ,
H. nummularium, 50 baschkirorum: , - . baschkirorum , .nummularium, . baschkirorum,
[7]. ,
.. . ,
: , [3]. , .
, , , . . , 30-40 .
, . 40-50 , . ,
,
. . .
90%. , ,
. ,
[.3.a, b ]. ,
,
. , , .
, (
).
,
( ).
. .
1 , . .
, .
.
, [3.c.] - [.3.d.].
- 12-15, 30 - 35.
. . , ( 3-5 ).
, .
. ( ) , . .
Polygonum- .
264
________________________________________________________________
265
___________________________________________________________________________
. 2. Helianthemum .
H. nummularium
H. baschkirorum
266
________________________________________________________________
. , . . . , , . ,
.
(15%), (4%). , .
. . , . 25%, ,
, -
, , ,
, , . (
60%). ( ), , , . ,
, -,
.
,
. .
( 70%). , , 50% , - - .
267
___________________________________________________________________________
.nummularium H. baschkirorum , .
, , ,
. , .
. ,
, . , ,
, . -,
, ( )
.
, . ,
.
1. .. . ( ). - .: 1982. - 80 .
2. .. . .: - ,1973. 384.
3.
..
.
1999.
http://www.ggause.com/ titpagru.htm.
4. .. .//
, . .- 1969. .66. . 151-154.
5. . / .. , .. , .. . : , 2001.- .I .207.
6. ( ): . / . ..
.. . - : - -,1996. .122.
7. .., .., .. Helianthemum . //
, 60 . : , 2006. . 190 -193.
8. .., .., .., .. - . . .: 1988.- .4-9.
9. .. . .: ,1980. 304.
10. ./ . .., .. .: - . 1964. .III 530c.
11. .. ( Cistaceae Lindl). // . -.: .- 1949. .XV .327-347.
e-mail: minogina71@mail.ru,
lidia.semkina@botgard.uran.ru
2) Ekaterinburg,
e-mail: zimn@list.ru
268
________________________________________________________________
632.51:633.85
..
, 634050,
. , . , 36.
e-mail: mikhailova.si@yandex.ru
:
Brassica napus L. Metzg., Brassica rapa L. ssp. oleifera
campestris Camelina sativa Crantz. , ,
. ,
, .
() , . (. )
[1]. ,
, , .
, , , , . ,
, .
. , , .
( , ), 20022008 .
. 50
. , .
1
2 , .
,
. , , :
, , [2].
269
___________________________________________________________________________
.
,
. 1,6 8,4%. , : (, , )
. ( 34 ) -
( ) ( ) . ,
, .
, .
Avena fatua L ,
Elytrigia repens (L.) Nevski
Echinochloa crusgalli (L.) Beauv. ,
Panicum miliaceum ssp. ruderale (Kitag.) Tzvelev
Setaria viridis (L.) Beauv. s.str.
Cannabis sativa L.
Fallopia convolvulus (L.) A. Lve
Persicaria lapathifolia (L.) S.F.Gray
Polygonum aviculare L. ,
Rumex acetosella L. ,
Chenopodium album L.
Amaranthus retroflexus L.
Elysanthe noctiflora (L.) Rupr. ( )
Spergula arvensis L.
Stellaria media (L.) Villars ,
Brassica campestris L.
Erysimum cheiranthoides L.
Neslia paniculata (L.) Desv.
Raphanus raphanistrum L.
Sinapis arvensis L.
Thlaspi arvense L.
Melilotus albus Medicus
Trifolium pratense L.
Vicia hirsuta (L.) S.F.Gray
Erodium cicutarium (L.) Her.
Viola arvensis Murray
Phacelia tanacetifolia
Lappula squarrossa (Retz.) Dumort.
Galeopsis bifida Boenn. ,
Galeopsis ladanum L.
Stachys annua (L.) L.
Stachys palustris L.
Plantago media L.
Galium aparine L.
Arctium tomentosum Mill.
Centaurea cyanus L.
Cirsium setosum (Willd.) Bess.
270
________________________________________________________________
Lapsana communis L.
Matricaria perforata Merat.
Sonchus arvensis L
40
. , ( )
. ,
, , , , .
, . 13 : , ,
, , , , ,
, , ,
, , (.).
, [3].
(%)
Camelina sativa ( .)
1
2
3
4
5
6
7
8
9
10
11
12
13
Echinochloa crusgalli
Setaria viridis (L.)
Persicaria lapathifolia
Chenopodium album
Amaranthus retroflexus
Elysanthe noctiflora
Spergula arvensis
Brassica campestris
Thlaspi arvense
Erodium cicutarium
Galeopsis bifida
Cirsium setosum
Sonchus arvensis
2002 .
100,0
71,4
85,6
100,0
100,0
100,0
85,6
14,3
85,6
57,1
85,6
100,0
71.4
2003 .
71,4
100,0
57,1
100,0
85,7
85,6
85,6
14,3
100,0
57,1
85,6
85,6
57,1
2004 .
50,0
75,0
87,5
87,5
100,0
100,0
37,5
12,5
100,0
75,0
37,5
100,0
87,5
2005 .
100,0
100,0
100,0
100,0
100,0
75,0
100,0
50,0
100,0
100.0
50,0
100,0
50,0
. , . , (. .), :
, , , ,
, , , , .,
, , . , (. ),
( , , ,
, , , , , ).
, ,
, , ,
. 40 , 13 .
, , . -
271
___________________________________________________________________________
, .
.
, .
1. .. . .: , 1987. 160 .
2. .., .. . .
2-, . . .: , 1978. 288 .
3. .., .., .., .., .. : // . .
. . , 2007. . 231234.
S.I. MIKHAILOVA
Siberian Botanical Garden
Tomsk State University,
pr. Lenina 36, Tomsk, 634050,
Russia
e-mail: mikhailova.si@yandex.ru
272
________________________________________________________________
630*181.8: 582. 717.4 (470. 57-25)
HYDRANGEA
DUMORT.
..
- , . ,
. 195, . 3, 450080
e-mail: murzabulatova@yandex.ru
6 Hydrangea Dumort., .
.
.
: , , , -
, - .
,
.
- [5, 8, 9].
[9],
. Hydrangea Dumort., . ( ),
,
.
Hydrangea Dumort. - . , 11 .
6 , : Hydrangea arborescens
L. (), H. bretschneideri Dipp. (), H. heteromalla D. Don. (), H. cinerea Small. (), H. paniculata Siebold (), H. radiata Walt. ().
.
[6], 3-
6 (2005-2010 .). : , , , , , , , , .
.. [1].
, . , (. 1).
. , -
273
___________________________________________________________________________
, ,
.
(. )
H. arborescens
H. bretschneideri
H. heteromalla
H. cinerea
H. paniculata
H. radiata
(3,0)*
(1,19)
(3,0)
(0,58)
(3,0)
(0,54)
(2,0)
(1,04)
(10,0)
(1,38)
(2,5)
(0,86)
()
()
( )
* (3,0) - (, )
Hydrangea Dumort
H. arborescens
H. bretschneideri
H. heteromalla
H. cinerea
H. paniculata
H. radiata
27.0410,28
7.050,88
12.0720,26
21.1017,50
21.097,22
18.102,98
29.0410,58
9.0510,28
1.077,86
28.089,04
22.094,20
12.1012,48
28.0410,58
10.0510,46
3.011,50
28.089,04
20.093,10
12.1012,96
1.056,22
30.0413,46
1.054,16
29.040,63
11.056,88
10.058,42
10.056,86
10.050,56
23.078,36
4.074,44
30.0713,32
12.074,78
16.1015,48
2.0929,48
30.0915.48
21.0910.24
28.0911,24
28.098,52
28.0912,08
24.091,50
27.109,20
18.106,62
24.105,70
18.102,44
- ,
.
. H. bretschneideri H. heteromalla (21 22 ) (. 3). H. radiata (14 ). . 2 . ,
. H. paniculata ,
[10].
- -, - [4].
(<0,001),
.
34 (. . 3). H. bretschneideri H. heteromalla ( 13,7 ), H. radiata
H. cinerea (53,0 71,8 ).
.
274
________________________________________________________________
[2] , H. bretschneideri H. cinerea
, (H. arborescens, H. heteromalla) , . ,
(, ) [7], 2 , , 10-12 .
3
H. bretschneideri
H. heteromalla
H. paniculata
H. cinerea
H. arborescens
H. radiata
, 1
113,32,90
I
114,02,30
I
119,42,50
I
129,28,03
II
131,06,04
III
135,83,19
III
,
13,71,20
13,01,73
29,66,21
71,810,48
23,36,69
53,018,17
20 15 . (
2,5 ),
.
1
30 ( 12 ). H.
bretschneideri (1 ), H. heteromalla (3 ) H. paniculata (5 ). H. radiata (30 ).
28
16 , 19 . H. bretschneideri H. heteromalla (28 ), H. cinerea (16 ).
23 .
H. paniculata (30 ), H. bretschneideri H. heteromalla (14 ),
H. radiata (30 ).
.
4 30 , 20 .
H. heteromalla (4 ), H. paniculata (27 ) H. cinerea (30 ).
.
. H. heteromalla H. arborescens (20 21 ). H. cinerea, H. paniculata, H. radiata (28
).
. 12 27 , 18 . H. bretschneideri H. heteromalla (12 ),
H. cinerea (27 )
(. 4)
275
___________________________________________________________________________
4
Hydrangea Dumort
H. arborescens
H. bretschneideri
H. heteromalla
H. cinerea
H. paniculata
H. radiata
27.04-23.10
29.04-17.10
28.04-12.10
1.05-29.10
30.04-27.10
1.05-14.10
,
174,56,00
166,210,35
167,010,40
178,65,64
174,47,16
176,14,48
. - 173 .
[1]
(. 5).
Hydrangea Dumort .
H. arborescens
H. bretschneideri
H. heteromalla
H. cinerea
H. paniculata
H. radiata
0,088
-0,742
-0,789
0,887
-0,038
0,690
[: , 1981]
5
4
4
5
4
5
,
(H. bretschneideri, H. heteromalla, H. paniculata). 4, . . () . 3 (H. arborescens, H. cinerea, H. radiata) 5
() .
, , , .
6 . .
. .
( 2,5 ), .
.
1. . . . .: , 1981. 119 .
2. .. . ., ,
1975. 547 .
276
________________________________________________________________
3. .. .
: . -, 1982. 128 .
4. .. . // .
. . . 2010. 2. 38-39.
5. .. . :
, 1976. 390 .
6. . ., 1975. 28.
7. .. // . . . . 1978. . 109.
50-54.
8. .. 5. Hydrangea L. // . .,
.: - , 1954. . 3. . 162-173.
9. .. . :
, 2007. . 85-87.
10. .. , . : . . -, 1984. 244 .
Six species of Hydrangea Dumort. introduced in Bashkir CisUrals are studied.. All the species are stacked by their phonological
cycle into vegetation period of the introduction region.
Key words: Hydrangea, species, phenology, flowering.
277
___________________________________________________________________________
582. 677.1
..
..
-
,
. ,
. , 142; 690024
e-mail: Lubavar1188@mail.ru
M. sieboldii K.
Koch. . M. sieboldii ,
0-40 20 .
.
39 52%.
: Magnolia sieboldii, ,
, .
278
________________________________________________________________
Magnolia sieboldii K. Koch. .
Magnolia sieboldii - 8 .
. , -. . , 6-15 , -, ,
-, . 1-2 . , . , 7-10 , , ,
3-6 , ; , , , , . , .
,
() [3, 4].
Magnolia sieboldii,
.. [5].
[6],
Magnolia L.,
0-50. Magnolia sieboldii
( ). 0-40 +16-200, 20 . .
. . [7]. () , ()
, , .
- , . (),
.
-
70 M. sieboldii
. M. sieboldii
9- . 2 , (
, ), . 3-
.
11 . M. sieboldii, ,
, 7-8 . ,
.
25 29 .
, M. sieboldii
. 1 21 .
. 29 7 (.
1). 161 .
M. sieboldii, 13 . 36 . M. sieboldii,
,
279
___________________________________________________________________________
[8]. M. sieboldii ,
.
1
31.05
18.06
13.06
10.06
30.06
29.06
07.07
28.07
18.07
12.05
13.06
2.06
07.05
25.05
16.05
30.04
06.06
09.05
19.04
30.05
29.04
Min
Max
24.09
08.10
03.10
18.10
29.10
23.10
05.10
10.10
07.10
, , .
. :
,
. ,
.
M. sieboldii, , , . , 34-48 . , , , 18-19 1 . , 39-52%.
5,4 1,4 .
: 4,2 0,14 1,97 0,04
(. 1). 0,5 0,02 0,48 0,02 .
, , , [6,8].
, ( ) 60%,
80%.
77% .
,
. 1.
- 55%.
Magnolia sieboldii
,
K. Koch
- 50%, - 26%.
10 ,
. , ,
33 , . ,
. ,
0-40 20 (69-90%), (50%) +16-200 [8].
280
________________________________________________________________
M. sieboldii
, . M. sieboldii
0-40 20 . .
39 52%. M.
sieboldii , .
1. .. // . . . .
2: . : , 2000. . 218-225.
2. .., .. . : . , 1987. 184 .
3. .., .., .. Magnoliaphyta. - .:
, 2009. 400 .
4. .. // . . ., 1995. . 128-130.
5. .. // . . 1967. . 65. . 13-18.
6. .., .., .. . .: , 1985. .198.
7. .. // . , 1974. . 59, 6. . 826-831.
8. .. . :
, 2003. 100 .
281
___________________________________________________________________________
630.62
1
.. 1
.. 2
1)
. .. , 127276,
, ., 4
e-mail: ser-rysin@yandex.ru
, 690041,
, . , 7
2)
,
(). .
.
: ,
e-mail: elepeshkin@amur.wwf.ru
,
.
, . ,
. ,
,
.
.
,
. , , , .
