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O As early as 5,000 B.C., hybrid production of OIn 1933, < 1% of corn planted in the U.S.
corn was indirectly encouraged by Indian tribes represented hybrids. During this period the
interested in the semi-controlled mixing of development of inbreds, and crossing with
endosperm colors during religious ceremonies other inbreds to produce double-cross hybrids
O These attempts not focused on high yields and resulted increased grain yields by 10-40%
it wasn’t until the early 1900’s that breeders over traditional OP varieties.
developed a method of self-fertilization to
select homozygous inbreds O # of dbl-cross combinations and their
O Many deleterious recessive genes were superior performance caused rapid adoption
expressed - often resulted in greatly reduced of corn hybrids by farmers.
yields while simultaneously encouraging the
expression of specific traits
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Agricultural Importance Vegetative Development
O Worldwide, corn ranks behind only wheat in O Germination of corn seeds is similar
to that of many other grasses.
total production. O Germination begins when the seed
O ~ 80% of the corn produced goes to livestock reaches a moisture content of about
30%
and poultry feed, while ~ 20% is used in food O Primary radicle is followed by
& industrial applications ( these values are seminal roots as the first organs to
changing w/ demand for ethanol, etc) emerge from the seed
O About 12 h later, the coleoptile,
O In many countries (Africa, Latin America) it is which covers the leaves, ruptures
an important crop for food as well as for the seed coat and is pushed ot the
surface by the elongating mesocotyl.
livestock feed.
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O Ear initials are found on mature plants in O Because the tip ear silks appear after
axillary buds of lower leaves up to the last basal ear silk emergence, they may not be
axillary bud that produces the ear. fertilized if pollen shed has ceased,
O Rapid growth of the ear shoots occurs resulting in a barren tip ear.
simultaneously with tassel emergence O Extreme heat or drought also have a much
and the most rapid growth is in the greater effect on the timing of silk
highest position. emergence than pollen shed or pollen
O Silk elongation is initiated first from ovules viability, and is linked with poor yields
at the base of the ear and last from those under stress conditions.
at the tip of the ear.
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O The seed is botanically classified as a
caryopsis because it is a dry (at
maturity), indehiscent, single-seeded fruit
that includes the tightly fused ovary wall.
O Corn seeds differ dramatically in size and
shape.
O Position on the ear is primarily
Black layer: responsible for these differences
O Large round seeds tend to be at the base
of the ear and small round seeds at the
tip.
(Pictures from: Mississippi State University extension service, http://msucares.com/crops/corn/corn2.html)
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Flint corn primarily grown in
O
Europe, Asian and Central O Sweet corn is grown for direct human
and South America. Kernels consumption, and differs from dent corn
are smooth and rounded
with no denting and posses by only 1 recessive gene that prevents
a thick, hard, vitreous
endosperm surrounding a the conversion of sucrose to starch in the
small granular center. kernels.
O Floury corn is primarily
grown in South America and O Popcorn is a small flint type that
southern Africa.
O One of the oldest types and
possesses a hard, vitreous endosperm
possesses a soft endosperm that explodes when between 13 and 15%
with no denting.
O This corn is often ground to
moisture content after heating.
produce flour for cooking.
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Composition as Percentages of Whole and
Fractionated Corn Kernels.
Seed Production
O Corn is a warm-season crop.
Fraction Kernel Starch Sugar Protein Oil
O Breeders have developed lines that can grow in almost any
climate, but corn typically grows best when average temps of
21-32C (70-90F) and 15-20 cm (6-8 in) rainfall are
Kernel 100.0 71.5 2.0 10.3 4.8
encountered for 3-4 consecutive months.
Endosperm1 81.9 86.4 0.6 9.4 0.8
O These conditions abound in the US corn belt. Iowa,
Germ 11.9 8.2 10.8 18.8 34.5
Nebraska, Indiana and Illinois are the center of this region.
Bran 5.3 7.3 0.3 3.7 1.0
O Major corn seed companies are located in these states.
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O Agronomic practices for the production of seed corn are the
same as those for grain except that some additional
1. Composed of starchy and aleurone fractions.
requirements exist.
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O One of the greatest concerns in hybrid O Quality assurance (QA) specialists from
seed corn production is genetic purity. seed companies constantly monitor
production fields to ensure that only
O Because detasseling operations are
hybrid seed is produced and marketed.
seldom successful in eliminating all
O Field inspections provide one measure of
tassels, some level of selfing occurs.
this success.
O Contaminating pollen from other seed
fields creates problems by producing O Many companies have adopted lab
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O To preserve crop identity, a grower should use the
Identity Preserved (I.P.) Production same generally accepted practices to manage pollen
flow that are used in any of the currently grown I.P.
O Some growers may choose to preserve the
corn crops.
identity of their non-genetically enhanced corn to
O These include adequate isolation distances, buffers
minimize the potential for movement of pollen
between hybrids, border rows, planned differences in
from fields containing any genetically improved maturity between adjacent fields and natural barriers.
traits or any such traits not yet approved for O Examples: production of seed corn, white, waxy, high
export markets (such as the European Union) oil, high amylose, high lysine and other corn crops that
O Accepted practice with I.P. production is that meet specialty needs, including organic and non-
each grower has responsibility to provide any genetically enhanced specifications.
necessary crop isolation that might be required.
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O Planting pattern varies, but most common is 1
row of male parent to 4 rows female parent.
O Selection of parents must be planned so that the