, , ,
. ,
,
, . , , , , 1
: . : .
282
________________________________________________________________
, ( )
[2].
, . , .
, [1].
, ,
[3, 4], 29
, : , . ( 0 4 ). ,
. ( 0 1)
.
.
(). I
;
II , ;
III ;
IV , .
,
.
,
. ,
.
,
[4]. . ; ,
[5].
, ,
, , . , ( )
().
283
___________________________________________________________________________
() , . 1983 . .
11,6 . ; 80% .
38%, 42%,
16% , . . - 1 . . . - (,
, ), . ,
, .
( 290 ),
. . , 1950- . ..
.. . ,
, ,
, . ,
, ;
. ,
[5].
(Torup) ( ),
8 . - .
955 , 360 , 38%
. , , . , . , ,
- (Bokskogen), .
(, , ),
. ,
.
, . ,
, , ,
.
, , ,
. . , .
, , , , . , , , . ; .
.
284
________________________________________________________________
1.
(1-4 , =3,25),
,
(0-4 ). -
2,69. ; 11 25 (=3,60).
, - .
: ,
5 (=1,71), (=1,08). (=3,61) , ,
(=2,56).
, ( 2,36 2,16), , ,
, , . , , . (2,16 ),
(0), (4).
: , , ( ),
. ,
. .
, . ,
(V ) - , , .
, ..
( 2,50 2,77).
- 1 ( 7% , ) 4
(15% , ); 2,70. ,
- 90%
(3-4 , =3,32). , . ; (3-4 ) 90% ,
3,46. 60% ,
(2,56).
1
() ().
285
___________________________________________________________________________
- 0 ( 5 ) 4 ( 50 ).
, 45%
(=1,84),
,
(=2,33).
,
.
, .
, , ;
1-4 . ( );
.
, ( ) - 0,61, .
, .
, . , (
), . , , . ,
, , . , ,
.
.
. (=3,99). ,
,
( 0 4; =3,45). , ,
,
(=2,69, =2,94).
- , 30% , .
( , ) ,
(=1,03).
: , - ,
, , , (=0,96).
. -
286
________________________________________________________________
. ,
, (=3,97).
, (4 )
( 40% ),
, (=3,34). ,
;
.
4 (=2,87).
,
(=3,00).
,
: , - ,
. , -.
, (=1,58).
(-) , ( =2,04). ,
,
, , .
(=2,13, =2,08); , . (80% 90% )
. ,
, ,
.
, (3,92 3,93). - - (), . (=0,70; =0,66)
.
,
, ,
- . ,
, .
. (0 1
)
, ; (3 4 ) ,
, , .
.
, ( )
(=3,26). -
287
___________________________________________________________________________
, , ;
, ( ) =2,78.
1/3
(4 ); ,
, . 4% .
,
.
,
(=1,34). (60%)
1 , 15% (, )
(=1,19).
( 0,29
0,50);
( 0,09 0,05 ).
, , , ( 3 );
(
).
,
,
. , , . , , 60 . .
, . (=0,95) , 1/3
- .
(=1,34), ,
. , ,
.
,
, (=0,51;
=0,49)
.
. ,
, (, )
. -
288
________________________________________________________________
, ,
. ,
.
IV ,
, . 5%
, 14%.
. , , ,
- . , ,
, .
, , , .
. , ,
.
, .
. 1) , , .
2) , . :
( , );
,
;
- ;
(
. . , );
,
( - , .).
. 1) , ,
.
, .
2)
,
( , ).
3)
. -
,
.
289
___________________________________________________________________________
. , ,
.
1. : . /
. .. , .. . .: , 2004., - 235 .
2. . , 1997. 40 .
3. , .. / .. // , 2003, - 1. . 17-27.
4. , .. / .. // . .: , 2006. . 143164.
5. , .. / .. , .. // . . .: , 2008. . 185-208.
e-mail: ser-rysin@yandex.ru
Institution of Russian academy
of sciences Pacific Institute
of Geography, Far Eastern Branch
of RAS; Radio str., 7, Vladivostok,
690041, Russia
2)
e-mail: elepeshkin@amur.wwf.ru
290
________________________________________________________________
574.21
..
. .. ,
127276, ,
., 4
e-mail: ser-rysin@yandex.ru
.
: , -,
,
.
- (. .
) , .
, : , , , , . - ( , , .), . -
, , . [1].
A. ( ) , . . , .
,
. , , .
B. (, ,
, .), .
: , .
C. , .
, .
D. , ( ).
, :
;
.
, -
291
___________________________________________________________________________
(, , , ,
, , ).
.. [2] :
- : , , ;
- :
; ;
- :
, ; , .
(A, B, C, D) 5- (. 1).
1
( )
,
,
,
0
1
2
3
4
() , (1):
C
16
D
,
(1)
A, B, C, D
.
(. 2).
2
I
II
III
IV
V
0 0,20
0,21 0,40
0,41 0,60
0.61 0,80
0,81 1,00
I , , . .
, II, . - .
, , .
III ,
. -
292
________________________________________________________________
. , , , .
IV ,
.
. , , .
V
. , , .
,
.
(a, b, c), . , . 3.
3
I
II
III
IV
a
b
c
0,06 0,13 0,19
0,25 0,31 0,38
0,44 0,50 0,56
0,63 0,69 0,75
250 ,
[1]. ,
( . 4.)
, .. [3]. (2):
kx
,
k
(2)
N (A , B , C , D
); ; k
.
k
- (. 5).
293
___________________________________________________________________________
4
()
Achillea millefolium L.
7
8
Aconitum
septentrionale Koelle
Actaea erythrocarpa
Fisch.
Actaea spicata L.
Adoxa moschatellina L.
Aegopodium
podagraria L.
Agrimonia pilosa Ldb.
Agrostis canina L.
10
11
12
14
15
Alchemilla vulgaris L.
16
Allium ursinum L.
17
Alopecurus pratensis L.
Amoria montana (L.)
Sojak
Amoria repens (L.)
C.Presl
2
3
4
5
6
13
18
19
2
0
21
22
23
24
Anemone sylvestris L.
Anemonoides
nemorosa (L.) Holub
Anemonoides
ranunculoides (L.)
Holub
Angelica sylvestris L.
Antennaria dioica (L.)
Gaertn.
()
0,06
1a
0,75
4c
0,50
3b
4
4
4
4
0
0
0
0
0,50
0,50
3b
3b
0,31
2b
4
2
3
2
0
1
0
0
0,44
0,31
3a
2b
0,13
1b
2
2
2
2
2
2
0
0
0
0
0
0
0,25
0,25
0,25
2a
2a
2a
0,19
1c
0,50
3b
0,13
1b
0,94
5b
0,25
2a
0,31
2b
0,13
1b
0,75
4c
0,63
4a
0,56
3c
0,50
3b
0,38
2c
5
75% ( )
50-75%
25-50%
5-25%
5%
( )
-
5
4
3
2
1
r+
5
4
3
2
1
0,1
(. 6).
294
________________________________________________________________
6
()
0 0,80
0,81 1,60
1,61 2,40
2,41 3,20
3,21 4,00
(),
:
0,80,
I ;
0,81
1,60, 0,80,
II ;
1,61
2,40, 2,40, III ;
2,40,
, IV ;
3,21, V .
. .
,
. ,
-
, . 9
(. 7).
, ,
1957 .
. ,
( ) B
( ).
C ( ) D ( ). , ( , V IV ).
7
()
1957
1990
1995
2003
A
B
C
D
3,36
2,93
0,24
0,14
3,45
3,20
0,10
0,07
3,42
2,66
0,13
0,10
2,77
2,68
0,12
0,09
295
___________________________________________________________________________
, . , .
1. . // .., .., .. . : , 2004. 302 .
2. , .. / .. // . . / . . 1950. . 3: . . 6
..: . 7204.
3. , ..
/ .. / .: , 1983. 197 .
296
________________________________________________________________
582.973(470.13-924.82)
LONICERA L.
..
..
..
, 167982, . ,
. , 28
e-mail: mryabinina@ib.komisc.ru
48 , . , - .
: , Lonicera, , , .
297
___________________________________________________________________________
,2003). 2003 . 11 (, 1994
.). 19 2 . 26 .
, ,
(, 1980), 3 , 9 Lonicera ( ) 2 Caprifolium (
).
Spinosae
Caeruleae
Vesicariae
Distegiae
Adenostegiae
Alpigenae
Rhodanthae
Tataricae
Ochranthae
,
Lonicera
Isoxylosteum
L. alberti Regel
Isika
L. altaica Pall., L. caerulea L., L. edulis
Turcz. ex Freyn, L. pallasii Ledeb.
L. ferdinandi Franch., L. vesicaria Komar.
L. involucrata (Richards.) Banks ex Spreng.
L. tolmatchevii Pojark.
L. alpigena L.
L. maximowiczii (Rupr.) Regel, L. nigra L.
Lonicera
L. korolkowii Stapf, L. sovetkiniae V. Tkatschenko, L. tatarica L., L. tatarica f. alba
L. chrysantha Turcz., L. demissa Rehd., L.
maackii Rupr., L. morrowii A. Grey, L. ruprechtiana Regel, L. xylosteum L., L. xylosteum f. nana
Caprifolium
Cypheolae
L. dioica L., L. prolifera (Kirchn.) Rehd.,
Caprifolium
L. caprifolium L., L. periclymenum L.,
L. x heckrottii Rehd.
- 39 % - , , , , 19 % , 15 % - , 12 % - (.). , . 53 ,
(, 1958), - 15.
35-50 - L. altaica, L. maackii, L.
morrowii, L. xylosteum, L. tatarica 75 .
10-13 .
, . ( I
( , 1975)) 11 (44%) .
Caeruleae, L. xylosteum, L. korolkowii, L. maackii, L.
morrowii, L. nigra, L. vesicaria. 24 % : L. alpigena, L. chrysantha, L. involucrata, L. maximowiczii,
L. ruprechtiana, L. sovetkinae. , , . , L.
alpigena, , , -
298
________________________________________________________________
,
. L. involucrata , .
( III-IV) 16 % . L. demissa. ( Caprifolium).
. , ,
.
, . , .
, ( VI), L. alberti ( ) L. ferdinandi.
17 . ,
, . (L. caprifolium) , , ,
- . L. alpigena ( , ), L. involucrata (
), L. nigra ( ).
, : L. tatarica (c Alba ), L. maackii, L. maximowiczii, L. morrowii, L. korolkowii, L. xylosteum .
. ( 1 )
L. alpigena, L. altaica, L. ferdinadi, L. korolkowii, L. demissa. , 1 (L. xylosteum f. nana). (1.5-2.0 ) : L. caerulea, L. edulis, L. involucrata, L. maximowiczii, L. pallasii, L. xylosteum. , 2-4 , L. maackii, L.
morrowii, L. ruprechtiana, L. tatarica.
. , L. tatarica, , 1.2
4.5 . , , . , , ,
, .
: L. alpigena, L. altaica, L. caerulea, L. edulis, L. involucrata, L. korolkowii, L. maackii, L. maximowiczii, L. morrowii, L. nigra, L. pallasii, L. ruprechtiana, L.
sovetkinae, L. tatarica, L. xylosteum. ,
,
L. caprifolium. (L. periclymenum, L. prolifera) .
L. chrisantha, L. demissa, L. dioica,
L. heckrottii, L. tolmatchevii, L. vesicaria. , ( 1999 2008 .)
L. tolmachevii (. ). ,
(2008).
(. -) 1999 . . 4 .,
2 . 2007 . 59 , . - , .
L. albertii, L. ferdinandii.
L. albertii, L.x heckrottii L. tolmatchevii,
299
___________________________________________________________________________
L. maximowiczii (
(, 2003)).
Caeruleae ( ) , , . -
41 (L. caerulea s.l.) .
.. , , . .. ,
- , .
L. edulis - ( 1996 .), 22 L. pallasii,
. 2000 . . ... .. 2002-2003 . , .
,
Caeruleae 1-4 ( ) (, , 2002). ,
L. caerulea ( ), , " ". -, , ..
, -, -
. , , , , ,
, , , , ,
(L. kamtschatica (Sevast.) Pojark.),
68 - (L. turczaninowii Pojark.)
, .
,
.
. (
-) 3-4 , , ,
,
- . , , , ,
(0.9-1.4 ) . L.
caerulea, L. pallasii , , .
(
) , L. altaica
, 4-11-59, L. kamtschatica , , , , ,
L. turczaninowii , (, 2007).
,
15. , - .
. .
( Caeruleae) - .
.
300
________________________________________________________________
1. . .: , 1973. .20-26.
2. . ., 1975. 547 .
3. (
50 ; .III) / .., .., ..,
.. .: , 2003. 214 .
4. .. . ., 1974. 704 .
5. .. // , ., 1958, .23.
6. .. Lonicera caerulea L. // . ( .
. ., 5-8 2007 ., . -). ., 2007. . 351-353.
7. .. . .: , 1980. 160 .
8. .., .. : . .: , 2002. 160 .
9. - . .: , 1977. .IV. . 144-148.
-
.
.,
L. alpigena
1964, 1995,
2004
, ,
0.8-2.0
I-III
L. altaica
, ,
1964
L.
caprifolium
1955, 1978,
1995-1997,
1999, 2000,
2001
II-V
L. chrysantha
1999-2001
II
L. demissa
1999, 3.
(),
-
1.5
II-III
2000, 2002
1998
--
1.0
IV- VI
1.7
I - III
()
1.5
1976,
2.5
I - II
1976,
1.5
II
L. edulis
L. ferdinandi
, , ,
L. involucrata
L. korolkowii
L. maackii
L. maximowiczii
,
, ,
,
1998, 3
1999, 3
, , , ,
(
.),
, -, ,
0.6
1.0-1.2
301
___________________________________________________________________________
.
L. morrowii
1948; 1976
L. nigra
1999, 3
()
1976
10
L. pallasii
L. periclymenum
L. ruprechtiana
L. sovetkinae
L. tatarica
L. tatarica f. alba
L. tolmatchevii
L. vesicaria
L. xylosteum
,
,
, ,
. .
,
2000, 3
1939,
1999 (3. .)
1999,
1936, 1999
1936
1999,
?
1976
2.2
1.5-1.8
III-IV
3.2
I-II
1.9
I-II
10
1.2-4.5
4.5
0.6-0.9
3 .
I-II
,
(
.)
,
(
.),
1.2
1.5
302
________________________________________________________________
58.087: 634.511
JUGLANS
-
..
..
..
, ,
. , . , 85
e-mail:Rezanova@bsu.edu.ru
sorokopudov@bsu.edu.ru
nazarova@bsu.edu.ru
- Juglands : J. regia L., J. manshurica Max., J. siboldiana Maxim., J. cordiformis Max., J. nigra L., J. cinerea
L., J. rupestris Engelm. , , , ,
. ,
, , ,
.
: Juglans, , , , , .
: 2,
t : ,
.
- ,
[9].
.
,
.
.
. ,
.
,
,
. Juglans ,
, [2, 3, 5].
75% , 22%, 10%, : , ,
, .
, , , , , ,
[16].
,
.
Juglans,
, 2001 : J. regia L., J. manshurica Max., J. siboldiana Maxim., J. cordiformis Max., J. nigra L., J. cinerea L., J. rupestris
Engelm. J. regia L.,
.
.
(7- )
303
___________________________________________________________________________
, .
: , , (1:1:1) [14].
[15].
13, -5, 10 Quanta 200 3D
, -.
,
50- .
, , () [8],
. ,
,
.
.
, , [1].
, [4].
. . (1984), Microsoft Office.
[7].
.
.
,
.
.
,
, ,
,
.
,
, -
,
[7].
.
[16]. .
, ,
.
, ,
, (. 1).
[9]. Juglans
[1]
[7].
304
________________________________________________________________
J. cordiformis
J. siboldiana
J. cinerea
J. nigra
J. rupestris
J. regia
J. manshurica
. 1. Juglans ( 8000).
, , , ,
, .
[1].
.
,
.
().
.
,
(. 1).
,
.
305
___________________________________________________________________________
.
, ,
,
[13].
Juglans
J. manshurica
J. siboldiana
J. regia
J. rupestris
J. cinerea
J. nigra
J. cordiformis
S , 2
265,828,64
278,5017,29
274,5517,25
375,5713,43**
357,5416,89**
250,6211,27
307,068,81
10,471,35
15,353,30*
6,171,65
8,740,466
7,911,27
8,120,958
7,480,847
1 2
150,013,50
155,533,0
81,333,0
120,34,66
83,3312,7
101,59,58
73,58,47
* 0,95; ** 0,99
( J. Regia ); .
, ,
,
.
( , ,
, ). 430,37-677,26 .
,
- (. 1).
J. regia
J. rupestris
J. nigra
J. cinerea
J. manshurica
J. siboldiana
J. cordiformis
. 2. Juglans
306
________________________________________________________________
,
.
.
.
,
[13].
.
[11,12].
,
(. 2). ,
, , , ,
.
, .
.
, .
1. Juglans
,
.
2.
,
.
1. .. /
.. / . .. . .: . , 1988. 208 .
2. .. Juglans L. /
.. , .. // . .
(23-25 2003 ., -).
2003. . 179-180.
3. .. //
. 2002. . 274. . 444-448.
4. , .. /
.. . - .:, 1989. - 204.
5. ..
(POACEAE) // .
2009. . 94, 8. . 1204-1215.
6. .. //
. . 2009. 5. . 60-62.
7. .. /
.. // . 2009. 7. . 17-18.
8. , ..
/ .. , .. //
( -
. 19-22 2006). : , 2006. . 344.
9. .. /
.. , .. : , 1991. 307 .
10. ..
/ .. , .. , .. //
. 2010.
. 42, 1. . 3-21.
307
___________________________________________________________________________
11. , ..
HAMAMELIDACEAE / .. , .. // .
2010. . 95, 3. . 338-345.
12. ..
/ .., .. // I
. . , 2003. 2. 83.
13. .., .. ..
Populus (Salicaceae) // . 2002. - . 87, 1. - 63-72.
14. .. - Ribs americanum
Mill. .
. 2010. 20.
15. .., .. -
Ribs americanum Mill. //
: , , ,
.-. ., .: 2008. .133-135.
16. , .. -
/ .. , .. //
. 1992. 2, . 24. . 107-116.
17. , .. / .. , .. , ..
. .: -, 1978. 256 .
308
________________________________________________________________
581.5
..
..
, 308007,
. , . 85
e-mail: martynova @bsu.edu.ru
tokhtar@bsu.edu.ru
.
.
. ,
.
: , , .
.
, , . ,
, , , .
, - .
, . .
.
, , . .
,
, .
- .
,
, . -
309
___________________________________________________________________________
. ,
: -, - ,
.
-
, .
. (Ellenberg, 1974,) . (Landolt, 1977).
: ,
-, , .
, ,
, , . ,
,
, .
, , - .
. ( ) ( ).
, . , , ,
(, , .),
(pH 6.3 4.3) , , . (, 2005).
.
, , - . , , . , ,
.
: , , , . , .
: , , , ,
, . ,
, .
, ,
. . .
310
________________________________________________________________
Berberis vulgaris, Ulmus parvifolia, Caragana frutex, Kerria japonica, Frangula alnus,
Corylus avellana, Elaegnus angustifolia, Juniperus sabina, Hippophae rhamnoides,
Rosa cinnamomea, Rosa canina, Pinus silvestris
Armeniaca mandshurica, Armeniaca vulgaris, Amorpha fruticosa, Aralia elata, Aronia
melanocarpa, Berberis coreana, B. vulgaris, B. thunbergii, Phellodendron amurense, Betula pendula, Euonymus verrucosa,E. bungeanus, E. europaea, E. alatus, E. atropurpurea, Euonymus latifolia, Biota orientalis, Ligustrum vulgare, Crataegus submollis, C.
holmesiana, C. ellwangeriana, Sambucus canadensis, S. racemosa, Sambucus sibirica, S.
nigra, Fagus orientalis, Gymnocladus dioicus, Vitis labrusca, Cerasus mahaleb, C.
fruticosa, Ulmus parvifolia, Gleditsia triacanthos, Carpinus orientalis,C. betulus, Pyrus
elaeagrifolia,Pyrus communis, P. ussuriensis, Partenocissus guinguefolia, Cornus alba,
C. sanguinea, C. stalonifera, C. siricea, Quercus rubra, Q. robur, Picea pungens,
Lonicera alpigena, L. flava, L. canadensis, L. xylosteum, L. olgae, Iberis sempervirens,
Salix alba, Salix purpurea, S. fragilis, S. acutifolia, S. cinerea, Salix pentandra, S. rossica, S. dasyclados, Viburnum opulus, V. trilobum, Caragana arborescens, C. frutex, C.
pigmea, Catalpa bignonioides, Aeskulus hippocastanum, Kerria japonica, Cotoneaster
lucidus, Cotoneaster melanocarpus, Acer ginnala, A. platanoides, A. campestre, A.
saccharinum, A. tataricum, Juniperus sabina, Juniperus scopulorum, Hippophae
rhamnoides, Alnus incana, A. glutinosa, Juglans regia, J. manshurica, J. cinerea, J.
nigra, Artemisia abrotanum, Robinia viscosa, Robinia pseudoacacia, Rosa woodsii,
Rosa acicullaris, R. cinnamomea, R. multiflora, R. rugosa, R. nutcana, R. pendulina, R.
glauca, R. canina, Sorbus aucuparia, Sorbus sibirica, Sorbaria sorbifolia, Securinega
suffruticosa, Syringa joseikaea, S. vulgaris, Cotinus coggigria, Prunus spinosa, Prunus
divaricata, Ribes alpinum, Ribes aureum, Ribes rubrum, R. lucidum, R. nigrum, Pinus
pallasiana, Pinus silvestris, Spiraea chamaedryfolia, S. trilobata, Rhus typhina,
Tamarix ramosissima, Populus alba, P. berolinensis, P. tremula, P. maximowiczii, P.
simonii, P. nigra, Thuja occidentalis, Forsythia europaea, Padus avium, P. sirotina, Philadelphus coranaris, Morus alba, Shepherdia argentea, Malus silvestris, Fraxinus lanceolata, Fraxinus excelsior, Fraxinus ornus
Berberis vulgaris, Betula pendula, Ulmus parvifolia, Caragana frutex, C. pigmea, Kerria
japonica, Frangula alnus, Corylus avellana, Elaegnus angustifolia, Juniperus sabina,
Hippophae rhamnoides, Rosa acicullaris, R. cinnamomea, R. canina, Cotinus coggigria,
Pinus silvestris, Rhus typhina
Armeniaca mandshurica, Armeniaca vulgaris, Aralia elata, Aronia melanocarpa,
Berberis vulgaris, Betula pendula, Biota orientalis, Sambucus canadensis, S. nigra,
Cerasus mahaleb, C. fruticosa,. Ulmus parvifolia, Gleditsia triacanthos, Picea pungens,
Salix purpurea, S. acutifolia, Caragana arborescens, C. frutex, Catalpa bignonioides,
Aeskulus hippocastanum, Acer saccharinum, A. tataricum, A. negundo, Frangula alnus,
Corylus avellana, Elaegnus angustifolia, Juniperus sabina, Hippophae rhamnoides,
Alnus incana, A. glutinosa, Juglans regia, J. manshurica, J. cinerea, J. nigra,
Physocarpus opulifolius,. Robinia pseudoacacia, Rosa acicullaris, R. cinnamomea, R.
rugosa, R. canina, Sorbus aucuparia, Sorbus sibirica, Sorbaria sorbifolia, Cotinus
coggigria, Prunus divaricata, Ribes aureum, Pinus pallasiana, Pinus silvestris, Rhus
typhina, Tamarix ramosissima, Populus alba, P. maximowiczii, P. simonii, Populus x
sowietica cv. Pyramidalis, P. nigra, Thuja occidentalis, Forsythia europaea, Padus
avium, Morus alba
Salix pentandra, Caragana frutex, Kerria japonica, Elaegnus angustifolia, Juniperus
virginiana, Juniperus sabina, Juniperus scopulorum, Hippophae rhamnoides, Physocarpus opulifolius, Pinus silvestris.
, , I ,
2.5 0,8 ., 1.5 0.5 ., . 3-5 .
- , II
, -
311
___________________________________________________________________________
5 , 2.5
0,8 ., 1.5 0.5 . .
, III ,
2.5 0,8 ., 1.5 0.5 . .
,
, 5- 90 % .
, , : , , ,
, , ,
, , .
() , : ,
, , .
""
: , , , .
.
, .
, , 1:1, : , ,
, , . .
,
: (Sorbus sibirica Hedl.), (Caragana arborescens L.), (Crataegus ellwangeriana Sarg.), (Cotinus coggigria Scop.), (Elaeagnus argentes
Pursh.), (Sambucus sibirica L.), (Swida alba, Thelycrania alba L.), (Aronia melanocarpa Elliot.), (Physocarpus opulifolia Maxim.), (Syringa josikae Yacg.),
(Spiraea chamaedryfolia L.) (Acer tataricum
Z.), (Rosa rugosa Thunb.) . . , , .
. . .
155
, , . ,
. 50
% .
25-30%.
. -
312
________________________________________________________________
, , , .
, . .
, . ,
.
1. ..
( ).
, 2010 50 .
2. .. ( -
).
, 2005 36 .
3. Ellenberg H. Zeigerwerte der Gefasspflanzen Mitteleuropas. Gottingen: Goltze. 1974. 97 s.
4. Ellenberg H. Vegetation Mitteleuropas mit den Alpen in okologischer, dynamischer und
historischer Sicht. 5. Aufl. Ulmer, Stuttgart. 1996. 1096 s.
5. Landolt E. Okologische Zeigerwerts zur Sweizer Flora. Veroff. Geobot. Inst. ETH. Zurich.
1977. H.64. S. 1-208.
313
___________________________________________________________________________
634.722.581
SBUCUS L.
..
..
, 308007,
. , . 85
e-mail: sorokopudov@bsu.edu.ru
.
Sambucus L.
.
: , , Sambucus L.,
.
- .
,
.
- , . , . .
, , , . (Sambucus
L.) ;
.
,
, .
() 0,088 0,179 . , .
- ,
.
, ,
. 20-30 (King, 1975),
. :
, , ,
, ,
. . (, 1976; , 1982; , 1976; , 1937), , 1
582 .
5-15 .
, , -
314
________________________________________________________________
,
. , ,
.
, , .
, .
, , (, 1983).
- , .. . (- -,
, 1986). -1. (2) (1)
. , -1 10-60 0,01 , (35-40) 1 ; . 30-
. 2/1=0,9-0,8 ,
0,7-0,6 , 0,5-0,4 .
,
, .
Sambucus racemosa 'Plumosa'
L., ( 2)
(1) (.). , ,
,
. 2/1
Sambucus racemosa 'Aurea' L. 0,820,05 ,
,
. , 2/1
. ,
3% (Sambucus nigra L.) 8% (Sambucus racemosa L., Sambucus sibirica Nakai.). , .
, .
315
___________________________________________________________________________
Sambucus L.
, .
S. nigra L.
S. canadensis L.
S. racemosa 'Aurea' L.
S. racemosa L.
S. racemosa 'Plumosa' L.
S. sibirica Nakai.
S. canadensis 'Plumosa' L.
S. coreana Nakai.
S. nigra L.
S. canadensis L.
S. racemosa 'Aurea' L.
S. racemosa L.
S. racemosa 'Plumosa' L.
S. sibirica Nakai.
S. canadensis 'Plumosa' L.
S. coreana Nakai.
2/1()
19425
13
21612
6
19923
12
26446
17
31253
17
25638
15
23816
7
24547
19
19425
13
21612
6
19923
12
26446
17
31253
17
25638
15
23816
7
24547
19
16622
13
18716
9
16317
10
21847
22
28260
21
21032
15
20918
9
21442
19
16622
13
18716
9
16317
10
21847
22
28260
21
21032
15
20918
9
21442
19
0,860,03
3
0,870,04
5
0,820,05
6
0,820,06
8
0,900,04
5
0,830,06
8
0,870,03
4
0,880,03
4
0,860,03
3
0,870,04
5
0,820,05
6
0,820,06
8
0,900,04
5
0,830,06
8
0,870,03
4
0,880,03
4
1. Sambucus L.
.
2.
.
1. .., .., .. . .: , 1976. - . 6-17.
2. .. - , .: , 1982. 280 .
3. .. // , 1983, 1, 15-24.
4. .., .., .. -
-, , -
, 1986. - 39 .
5. ..
, .: , 1976.- 54-60.
6. .. , .: , 1937, . 3. 435 .
7. King M.C., Wilson A.C. // Sciensa, 1975. 188. 107-116.
316
________________________________________________________________
DROUGHT RESISTANCE OF SOME REPRESENTATIVES OF SORT SBUCUS L.
IN THE CONDITIONS OF THE BELGOROD REGION
S.V. Koltsov
V.N. Sorokopudov
Belgorod National Research
University, 308007, Belgorod,
Pobeda-str., 85
e-mail: sorokopudov@bsu.edu.ru
Adaptation kinds of an elder in the conditions of a droughty climate of the Belgorod region is studied. Plants completely pass a phenological cycle of development at influence on them of the raised temperatures of air. Plants of sort Sambucus L. re carried to number perspective for cultivation in the conditions of the Belgorod region.
Keywords: a sort, a kind, Sambucus L., drought resistance.
317
___________________________________________________________________________
581.95(470.325)
-
-
1
..
..
..
..
, 308015,
. , . , 85
- .
,
.
: , ,
, .
e-mail: tokhtar@bsu.edu.ru
,
. [1]. . , , , - .
.
-
.
- . - .
- , , [2, 3].
, , -: , , -, , .
- , .
, :
( ), ( ), ( ), ( ), . ( ). .
2008-2011 .
189 41 .
1
, 16.740.11.0053 01.09.2010.
318
________________________________________________________________
10 . : Asteraceae 18,0%, Fabaceae
11,6%, Lamiaceae 8,5%, Poaceae 8,5%, Brassicaceae 6,3%, Ranunculaceae 5,8%, Rosaceae
4,8%, Apiaceae 3,7%, Scrophulariaceae 3,7%, Boraginaceae 2,1%. 27,0 %, 19 , 9 . . , 52,9%, 23,3% . 7,4%, 6,9%, , 2,1%. , . ,
, ( ), ,
.
,
, 22 , : Adonis
vernalis L., Anemone sylvestris L., Asperulla tephrocarpa Czern. ex M. Pop. et Crishan, Astragalus albicaulis DC, Carex humilis Leyss., Cephalaria uralensis (Murr) Schrad.ex Roem.et
Schult., Clematis integrifolia L., Diplotaxis cretacea Kotov, Gentiana cruciata L., Hyacinthella
leucophaea (K. Koch), Linum perenne L., Linum ucranicum Czern., Onosma simplicissima L.,
Polygala sisbirica L., Primula veris L., Pulsatilla patens L., Silene supina M. Bieb., Stipa pennata L., Hlianthemum canum (L.) Hornem, Schivereckia podolica Andrz., Scutellaria supina L.,
Thymus cretaceus Klok. et Shost., 3 (Helictotrichon desertorum (Less.) Nevski, Oxytropis pilosa (L.) DC, Ranunculus pedatus Waldst. et Kit) [2]. , ,
25 , 13,23% .
214 , 34 . : Asteraceae 20,1%, Fabaceae 9,8%, Lamiaceae 8,9%, Poaceae 8,4%,
Brassicaceae 5,6%, Scrophulariaceae 5,1%, Apiaceae 4,7%, Rosaceae Liliaceae 3,7%,
Ranunculaceae 3,3%. 26,7% Rubiacecae Caryophylaceae 6 , Boraginaceae 5 , Euphorbiaceae 4 , 5 3 2 9 1 . , Asteraceae, Fabaceae, Lamiaceae, Apiaceae.
. , 52,8%,
26,5%, 11,7%, 5,2% 3,8%. 24 .
, () .
125 , 40
. : Asteraceae (20,8%
), Fabaceae (12,8%), Lamiaceae (7,2%), Rosaceae (5,6%), Scrophalariaceae (4,8%), Brassicaceae (4,8%), Poaceae (4,8%), Caprypholiaceae (3,2%), Ranunculaceae (2,4%), Cyperaceae (2,4%), Plantaginaceae (2,4%), 7
, 22 . , (.). () , : : III .
,
: (), ( . , . . . . , . ),
(. ) ( , 2004). -
, .
319
___________________________________________________________________________
3% 4%
17%
18%
58%
,
109 38 .
: Asteraceae (15% ), Poaceae (9%), Fabaceae
(7%), Lamiaceae (6%), Rosaceae (6%), Caprypholiaceae (4%), Ranunculaceae (4%), Polygonaceae (3%), Scrophulariaceae (3%), 6 3 , 7 , 16
. 171 , 45 . : Asteraceae (11,7% ), Lamiaceae (9,36%), Fabaceae (9,36%), Poaceae
(6,43%), Scrophulariaceae (5,26%), Ranunculaceae (4,68%), Apiaceae (4,09%), Rosaceae
(4,09%), Brassicaceae (3,51%), Boraginaceae (2,92%), Liliaceae (2,92%), 3 , 3 , 12 , 16 . , , 59.
62 , .
. 41%,
5% , (36%).
,
, , . ,
, , .
, .
,
, .
. 171 . , 45 , , ,
Asteraceae. Asteraceae (12,87%), Fabaceae (8,19%), Lamiaceae (7,6%), Rosaceae (6,43%), Poaceae (5,26%), Apiaceae (5,26%), Brassicaceae (4,68%), Ranunculaceae
(4,68%), Scrophulariaceae (5,51%) Liliaceae (3,51%).
, , - , ,
. ,
320
________________________________________________________________
.
.
Rosaceae Brassicaceae. Brassicaceae ,
, , . Rosaceae , , , . , , , .
, . ,
, .
, , , ,
c . - - . q ,
, , [4],
. , .
, .
1. .. . : , 1991. -168 .
2. ... .. - // . . . . 2010. - . 1716. . 108-111.
3. .., .., .., .., .. // . 1. - 2009. .121-124.
4. .., .. :
// . 2008.
. 50-54.
Local florae of natural-reserved fund territories of the southwest of Central Russian upland were studied. Structures of these florae
and the florogenetic processes which occure here are similar. It is connected with anthropogenous impact which leads to changes in taxonomical and typological spectra. The data obtained allow to unite the
florae in the common anthropogenously transformed type of flora in the
southwest of the Central Russian upland.
Keywords: flora, anthropogenous transformation, structure, florogenetic processes
321
___________________________________________________________________________
581.4:582.477.6
.
, 450080,
. , . , 195,
. 3
: 6,12 , 5,48 ,
1000 47,49 , 4,42 ,
2,61 , 2,23 ., 1000 10,61 .
-
.
e-mail: gfbelal@mail.ru;
vpp99@mail.ru
: ,
, , .
..
..
. ,
, ,
[17]. .
.
10 . , ()
, (-) .
2006-2009 . [1, 16]. 0,10 0,31 . 15-30 . 10 .
1000 .
( 220 . ) 1000 100
( ) [4].
Juniperus L.
8 . , 5-9 , -, - , . 3 ( 1-2),
-, , 1-3 , , 3,5-5 , 2,5-3 [3, 5, 6, 11, 13-15].
322
________________________________________________________________
(.), 5,41 ( )
6,55 (), 4,76 () 6,40 ().
( )
. ,
.
, ,
,
.
(, )
, .
, , . , .
[18]. , 7,2 , 5,7
; : 4,6 , 3,8 .
, , . 25- ( ) [9]
4,94 6,56 , 4,13 6,49 . ..
[9] ,
, . , , .
(. .) : 32,42
( ) 73,56 (). (
, , , , ..
).
( , . ), ,
, .
(. ), -, .
.. [2] (
1 . 1000 .) (114 ), - (37 ).
, 0,04-0,185 ( 40
185 1000 .) [12]. [18] 100 . 4,6 (.. 46 1000 .). ,
( 47,49 ; . .)
( )
.
(. .) 4,24 ( ) 4,57 (), 2,42 ( ) 2,88 (-). . [2] (4,62-4,75 ). .. [9],
3,6 4,55 ( 3,98
323
___________________________________________________________________________
.. ), 2,04
3,05 ( 2,43 ). ,
, , ,
, .
,
,
,
,
1000 .,
1000 .,
, .
(- -
c )
6,180,1
5,190,0 45,492,8 4,450,0 2,830,0
14,270,6
1.
1,520,094
28
98
15
93
41
60
6,150,1
5,220,0 45,801,7 4,490,0 2,880,0
14,960,6
2.
1,430,056
21
70
40
78
33
93
6,170,11 5,040,0 40,192,8 4,360,0 2,730,0
11,850,8
3.
1,550,104
1
85
15
68
61
66
6,520,1
6,400,1 73,568,0 4,570,0 2,450,0
2,940,03 9,720,29
4.
37
65
96
89
65
7
6
6,370,1
6,220,1 66,102,9 4,470,0 2,420,0
10,160,4
5.
2,920,031
37
16
38
70
39
54
6,280,0 5,610,2 54,236,5 4,470,0 2,660,0 2,0720,35 12,281,1
72
87
56
34
96
1
02
(- )
5,690,1
5,250,2 46,675,5 4,430,0 2,560,0
8,520,97
6.
2,610,102
42
10
91
92
59
2
2,330,111
69
37
80
97
76
84
5,890,1
5,300,1
32,421,7
4,510,0 2,680,0
2,270,08
9,260,3
9.
19
27
39
94
57
7
85
5,410,1
4,760,1
35,152,4
4,370,0 2,470,0
7,290,49
10.
2,370,095
33
39
67
87
82
3
5,960,2
5,350,1 40,762,9 4,380,0 2,560,0
2,390,05 9,020,52
04
99
1
45
52
9
1
- 6,120,11 5,480,1 47,494,0 4,420,0 2,610,0
10,610,7
2,230,176
4
70
59
30
54
65
(. .)
1,43 . (-) 2,94 . (). ( , ) , 4, 5 6 . . ,
( ) ( 3 ), (1-2 ). 2-3 .
1,4 2,3 [2],
2-3 [10]. [7, 8], ,
1-2, 3 ( 4 5) .
[9] 1,2 2,68 . 1,93 .
.. [9]. ,
324
________________________________________________________________
, , , ( ).
1000 . (. .) 7,29 () 14,96 (-).
(, ) , ( ; . )
( 12,28 ) - (9,02 ).
[2]
1000 . 7,18 11,15 . 7,15-9,08 [10]. 1000 . [18] 9,6 , 13,5 . . ,
(
14,96 ). , , (16,0 ), [: 2].
. (r = 0,7).
, 1000 (r = 0,63 r = 0,85 ). , (r = 0,64),
.
(r = -0,88), .. .
[9]. ,
(r = 0,8), .. .
, (r = -0,77)
1000 . . ,
, 1000 . .
, , , ( , ); .
, .
:
6,12 , 5,48 , 1000 47,49 , 4,42 , 2,61 , 2,23 ., 1000 10,61 . .
, .
.
1. .., .., .. . .
.: , 2002. 240 .
325
___________________________________________________________________________
2. .. -
(Juniperus communis L.) : . . . .-. .
06.03.03. , 2007. 18 .
3. .. . .
, 2004. 207 .
4. 13056.4-67. . 1000
. . 01.07.68 . ., 1977.
5. .. . , 1969. - 93 .
6. .., .. . -: -
, 1976. 104 .
7. .. . . 2007. 1. . 65-69.
8. ..
(Juniperus L.) // - . , . 300-
. . ., 2007. . 102-104.
9. .. // . 2010. 5. . 36-44.
10. .., .. // .
1965. 1. . 57.
11. .., .. . : - , , 2002. 707 .
12. .. Juniperus
communis L. ( ) // . . 1999. . 35, . 4.
. 13-20.
13. .. . Juniperoideae Pilg. //
. , . . 1. .
.; .: - , 1949. . 340-376.
14. .., .., .. ,
. . 1. .: - . , 2006. 259 .
15. .. . .: . -, 1980. 96 .
16. . . 3. .; .: , 1964. 530 .
17. .., .., .. : - . .: , 2004. 276 .
18. ..
// . -, 1998. . 314-316.
Mean parameters of generative organs of Juniperus communis L. in the South Urals and Cis-Urals are determined: cone
length is 6.12 mm, cone width 5.48 mm, mass of 1000 cones
47.49 g, seed length 4.42 mm, seed width 2.61 mm, seed number per cone 2.23, mass of 1000 seeds 10.61 g. Cenopopulations
of J. communis in the region are characterized by intermediate position in the parameters of cones and seeds between Easterneuropium and Siberian populations.
Key words: Juniperus, cenopopulation, cones, seeds, the
South Urals.
326
________________________________________________________________
574.3 + 582.579.2 (471.61)
IRIS PUMILA L. 1
.. 1
.. 2
.. 2
1)
, 344006, . --,
. . , 105
e-mail: vfedyeva@gmail.com
2)
,
. --, 344041,
. , 7
e-mail:anshmaraeva@sfedu.ru
shishlova@sfedu.ru
2005-2010 .
Iris pumila L. 60
.
Iris pumila.
, , ,
, . ,
Iris pumila
,
. , ,
, ,
.
: , ,
, , , .
: : .
327
___________________________________________________________________________
[2, 4]. 60
( 100 ) Iris pumila
, .
, ()
- .
1
: - -, 2 ,
. , , - .
: 470 36.978 . ., 400 42.092 . .
: .
: - ; : Stipa pulcherrima Agropyron pectinatum Artemisia marchalliana + .
: ,
,
. , .
50 %.
( , ):
1. Stipa pulcherrima cop2
38. Holosteum umbellatum sp3
2. Agropyron pectinatum cop2
39. Hylotelephium stepposum sp2
3. Artemisia marchalliana cop2
40. Hypericum perforatum sp2
4. Alyssum calycinum sp3
41. Iris pumila cop1
5. A. desertorum sp3
42. Jurinea cyanoides sp1
6. Artemisia austriaca sp3
43. Koeleria cristata sp3
7. Asparagus officinalis sp1
44. K. sabuletorum sp3
8. Astragalus pseudotataricus sp2
45. Lappula squarrosa sp2
9. A. varius sp3
46. Linaria genistifolia sp2
10. Bellevalia sarmatica sp1
47. Medicago minima sp1
11. Bromopsis riparia sp1
48. Meniocus linifolius sp2
12. Bromus squarrosus sp2
49. Oxytropis pilosa sp3
13. Centaurea carbonata sp3
50. Papaver rhoeas sp1
14. Cephalaria uralensis sp3
51. Phleum phleoides sp1
15. Chamaecytisus ruthenicus sp3
52. Poa crispa sp3
16. Chondrilla latifolia sp2
53. Potentilla argentea sp3
17. Cleistogenes bulgarica sp3
54. P. astracanica sp1
18. Crupina vulgaris sp1
55. P. humifusa sp3
19. Dianthus polymorphus sp2
56. Rumex acetosella sp1
20. Echinops ruthenicus cop1
57. Scabiosa ochroleuca sp3
21. Elytrigia repens sp3
58. Scorzonera mollis sp3
22. Ephedra distachya sp3
59. Senecio vernalis sp1
23. Eremogone biebersteinii sp2
60. Silene chlorantha sp1
24. Eryngium campestre sp3
61. S. supina sp1
25. Euphorbia seguieriana sp3
62. Stachys atherocalyx sp3
26. E. virgata sp3
63. Stipa capillata sp2
27. Falcaria vulgaris sp3
64. Teucrium polium sp3
28. Ferulago galbanifera sp1
65. Thymus dimorphus cop1
29. Festuca beckeri sp3
66. Trinia hispida sp3
30. F. valesiaca sp3
67. Tulipa schrenkii sp1
31. Galatella villosa sp3
68. Valeriana tuberosa sp1
32. Galium aparine sp1
69. Verbascum phoeniceum sp3
33. G. octonarium sp2
70. Veronica spicata sp3
328
________________________________________________________________
34. G. verum sp3
71. V. verna sp3
35. Gypsophila altissima sp3
72. Vincetoxicum maeoticum sp1
36. Helichrysum arenarium sp3
73. Viola arvensis sp3
37. Hieracium echioides sp3
74. Xeranthemum annuum sp3
, , , ,
.
, -
.
Stipa pulcherrima C. Koch
(), Festuca valesiaca Gaudin, Artemisia austriaca Jacq., Galatella villosa (L.)
Reichenb. fil., Falcaria vulgaris Bernh., Eryngium campestre L. .; Cleistogenes bulgarica (Bornm.) Keng, Ephedra distachya L.,
Gypsophila altissima L., Silene supina Bieb., Echinops ruthenicus Bieb., Thymus dimorphus
Klok. & Shost. .; Artemisia marchalliana
Spreng. (), Festuca beckeri (Hack.) Trautv., Koeleria sabuletorum (Domin)
Klok., Dianthus polymorphus Bieb., Helichrysum arenarium (L.) Moench, Rumex acetosella L.,
Astragalus varius S.G. Gmel. .
,
,
4000 . ; Iris pumila ,
. , . , 20-50 . 100 .,
, 20 %.
. 1 .
. , , , .
, .
. , 15 .
.
: .
: , , , .
2
: -, 3 ,
. , , .
: 4700 36.641 . ., 4100 09.558 . .
: .
329
___________________________________________________________________________
330
________________________________________________________________
. , 2:4:1. Iris
pumila ( 20-30 10
. ). , , 20-40 ;
100 . ( 20 %). .
-
( Stipeta ucrainicae, Stipeta lessingianae, Stipeta pulcherrimae, Stipeta dasyphyllae, Stipeta asperellae, Festuceta valesiaceae .)
.
15-20 , . Iris pumila , (
, ),
( 7085 ), . ,
( Bothriochloa ischaemum + Stipa pulcherrima Koeleria sabuletorum) Iris pumila ,
. 250-270 .
, ,
Iris pumila
. ( , , , , ).
, . , ,
:
; Iris pumila , ; . , Iris pumila -
33 %, [5].
, .
, Iris
pumila , ,
( 30) . Iris pumila , ,
. , Iris
pumila , ( )
( , )
. , Iris pumila ( -)
331
___________________________________________________________________________
(- -), ( -),
( -), ( -)
-- ( -).
Iris pumila . , , . , . ( -);
, ( -),
- ( -). 38,5 . .
( , , , ,
.), .
Iris pumila
, , 2008 .
165 . , 600
[6].
1. ( ). .: - , 2008. 855 .
2. . . 2. , . --: , 2004. 333 .
3. .., .., .., .., ..
// . . 1977.
1. .105-108.
4. .., .. //
() . ,
2005. .29-36.
5. .. // . - . . .
2008. 4(146). .75-79.
6. .., ..
// ,
. :
, 2010. .434-435.
e-mail: vfedyeva@gmail.com
Botanical garden of SFU,
Botanichesky spusk, 7,
Rostov-on-Don, Russia
2)
e-mail: anshmaraeva@sfedu.ru
shishlova@sfedu.ru
332
________________________________________________________________
:581.145:633.88:581.52
-
UPATORIUM CANNABINUM L., LOPHANTHUS ANISATUS BENTH.
MONARDA CITRIODORA CERV.,
.. 1
.. 2
1)
, 634050, . ,
- , 36
, 634050, . ,
- , 36
2)
e-mail: babichevaNV@yandex.ru
,
,
.
. ,
. -
.
, ,
[12].
: upatorium cannabinum L., Lophanthus anisatus Benth., Monarda
citriodora Cerv.. , . ,
, , .
, , , [5]. M. citriodora. , , , . L. anisatus, - , , , , , . . cannabinum , , , [5, 13].
,
.
333
___________________________________________________________________________
upatorium
cannabinum L., Lophanthus anisatus Benth., Monarda citriodora Cerv.,
, , , [6, 13].
10 . -. - . -, ,
0.6 [1].
: ..
[14], .. [11], .. [4], .. [2]; .. [3]. ,
. 10 , . :
; ; ; ; , ; .
( 15).
5 1040 10100. Laboval l4 Olympus 41. Microsoft Office Exell2007.
Eupatorium cannabinum L. Asteraceae
, . , , - , , ,
, .
15 3 4 , , ( ), , . [13].
Lophanthus anisatus Benth. Lamiaceae , , 180 . - , - - , , , , , 5 10 , 4-6 .
Monarda citriodora Cerv. Lamiaceae , , 80 160 . , 1752 ,
, ,
[9]. ,
, , , , [10].
M. citriodora L. anisatus , , [9].
E. cannabinum :
, , [13].
, 25.230.6 30.320.7 , .
.. [4] .
. ,
4.0 , ( 3.5 ). , . ,
334
________________________________________________________________
. M. citriodora , .
L. anisatus (. 2.) , - , . E. cannabinum
- , M. citriodora , .
(30 40 %), M. citriodora
(>90 %). M. citriodora E. cannabinum (. 1.) ,
, L. anisatus , - .
; ;
; ; ;
.
,
. .
E. cannabinum L. anisatus , M.
citriodora . E. cannabinum L. anisatus , M. citriodora -. .
, . E. cannabinum,
M. citriodora . L. anisatus
.
, 58.7631.23 435,515.71 . 12. 5.710.55
29.322.19 .
335
___________________________________________________________________________
( 1, 2).
E. cannabinum : 3-6- , 1 2 ; 5-8-,
; , . L. anisatus - ; , ,
. M. citriodora .
, -
, :
Eupatorium cannabinum L. , , ,
, , , . ..
[7], .., .., .. [8] , , ,
.
Lophanthus anisatus Benth. , : , , , . Monarda citriodora
Cerv. .
,
, (3 5).
.
336
________________________________________________________________
1. / . . .. . .: , 1975. 147 .
2. .., .., .. . .: , , 2005. 160 .
3. .. . .: 1985. 160 .
4. .. .
.: - . -, 1988. 205 .
5. .. . //
. .. , 2006. :
www.mgtu.rtisystems.net.
6. ., .. // , 2006. : www.argodp.narod.ru/
7. .. / . .. . .: , 1948.
. 279 280.
8. .., .., .. // XXI . 2006. - 3. . 26 28.
9. .
// , 2006. : www.lekrasprom.com.
10. ..
. .:1961. 14 .
11. .. . .: , 1980. 317 .
12. .. / . .. . : . , 1978, - 239 .
13. / . . .. . .: - , 1959. . XXV. . 19 22.
14. .. - /
. .. -. .: , 1979. 155 .
2)
e-mail: babichevaNV@yandex.ru
The Studied anatomist-morphological factors Eupatorium cannabinum L., Lophanthus anisatus Benth., Monarda citriodora Cerv., sprouting in the south Tomsk area. It is shown that for Eupatorium cannabinum
L typical hygrophytes nature sheet; Lophanthus anisatus Benth. pertains
to mesophyte; Monarda citriodora Cerv. pertains to mesoxerophyte.
Key words: anatomy sheet plate, epidermis, mesophyll, stomatal
cells, upatorium cannabinum L., Lophanthus anisatus Benth., Monarda
citriodora Cerv.
337
___________________________________________________________________________
581.821: 58.02
ROSA L.
(ROSACEAE JUSS.)
..
..
. .
, ,
. , . , 85
e-mail: Rezanova@bsu.edu.ru,
sorokopudov@bsu.edu.ru,
evg-svinarev@yandex.ru
-
Rsa L.
. , , , , . Rsa. .
, . R. maximowicziana Regel., R. acicularis Lindl., R. foetida Persiana., R. canina L., R. glauca C. Hartm., R. pendulina L. .
. R. foetida, R. glauca.
: Rsa , , , , , .
Rsa L. ,
. Rosa
cinnamomea L. . .
,
.
,
. : P (), 1, ., , ,
. , ,
18 %, 4,5 %, 3,7 %, : 2 %, 1,8 % .; , ,
, , - , , , , . , [4], [6].
Rsa, , 2001 : R. amblyotis
. . ., R. cinnamomea L., R. maximowicziana Regel., R. pendulina L., R. acicularis
Lindl., R. rugosa Thunb., R. nutkana C.Presl., R. canina L., R. foetida Persiana.,
R. glauca C. Hartm., R. sibrica. R. canina,
, .
.
, . -
338
________________________________________________________________
(7- ) , .
: , , (1:1:1) [14].
[13].
13, -5, 10 Quanta 200 3D
, -.
,
50- .
, , () [7],
. ,
.
. , , [1].
, 2 .
Microsoft Office.
. 12 Rsa L. : . R. cinnamomea, R. glauca, R. canina , R. acicularis, R. nutkana , R. sibrica , (. 1, 2, 3). 12,590,630 , 219,46 410,95
(. 1, .). R. glauca, R. nutkana
(. 3).
R. rugosa
: , , . 1377,9-1550,13 ./ 1
2, 58,604,44 (. 1). ,
.
. R. sibrica ( 3000)
. R. sibrica ( 400)
339
___________________________________________________________________________
. R. acicularis ( 500)
. R. rugosa ( 150)
. 1. Rsa
9-10 , 2 (. 2).
: 199,8021,67 ./ 1 2, 29,612,69
.
. R. nutkana ( 640)
. R. rugosa ( 2500)
. 2.
R. nutkana, R. rugosa
.
1015,85105,34 2, 773,6788,64 2 (. 3).
.
8 . . , , . , , , ,
, ,
(. 4).
[9]. , , ,
[1]
340
________________________________________________________________
, . , , , ,
. , . [5]. , ,
, , . (r=0,874*).
. 3. R. glauca ( 1200).
. R. sibrica ( 2000)
. R. amblyotis ( 4000)
. R. glauca ( 2000)
. R. nutkana ( 300)
. R. pendulina ( 350)
. R. foetida ( 3200)
341
___________________________________________________________________________
. R. cinnamomea ( 2000)
. R. Maximowicziana
( 2000)
. R. acicularis ( 900)
. 4. Rsa
[1]. , , , , , .
(). .
, (. 1).
, .
-.
. .
1
Rsa
/
1
2
3
4
5
6
7
8
9
10
R. maximowicziana
R. amblyotis
R. cinnamomea
R. glauca
R. pendulina
R. canina
R. acicularis
R. nutkana
R. sibrica
R. foetida
S ,
2
315,069,13**
159,837,37**
301,837,43**
379,9512,82**
369,759,94**
491,4917,42
544,8418,42
417,4813,37**
489,6512,49
222,5812,56**
1,140,021**
1,310,074
1,110,018**
1,290,063
1,270,057
1,440,060
1,100,032**
1,200,040**
1,240,070
1,590,405
21,610,529**
9,832,42
8,784,57
6,950,521*
5,790,368
5,070,514
13,010,672**
9,480,506**
8,510,456**
19,899,46**
133,668,74**
234,8735,77**
130,617,17**
154,116,10**
120,575,81**
89,565,73
74,194,91
90,944,39
78,545,08
334,4768,89**
* 0,95; ** 0,99
( R. canina); .
. ,
Rsa .
(r=0,915*).
, ,
[1].
. , -
342
________________________________________________________________
, , ,
, , (. 2, . 5).
, , , .
2
Rosa
S , 2 , S , 2 ,
R. maximowicziana
814,3430,29**
18,471,05
970,0428,10**
18,883,87
R. amblyotis
14,652,40*
234,8735,77**
10,242,40
R. cinnamomea
579,8716,28*
15,031,65**
11,680,449*
R. glauca
781,2728,73**
17,061,44
1056,3939,17**
12,320,863
R. pendulina
466,379,21*
14,880,776**
602,8328,37
12,110,701
R. canina
509,8212,53
20,560,646
653,1214,86
15,231,49
R. acicularis
384,8818,00**
18,151,20
661,3980,34
13,481,20
Rosa nutkana
359,6310,35**
12,220,661**
530,4913,49**
9,330,634
R. sibrica
529,1112,87
13,00,664**
551,1619,89**
13,040,801
R. foetida
1129,8831,97**
22,481,57
970,0428,10**
16,261,76
R. foetida
16,580,956**
14,381,29
R. woodsii
16,770,976**
10,610,990*
1
2
3
4
5
6
7
8
9
10
11
12
* 0,95; ** 0,99
( R. canina).
. R. sibrica ( 1000)
. R. glauca ( 1000)
. R. amblyotis ( 1000)
. R. woodsii ( 3000)
. R. foetida ( 3000)
. R. acicularis ( 3000)
343
___________________________________________________________________________
. R. cinnamomea ( 1000)
. R. nutkana ( 1500)
. R. Maximowicziana
( 2000)
. R. rugosa ( 1500)
. R. amblyotis ( 2000)
. 5. Rsa
.
. , ,
[9].
, , .
.
, , ,
.
, , , , , , . , , , . , .
.
Ros
, , , , , .
(. 3). (. 6).
: ,
, , . , .
(r=0.809*), (r=0.807*) .
344
________________________________________________________________
, , . .. ,
, . , , [10 - 12].
: , , .
. R. woodsii ( 1000)
. R. acicularis ( 800)
. R. cinnamomea ( 800)
. R. maximowicziana
( 1500)
. R. rugosa ( 800)
. R. nutkana ( 800)
. R. pendulina ( 800)
. R. sibrica ( 800)
. R. amblyotis ( 1500)
. 6. ,
Rsa
345
___________________________________________________________________________
. Rosa .
.
, [3].
Rsa
. ,
(. 3). , , ,
, , , , , , ; , .
. , , .
, , ,
, , , .
.
, .
.
. , ,
, .
. , .
(. 4).
3
Rsa,
/
1
2
3
4
5
6
7
8
9
10
11
12
R. maximowicziana
R. amblyotis
R. cinnamomea
R. glauca
R. pendulina
R. canina
R. acicularis
Rosa nutkana
R. sibrica
R. foetida
R. rugosa
R. woodsii
.
,
51,382,17**
40,961,64**
54,052,29**
62,723,43
51,012,22**
67,662,48
76,763,31
43,261,77**
47,462,08**
45,562,21**
44,732,38**
52,831,73**
. - . ,
,%
49,774,0
51,442,31**
20,721,87
66,602,62
26,952,50
66,972,55
41,722,79
59,982,28
33,342,48
60,731,59
39,312,07
63,321,51
40,054,48
66,382,95
24,511,95
64,012,28
29,252,97
62,531,66
41,761,53
52,041,26**
20,771,70
68,162,52
27,552,47
66,581,530
103,6311,02*
78,942,05**
93,537,34**
120,446,10*
99,872,88**
137,713,22
125,984,69
82,602,02**
92,274,87**
115,502,13**
86,662,24
88,302,71**
* 0,95; ** 0,99
( R. canina)
346
________________________________________________________________
4
Rsa,
1
2
3
4
5
6
7
8
9
10
11
12
R. maximowicziana
R. amblyotis
R. cinnamomea
R. glauca
R. pendulina
R. canina
R. acicularis
Rosa nutkana
R. sibrica
R. foetida
R. rugosa
R. woodsii
, .
2,40,15*
2,150,11
2,00
2,00
1,90,07
2,00
2,00
1,750,13
2,00
2,00
2,00
2,180,11
, .
4,450,16**
3,100,10**
3,300,15*
4,300,21*
3,500,17
3,700,21
3,400,16
3,100,10**
3,100
5,100,10**
2,700,15
3,360,17
V .
1147,21293,18**
480,6953,55**
1028,71142,56**
975,02113,85**
967,4796,34**
3798,02207,20
2688,58351,84**
1412,74264,85**
993,72160,95**
547,3537,28**
847,82100,97**
1154,61144,33**
1,490,191
2,320,190
2,350,329
3,330,479*
2,290,212
1,980,313
1,750,189
1,540,206
1,760,123
3,030,482*
1,830,235
1,780,287
* 0,95; ** 0,99
( R. canina).
Rosa ,
. . .
(R. amblyotis, R.
cinnamomea, R. pendulina, R. acicularis, R. rugosa, R. nutkana, R. canina, R. sibrica, R.
woodsii) (R. maximowicziana, R. foetida, R. glauca);
(R. cinnamomea, R. pendulina, R. acicularis, R. rugosa, R. nutkana, R. canina, R.
sibrica, R. woodsii, R. maximowicziana, R. glauca), (R. foetida, R.
amblyotis).
, .
(
), . [21].
R. maximowicziana, R. acicularis, R. foetida. R. canina, R. glauca,
R. pendulina .
.
R. foetida, R. glauca.
(R. rugosa) ,
.
() .
1. .. /
.. / . .. . .: . , 1988. 208 .
2. , .. / .. . - .:, 1989. - 204.
3. .. (POACEAE) // . 2009.
. 94, 8. . 1204-1215.
347
___________________________________________________________________________
4. .
/ .. , .. , .. , .. // . - 2007. 4. - . 9194.
5. .. /
.. // . 2009. 7. . 17-18.
6. . . / . . // ,
, . , 2001. - . 99-100.
7. , ..
/ .. , .. // ( - . 1922 2006). : , 2006. . 344.
8. .. / ..
, .. : , 1991. 307 .
9. ..
/ .. , .. , .. //
. 2010.
. 42, 1. . 3-21.
10. .. / .., .. // I . . , 2003. 2. 83.
11. .., .. ..
Populus (Salicaceae) // . 2002. - . 87, 1. - 63-72.
12. , .. HAMAMELIDACEAE / .. , .. // . 2010.
. 95, 3. . 338-345.
13. .. - Ribs americanum
Mill. . . 2010. 20.
14. .., .. -
Ribs americanum Mill. // : , , , .-. ., .: 2008. .133-135.
348
________________________________________________________________
631.541:631.811.98
..
. .. , ,
127276 . ,
. , 31
e-mail: bondo-irina@yandex.ru
.
,
. ,
.
: , ,
, .
,
, . ..
(, , 1976; , , 1999; ,
, , 2009).
. , , .
.
Abies concolor (Gordon ex Glend) Lindl.ex
Hildebr ., Pinus nigra Arn., Larix sibirica Ledeb., Picea omorica (Panic) Purk.
, , ,
- .
- . (, ,
, 1998).
, 38722594 .
- .
, .
60 :
S
n1
K
n2
(1)
349
___________________________________________________________________________
%, R ;
S , ;
K ,
;
n1 , ;
n2 ,
.
,
R>0. R ,
. R=0 .
, , R>0, .
(, 1984).
, . .. () - ,
, -, ,
.
1 , 30 30 ( 60 )
.
, P.omorica ( ) . (60 )
52,5% , 21,3% ,
(31,2%). , 50,1% 31,2% .
- - , .
(96,4%) (47,4%).
1
( 30 30 )
Picea omorica
Larix sibirica
Abies concolor
Pinus nigra
60 (%)
31,20,28
50,10,27
52,50,25
31,20,20
41,250,25
92,30,21
94,30,29
96,40,27
93,80,50
94,20,32
31,90,47
43,70,28
47,40,36
36,00,43
39,750,38
6,60,17
43,70,25
25,40,26
30,60,22
26,570,23
, R (%)
18,9
21,3
0
13,4
2,0
4,1
1,5
2,53
11,8
15,5
4,1
10,46
37,1
18,8
24,0
26,63
350
________________________________________________________________
* .
, 60 , 92,3%
. (94,3%) (93,8%).
P.nigra ( ). 6,6% . . 43,7% ,
37,1% , . , ,
.
, , A. concolor (
),
.
(43,7%), (25,4%)
(. 1).
, ,
, . (. 1) , , - L. sibirica. 60- 92,3% .
P. nigra (6,6%). ,
A.concolor (31,9%) P. omorica (31,2%).
,
, .
, ( ,
), ( ). ,
, -
,
.
, , ,
. . 1 ,
A. concolor. , R, . , , 100%, 0 ,
0 .
, ,
.
351
___________________________________________________________________________
. 1.
Larix sibirica: 1 - ; 2 ;
3 ; 4 ,
, (R = 21,3%),
(R = 4,1%).
. , , , 60 6,6% (. 1). (. 1). 60
92,3% .
,
, , ,
.
R=1,5%, R=2,0% R=4,1%.
,
, . , , ,
,
, -
352
________________________________________________________________
. , 60
,
Larix sibirica - 94,2%, Picea omorica 41,25%, Abies concolor
39,75% Pinus nigra 26,57%. ,
Pinus nigra (R .=26, 63%), Picea omorica (R .=13,4%), Abies concolor (R .=10,46%) Larix sibirica (R .=2,53%).
, , , , ,
. , .
,
. , (. 1, ), .
(. 2).
2
1
( 1 ; 2 )
1 ( )
2
( )
12008
F'
F
(P = 95%)
, %
15
800,53
746,02
248,68
4,23
3,86
6,21
10732,83
3577,61
60,85
3,86
89,38
529,15
58,79
100
4,41
= 50,44 1,08%.
: m = 1,08%.
: = 2,1%<5%.
, (. 1), .
(.2)
, . 2 , 1
.
.
2,
(F2 = 60,85),
(F'2 = 3,86) 95% .
. ,
353
___________________________________________________________________________
, F1 = 2,55,
, F'1 = 3,86.
. ,
. 89,38%
12008, 100%. 6,21%
( )
4,41%.
, , . , , , , .
, , ,
, . ..
.
= 2,1%, 5% .
, ,
, , : - .
3. , ,
, .
, , , ,
, ,
, , , .. . , R= -9,9%,
R= -5,0%.
3
(1 30 )
Picea omorica
Larix sibirica
60 , %
47,40,44
51,00,51
52,70,49
51,50,31
50,40,35
39,50,20
48,750,38
69,70,30
92,70,39
79,40,44
72,00,33
84,90,33
64,00,38
77,120,36
, R (%)
3,6
5,3
4,1
3,0
-7,9
1,62
23,0
9,7
2,3
15,2
-5,7
8,9
354
________________________________________________________________
. 3
Abias concolor
Pinus nigra
62,70,55
69,00,79
77,40,37
64,00,37
82,60,63
57,70,50
68,900,54
62,60,70
69,90,44
71,00,58
78,90,47
69,70,45
52,70,24
67,470,48
6,3
14,7
1,3
19,9
-5,0
7,44
7,3
8,4
16,3
7,1
-9,9
5,84
* 5 .
(52,7%), (92,7%), (82,6%) (78,9%).
, , ,
, , .
, (. 1)
, (. 3), ,
: P.omorica, L.sibirica A.concolor, P.omorica. , (P. omorica, L. sibirica, A. concolor) (.1),
, (L.sibirica A.concolor), P.nigra P.omorica (.3).
,
, (R = 23,0%)
, - (R = 19,9%)
, (R= 16,3%) (R=5,3%).
, , , . 60 77,12% , 68,90%,
67,47% 48,75%. 8,90% 1,62% . ,
(, 69,7%) (R. = 8,90%), (, 47,4%) (R. = 1,62%).
, .
, . ,
355
___________________________________________________________________________
, , - .
4
3
( 1 ; 2 - )
1
2
3725,95
19
196,10
2344,01
781,33
20,15
3,49
62,92
916,69
229,17
5,91
3,41
24,60
465,26
12
38,77
F' (P = 95%)
100
12,48
= 66,55%.
: m= 1,39%.
: = 2,09%<5%.
4.
(F1=20,15) (F'=3,49),
, . , , - , , .
.
F2=5,91 , F' =3,41, ,
, .
, , , ,
. , ,
. ,
, .
1. .. . .: .
- 1984. 424 .
2. ., .., .. //
:
. 2009. . 1. - . 9-23.
3. ., ., .. // .: , , . .
1998. . 151-152.
356
________________________________________________________________
4. ., . , // .: I . , 90-
- .. . .: , , . .. . 1999. . 183-184.
5. .., .
//. 1976. .5-6. - 3. .74-83.
In this article we can see data on regeneration of four coniferous arboreal plants after their cut surface has been treated
with different PAS. The conclusion has been made that during different terms of vegetation species react differently to the same
PAS. It has been stated that some PAS reveal themselves as inhibitors.
Key words: PAS, coniferous arboreal plants, stimulator,
ingibitor.
357
___________________________________________________________________________
UDC 577.182.22:543.544.123
R.S. Othman
V.I. Deineka
Belgorod National
Research University,
308015 Belgorod, Russia,
Pobeda str., 85
e-mail: razhanchemistry@yahoo.com
Introduction
Amoxicillin ((2S,5R,6R)-6-{[(2R)-2-amino-2-(4-hydroxyphenyl)-acetyl]amino}-3,3dimethyl-7-oxo-4-thia-1azabicyclo[3.2.0]heptane-2-carboxylic acid) belongs to a class of penicillin antibiotics that include also ampicillin, piperacillin, ticarcillin and some others.
NH2
NH
HO
S
N
O
OH
O
Amoxicillin
358
________________________________________________________________
Materials and methods
Materials
In the present work were used amoxicillin (Sigma), acetonitrile (HPLC-gradient
grade, Panreac, Spain); distilled water, phosphoric acid (Vecton, RF) and NaOH (REACHIM,
RF) of a reagent grade.
The amoxicillin brands Acamoxil 500 (Iraq), Glomox 500 (U.A.E.) and Cimoxil500 (CHAPHAR, China) were purchased in local pharmacies in Iraq while
() was purchased in Belgorod.
Instrumentation
HPLC method of amoxicillin determination was performed and amoxicillin concentrations were determined by an isocratic high-pressure liquid chromatographic system that
included high pressure pump Beckman 110 B, Rheodyne model 7125 injector with 20 l sample loop, 2504 mm Reprosil-Pur C18-AQ or 250 4 mm Reprosil-Pur C8 columns, Nicolet
LC/9563 Variable UV Detector. Detector signal was processed by MultiChrom 1.5 data management program.
UV-vis spectra were recorded in quartz cuvettes by spectrophotometer SPh-26 (RF).
The acidity of the mobile phase was controlled by pH-meter pH-150 with a combination
electrode.
Preparation of the standard solution of amoxicillin
Precise mass sample of the amoxicillin standard powder (in region 0f 9 11 mg) was
quantitatively transferred into 10 ml volumetric flask; then 4 6 ml of mobile phase were
added and after the sample dissolution the flask volume was made up by the mobile phase.
The solution was filtered through syringe filters (0.45 m). The solution was diluted to get
the calibration solutions with desired concentration (0.025 0.10 mg/ml).
Sample preparation of drug amoxicillin
Some 5-8 tablets of the drug were finely dispersed in a porcelain mortar. Then a precise
mass sample of the powder was quantitatively transferred into 10ml volumetric flask; 4 6 ml of
mobile phase were added and after the sample dissolution the flask volume was made up by the
mobile phase. Before injection the solution was filtered through syringe filters (0.45 m).
HPLC procedure
For mobile phase preparation mix 12.5 ml of acetonitrile, 5 ml 0.5 M sodium phosphate buffer (pH=5) and some 100-150 ml of distiller water in the 250 cm3 volumetric flask
and make up the volume with distilled water. Then equilibrate the chromatographic system
with 250 4 mm Reprosil-Pur C18-AQ with a guard column (effluent monitoring at 228 nm)
before sample injection.
The result of amoxicillin determination calculate as mean value of two sample preparations for each of them taking the mean value for two injection, using a calibration curve for
amoxicillin concentration calculation.
Results and discussion
Stationary phase choice
Amoxicilline is a highly polar compound thus the reversed-phase mode of HPLC
seems to be the most suitable for the chromatographic determination. Meanwhile there are
some types of reversed-phase stationary phases available differing by the length of the alkyl
radical chemically bound to the silica surface. The most popular are series of trademarks of
octadecyl (ODS ore C18) phases, proposed for amoxicillin determination [1, 3-5, 7,8]. Some
less popular are octyl (C8) phases, though just a Zorbax SB-C8 column was found to be more
suitable (than Zorbax SB-C18) for quantitative determination of amoxicillin in complex mixture with possible degradation products [5].
The background of a stationary phase choice may be proposed by the following way.
The solute retention in HPLC belongs upon some parameters (eq.1).
359
___________________________________________________________________________
nsp (i)
nmp (i)
N ( sp)
nmp (i) / N (mp ) N (mp )
X sp (i)
X mp (i)
K th (i)
(1)
k A (i)
k B (i)
k A (i)
( A)
,
( B)
( A)
( B)
k B (i)
(2)
a k B (i).
(3)
t R (i) A
a0
a1 t R (i) B
(4)
lg k A (i )
lg a
lg k B (i ).
(5)
So, the retention times of the solute i for column A vs that for column B should be a
linear function (eq. 4) for some mobile phase compositions of any solvent system. The linearity of the dependence for logarithms of capacity factors with slope closed to one is more convenient for the analysis.
If stationary phases under investigation are not chemically equal no linearity will be
found between retention times or equation (5) should be replaced by equation (6) with a
slope deviating from 1.
lg k A (i)
a0
a1 lg k B (i).
(6)
Experimentally for columns with C8 and C18 stationary phases of Reprosil trademark
a linear relationship for the retention times of amoxicillin were found with intercept closed to
zero. This finding indicates that both phases are chemically equivalent it means that mechanism of retention of amoxicillin is hydrophobic repulsion of the solute onto sorbent surface. Indeed in this case the length of alkyl radical of chemically modified silica could not influence upon solute retention.
Chromatographic columns 2504 mm Reprosil-Pure C8-AQ and Reprosil-Pur C18.
Mobile phases 2.0 5.0 vol.% of CH3CN in water phosphoric buffer (0.01 M
NaH2PO4), 1 ml/min.
As the consequence of the fact C18 stationary phases may be readily replaced by C8
or even C4-phases for the given mobile phase composition. By the way for the two reversedphase stationary phases built upon the same silica matrix dead times may be closed then the
intercept of the equation (4) must also be close to zero.
In spite of the findings column with C18 stationary phases has been explored for the
further method development.
360
________________________________________________________________
(7)
A composition, containing 5 vol.% of CH3CN has been chosen for amoxicillin determination.
Column: 250 4 Reprosil-Pur C18-AQ; mobile phase 0.01 M phosphate buffer pH=5
361
___________________________________________________________________________
2. Low organic modifier concentration in the mobile phase may course phase collapse
[9] for the ordinary reversed phases. Thus phases sustained in aqueous solutions are preferable. Stationary phases of Reprosil trademark agree the requirement.
3. Chromatographic determination of amoxicillin as ionisable compound must be performed in buffered mobile phases. The choice of buffering system is determined by UVspectrum on the amoxicillin solution enabling a simple and sensitive detection. The UVspectrum is presented on fig.3.
NH3
NH
HO
S
N
O
O
mV
100
0
0
362
________________________________________________________________
Column 2504 mm, Reprosil-Pur C18-AQ,; mobile phase 5 vol. % in 0.010 M phosphate buffer (pH=5), 1 ml/min. Detection 228 nm.
Method validation
For the developed method of amoxicillin determination taking a result as a mean value for two replicate injection for two replicate sample preparations the overall relative confidence interval (P = 0.95) is composed of relative confidence intervals (CI) of method repeatability (R), sample preparation (SP), and detector calibration (DC), eq.8.
CI
CI R2
2
CI SP
2
CI DC
(8)
2.57%
Data for three series of six replicates of amoxicillin solution injections indicates a high
repeatability is no more than 0.93%. CISR for three replicate sample preparations was found
to be 1.71; CIDC (1.68) was estimated for the calibration mode utilizing three different concentrations with two replicate injections for each of them. The calibration dependence proved to
be linear with not statistically significant intercept, fig.5.
Drug trademark
1
2
3
4
Sample mass,
mg
10.3
10.2
10.5
9.6
Mass of amoxicillin
found, mg
10.24
10.56
8.66
9.55
Conformity with
stated content, %
99.53.7
103.63.8
99.53.0
99.63.7
The results obtained by the developed method proved the validity of the method and
good quality of the drugs under investigation.
363
___________________________________________________________________________
Conclussions
For the amoxicillin assay in drugs the RP HPLC method may be explored. The stationary phase must be collapse phase resistant while the length of the alkyl-radical of modified
silica is not critical. The mobile phase for isocratic HLPC assay consists of 5 vol.% of acetonitrile in (0.01 M) water phosphoric buffer (pH=5); the rate of mobile phase 1 ml/min. Peaks
are registered at 228 nm.
References
1. Pires de Abreu L.R., Ortiz R.A.M., de Castro S.C., Pedrazzoli J. Jr. HPLC determination of
amoxicillin comparative bioavailability in healthy volunteers after a single dose administration // J.
Pharm. Pharmaceut. Sci. 2003. V.6. P. 223-230.
2. Alemzadeh I., Borghei G., Vafi L., Roostaazad R. Enzymatic Synthesis of Amoxicillin with
Immobilized Penicillin G Acylase // Transactions C: Chem. Chem. Engineer. 2010. V.17, No.1. P.
106-113.
3. Moore T.D., Horton R., Utrup L.J., Miller L.A., Poupard J.A. Stability of AmoxicillinClavulanate in BACTEC Medium Determined by High-Performance Liquid Chromatography and Bioassay // J. Clin. Microbiol. 1996. V.34. P. 13211322.
4. Hsu M.-C., Hsu P.W. High-Performance Liquid Chromatographic Method for Potency Determination of Amoxicillin in Commercial Preparations and for Stability Studies // Antimicrob. Agents
Cemotherapy. 1992. V.36. P. 1276-1279.
5. Raju Ch.B.V.N., Sharma H.K., Rao Ch.S., RAO G.N. RP-HPLC Method for Analysis of Related Substances in Amoxicillin Drug Substance // Acta Chromatogr. 2009. V.21. P. 5770.
6. Doua M., Hosmanov R. Rapid determination of amoxicillin in premixes by HPLC // J.
Pharmaceut. Biomed. Anal. 2005. V.37. P. 373-377
7. Nikam D.S., Bonde C.G., Surana S.J., Venkateshwarlu G., Dekate P.G., Development and
Validation of RP-HPLC Method for Simultaneous Estimation of Amoxicillin trihydrate and Flucloxacillin sodium in capsule dosage form // Intern. J. Pharm. Tech. Res. 2009. - V.1, No.3. P. 935-939.
8. Rose M.D.; Tarbin J.; Farrington W.H.H.; Shearer G. Determination of penicillins in animal
tissues at trace residue concentrations: II. determination of amoxicillin and ampicillin in liver and
muscle using cation exchange and porous graphitic carbon solid phase extraction and highperformance liquid chromatography // Food Additives Contamin. Part A: Chem. Anal. Control, Exposure Risk Assess. 1997. V.14. P. 127 133.
9. Matthew Przybyciel, ES Industries, West Berlin, New Jersey, USA, and Ronald E. Majors,
Phase Collapse in Reversed-Phase LC // LCGC Europe. 2005. October. P. 2-5.
..
..
, 308015 ,
, . .85
e-mail: razhanchemistry@yahoo.com
-
.
.
Reprosil-Pur C18-AQ (pH=5) 5% (228 nm). .
: - ,
, .
364
________________________________________________________________
615.322:543.544.123
1
..
..
..
..
C..
, 308015 ,
, . .85
e-mail: deineka@bsu.edu.ru
Ach87@mail.ru, Deyneka@bsu.edu.ru
sorokopudov@bsu.edu.ru
shevchenko_s@bsu.edu.ru
.
: -, 6 -, 2 , . ,
, Cerasus mahaleb 550 750 100
.
: , , Cerasus sp.,
Prunus avium, Prunus padus.
[1, 2].
[2]. ( , ) [3], ( ), [2].
, ,
- [4]. , .
, [4], ,
: 1) -3- (Cy-3-Glu), 2) -3-(2-) (Cy-32GRut), 3) -3- (Cy-3-Sopho), 4)
-3- (Cy-3-Rut) , , ; .
( 28 80 100
) [2]. [5]
, .
, , , , , .
1
365
___________________________________________________________________________
OH
O
HO
OH
HO
OH
OH
O
OH
HOCH2
OH
OH
OH
HOCH2
OH HO
OH
OH
Cy-3-Glu
O+
OH
OH
O
HO
OH
O
OH
OH
OH
CH3
HO
OH
OH
OH
OH
OH
O
OH
CH2OH
Cy-3-Sopho
OH
HO
OH HO
CH3
HO
Cy-3-Rut
OH
CH2OH
OH
OH
OH
OH
Cy-3-2GRut
, , .
, .
, , . , , 0.1 HCl. 0.5 3 , , (
), .
, -3- [6].
18. 5 ,
15-20 0.1 HCl.
(10-20 ). 2 0.1
HCl. , 20 .% . 1-2 . 2 .
. Agilent 1200
Infinity c - ( 370 600 )
515 ; 2504 Reprosil-Pur C18-AQ, 5 ;
: 10 .% ( ) 10 .% .
366
________________________________________________________________
[7, 8].
:
1) (RT), 6 ( ) :
Cy-3-Glu + Rhamn Cy-3Rut,
Cy-3-Sopho + Rhamn Cy-32GRut.
2) (2GT), 2 :
Cy-3-Glu + Glu Cy-3-Sopho,
Cy-3-Rut + Glu Cy-32GRut.
3-, :
Pg(Gly) Cy(Gly),
, .
Cy(Gly) Pn(Gly).
,
, ,
, , , - [9]. .
, . .1. , .1 .
-, , , - 3 , , , [8].
-3. -3-
, [10].
-, [11],
.
, , ( , ) ( ) , .2. , , () ,
10%- 0.5 18.
1
.
, , , Cy-3-Rut,
RT 2GT. .
367
___________________________________________________________________________
,
- Cy-32GRut.
Cy-32GRut
mV
Cy-3-Sopho
Cy-3-Rut
500
Cy-3-Glu
()
250
10
.1.
1
4
2
5
.2. 3- (1),
(2) (3), (4) (5)
368
________________________________________________________________
1
*, %
Cy-3Pg-32GRut
Sopho
Glu
Rut
Glu
Rut
, Prunus cerasus L. Cerasus vulgaris Mill.
10.8
63.5
2.2
12.4
0
0
1.2
64.9
6.0
23.3
0
0
2.6
62.8
4.8
25.9
0
0
<0.5
63.6
8.9
23.7
0
0
1.1
62.4
5.8
25.4
0
0
3.0
4.4
4.4
85.5
0
0
<0.5
4.6
5.5
81.8
0
0
<0.5
<0.5
2.0
83.1
0
0
<0.5
3.5
6.3
83.2
0
0
<0.5
<0.5
1.9
79.8
0
0
, Prunus avium
0.8
0.9
13.0
78.6
0
0
, Prnus pdus
4.5
<0.5
33.7
59.1
0
0
<0.5
<0.5
7.7
70.4
0
0
, Cerasus mahaleb
<0.5
<0.5
43.7
51.6
<0.5
0
() Cerasus
3.3
59.7
4.2
22.2
0
0
fruticosa
, Cerasus fruticosa
<0.5
6.1
4.7
82.3
0
0
, Cerasus tomentosa
<0.5
<0.5
2.8
12.4
4.3
77.0
/
Pn-3Rut
0
0
0
0
0
0
0
0
0
0
9.1
1.6
4.0
3.7
2.3
2.7
8.1
14.9
7.0
18.3
3.5
0
18.8
3.1
1.8
3.1
4.7
6.7
0
0
6.8
3.5
:
1) RT,
, :
2) 2GT, , 2:
2 ,
RT, y-3-Rut Cy-3-2GRut. , , , : ,
, , , , ,
, , , , , , 2GT 84 44 %.
, ( ) , , Cy-3Rut. 2GT: , , -
369
___________________________________________________________________________
, ; . : , , , .
, -3 , .1, .
0.030
0.160 100 .
(360 100 ),
(180 100 ).
550 750 100 , , (, !,
).
2
1
2
3
4
5
6
7
8
9
10
11
12
13
14
Cerasus vulgaris
Cerasus
, Cerasus mahaleb
(), Cerasus
fruticosa
, Cerasus fruticosa
, %
RT
2GT
85.4
87
92.5
71.5
92.3
70.9
90.7
70.1
92.7
68.5
92.4
8.01
94.0
4.89
97.6
0
93.0
0
97.7
0
90.1
0
54.1
0
*
0.025
0.114
0.102
0.117
0.086
0.058
0.05
0.069
0.12
0.025
0.193
0.629
91.6
68.8
0.103
95
6.42
0.156
* - -3-.
1. Lila M.A. Anthocyanins and Human Health: An In Vitro Investigative Approach // J. Biomed. Biotechnol. 2004. V.2004. P. 306313.
2. Blando F., Gerardi C., Nicoletti I. Sour Cherry (Prunus cerasus L) Anthocyanins as Ingredients for Functional Foods // J. Biomed. Biotechnol. 2004. V.5. P. 253258.
3. Wang H., Nair M.G., Strasburg G.M., Chang Y.-C., Booren A.M., Gray J.I., DeWitt D.L. Antioxidant and Antiinflammatory Activities of Anthocyanins and Their Aglycon, Cyanidin, from Tart
Cherries // J. Nat. Prod. 1999. V.62. P. 294-296.
4. imuni V., Kova S., Gao-Soka D., Pfannhauser W., Murkovic M. Determination of anthocyanins in four Croatian cultivars of sour cherries (Prunus cerasus) // Eur. Food Res. Technol.
2005. V.220. P. 575578.
5. Chaovanalikit A., Wrolstad R.E. Anthocyanin and Polyphenolic Composition of Fresh and
Processed Cherries // J. Food Sci. 2004. V.69. P. FCT73-FCT83.
6. Giusti M., Wrolstad R.E. Characterization and Measurement of Anthocyanins by UV-Visible
Spectroscopy // Current Protocols in Food Analytical Chemistry. 2001. - F1.2.1-F1.2.13.
7. .., .., .., .. Rubus L. . //
. - 2005. - 4. - . 61-65.
370
________________________________________________________________
8. .., .., ..
// . : .
- 2008. - 7(47), .7. - . 71-78.
9. .., .., .., .., ..,
.. Rosaceae //
. - 2005. - .1. - . 91 -98.
10. .., .., .., .., .., .. // . .
. 2009. .9, .4. . 529-536.
11. .., .. . // . . . - 2004. - T. 78, 5. - .923-926.
371
___________________________________________________________________________
615.322:615.074
1
..
..
..
..
..
, 308015 ,
, . .85
e-mail: deineka@bsu.edu.ru
prizrac1@yandex.ru
Deyneka@bsu.edu.ru
sorokopudov@bsu.edu.ru
dzelu@yandex.ru
20
- . , 5.5 24.0 % , ;
: - . - - .
: Hippophae rhamnoides, ,
,
(Hippopha rhamnoides
L.) , [1].
[2], , ,
8-9 . %,
, , , , , , , , , . [1, 3].
[4, 5], . [4 - 9]. [4] , 20.76 % ( 425 % - ); - 14.7 36.9 % ,
.
168 1089 %. -
, -
, .
, ,
. ,
, [5], -, -
-, , -- .
[6] - (18.4 %),
- (15.8 %), (14.2 %) (12.4 %), - (7.3 %), (6.0 %) (5.8 %).
.
, - - .
1
- 2009-2013 , -174.
372
________________________________________________________________
, [7],
, , -, -
- .
[8] , . , [9], ,
, ,
, ; - 2.5 ,
( 6.5 ), .
( )
,
[10]. , , , , [11].
, ,
, , [12]. , .
,
. (0.5 )
,
(5-10 ). 25 3.
.
( -56, ) . - .
- .
. Agilent 1200
Infinity c - ( 370 600 ) 450 . -
2504 Reprosil-Pur C18-AQ, 5 ; : 20 .% ( )
, 1 /. - 2504
300, 5 ; 30 .% -, 1 /.
(ALOGPs 8.30 11.01, [13]), ,
.
[14] (
) - - .
, - , .1.
2
1
373
___________________________________________________________________________
.1.
- . : 2504 300, 5 ,
: 30 .% , 1 /; 450
( 1 - 5), ( 6), ( 7)
( 8). 6 7 , , , .2,
[15].
II
.2. 6 7 .1. ,
30 .% -
374
________________________________________________________________
.3. 1 .1. ,
30 .% -
- , ,
. .. -, . , , , .
,
.
2, .4.
.4. 2 .1
, -, , . ,
2 , .
5 , .5.
, ,
. 3 4 , ( ) , .6.
375
___________________________________________________________________________
.5. 5 . 1
CH2OH
.6.
, 2010 ,
.1
20
Car.
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
1
2
2
1
2
278439
13.8
15.2
17.6
19.3
21.5
22.2
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* - .
1. Zeb A. Important Therapeutic Uses of Sea Buckthorn (Hippophae): A Review // J. Biol. Sci.
2004. V.4. P. 687-693.
2. .. . .: , 2005. 1200 .
3. Kumar R., Kumar G.P., Chaurasia O.P., Singh S.B. Phytochemical and Pharmacological Profile of Seabuckthorn Oil: A Review // Res. J. Med. Plant. 2011. V.5. P. 491-499.
4. Bekker N.P., Glushenkova A.I. Components of certain species of the Elaeagnaceae family //
Chem. Nat. Comp. 2001. - V.37. P. 97-116.
5. Sergeeva N.V., Shapiro D.K., Bandyukova V.A., Anikhimovskaya L.V., Narizhnaya T.I. Carotenoids of the fruit of some varieries of Hippophae rhamnoides // Chem. Nat. Prod. 1979. V.5.
P.87-88.
6. Kudritskaya S.E., Zagorodskaya L.N., Shishkina E.E. Carotenoids of the sea buckthorn, variety Obilnaya // Chem. Nat. Comp. 1989. V.25. P. 724-725.
7. Pintea A., Varga A., Stepnowski P., Socaciu C., Culea M., Diehl H.A. Chromatographic Analysis of Carotenol Fatty Acid Esters in Physalis alkekengi and Hippophae rhamnoides // Phytochem.
Anal. 2005. V.16. P. 188195.
8. Parlog R.M., Vodnar D.C., Dulf F.V., Leopold L., Socaciu C. HPLC-PDA and UV-VIS spectrometry analysis used to fingerprint sea buckthorn (Hippophae rhamnoides L.) berries comparatively
with leaves and seeds extracts // Bull. UASVM. Agric. -2099. V.66. P. 409-414.
9. Andersson S.C., Olsson M.E., Johansson E., Rumpunen K. Carotenoids in Sea Buckthorn
(Hippophae rhamnoides L.) Berries during Ripening and Use of Pheophytin a as a Maturity Marker //
J. Agric. Food Chem. 2009. V.57. P. 250258.
10. Krinsky N.I., Landrum J.T., Bone R.A. Biologic mechanisms of the protective role of lutein
and zeaxanthin in the eye // Annu. Rev. Nutr. - 2003. V.23. P. 171201.
11. Weller P., Breithaupt D.E. Identification and Quantification of Zeaxanthin Esters in Plants Using
Liquid ChromatographyMass Spectrometry // J. Agric. Food Chem. 2003. V.51. P. 7044-7049.
12. .., .., .., .., ..
/ ,
: . . . / . .. .
: , 2009. . 64. - . 272-274.
378
________________________________________________________________
13. Tetko I. V., Gasteiger J.. Todeschini R., Mauri A., Livingstone D., Ertl P., Palyulin V. A., Radchenko E.V., Zefirov N.S., Makarenko A.S., Tanchuk V.Y., Prokopenko V.V. Virtual computational
chemistry laboratory - design and description // J. Comput. Aid. Mol. Des. 2005. V.19.
P. 453-63.
14. .., .., ..
. // . . . 2009. .9, .3. . 374-382.
15. .., .., .., .., .., ..
// . . - 2010. - 21 (92), .13.
. 143-148.
16. Rodriguez-Amaya D.B., Kimura M. HarvestPlus handbook for carotenoid analysis.
HarvestPlus Technical Monograph Series, 2. 2005. - 57 p.
17. .., .., .., .., ..
// . : . 2010. 15(86). .12. . 140-144
379
___________________________________________________________________________
615.322:543.544.123
(
- )
haenomeles japonica C. maulei. , 3- 3 ,
-3- (F-3-H),
.
: haenomeles japonica, C. maulei, , , -.
[1],
Chaenomeles Lindl. ( )
, . : (I, C. japonica (Thunb.) Lindl. ex Spach.), (II, C. speciosa (Sweet) Nakai) (III, C. cathayensis (Hemsl.) Schneid.) [1, 2],
C. thibetica. : C. superba (Frahm) Rehder;
C. japonica C. speciosa, Superba, C. clarkiana Weber (C. cathayensis C. japonica, Clarkiana), C. vilmoriniana Weber (C. cathayensis C. speciosa, Vilmoriniana) C. californica Clarke ex Weber [C. cathayensis (C. superba), Californica]. : I, III, ( , C. maulei (Mast.) C.K. Schneid), (C. sinensis
(Thouin) Koehne) [3], C. maulei . japonica
[4], C. sinensis
.
Chaenomeles ,
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, , , ,
[5], .
II [6], C. sinensis
, [7].
2002
: C. japonica C. maulei.
( - )
( ).
.
, C. japonica C. maulei , 1
380
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positive, fragm. 200V). 4.6250 Symmetry 18; 10 .% 8 .% , 1 /.
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490 500 , ,
.
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( ).
.2 ,
[11], ( ) -3-, Dp-3-Glu, -3-, Dp-3-Rut,
-3-, Cy-3-Glu, -3-, Cy-3-Rut.
381
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. 1. Chaenomeles sp.
: C. japonica 1, 2, 5; C. maulei 3, 4
Cy-3-Rut
Dp-3-Rut
Dp-3-Glu
Cy-3-Glu
5
A
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; ;
1 Cy-3-Hex-Gala; 3 Cy-3-Gala; 4 - Pg-3-Hex-Gala; 5 Pg-3-Gala; 2, 6, 7 .
,
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,
-3-, Cy-3-Gala, Cy-3-Glu, -3, Cy-3-Ara, -3-, Cy-3-Xyl, [9], 3 Cy-3-Gala, .
3 ,
3 , Cy-3-diGly. , 4 5 -3-
Pg-3-diGly Pg-3-Gala. , .4.
1,2,3,7
4,5,6
382
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. 4. - 1,2,4 5 . 2
383
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, (
).
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;
, 5:
(1)
Si(Cy) ;
Si(Cy+Pg) .
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Pg-3-Hex-Gala
-3-Gala
Pg-3-Gala
. 5.
. maulei ( ) C. japonica ()
1
Chaenomeles
. japonica
C. maulei
2-14
2-01
5-29
6-8
6-5
6-1
(F3H),% %Hex
62.8
72.2
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384
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, . japonica C. maulei.
-3- [14] 120 180 100 ,
C. japonica (5-29) 400 . ,
.
385
___________________________________________________________________________
Spectrophotometry and HPLC-MS were explored for the determination of main anthocyanins of haenomeles japonica and C. maulei
petals. It has been established that all samples under investigations were
composed of 3-galactosides as well as 3-hexosilgalactosides of cyanidine
and pelargonidine. The main anthocyanin composition differences were
due to relative activity of flavonoid-3-hydroxylase being the base for differentiation of the Chaenomeles species.
Key words: haenomeles japonica, C. maulei, flowers petals, anthocyanins, HPLC-S.
386
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51-72:530.145
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e-mail: Chekanov@bsu.edu.ru
2
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2. .. // . 1987. .32. . 1316-1321.
3. .. // . 1978. .75, .4. . 1261-1268.
4. Gutzwiller M.C. Chaos in Classical and Quantum Mechanics. New York: Springer, 1990. 432 p.
5. Henon M. Integrals of the Toda lattice // Phys. Rev. 1974. Vol. 9, 4. P. 1921-1923.
The receiving procedure of classical Hamiltonian to the BirkhoffGustavson normal form is described. With help of the Weyl correspondence
rule for the classical normal forms of some integrable and nonintegrable systems their quantum counterparts are constructed and approximated energy
spectra and wave functions are found. It is shown that thus obtained energy
spectra are represented good exact ones in an energy domain where the classical motion is regular but agreement is worsen strongly at the energy domain
where the classical regular motion is going into chaotic one. It is established
that .
Key words: classical Hamiltonian, Birkhoff-Gustavson normal
form, Weyl correspondence rule, energy spectra, wave function, method of
quantization.
392
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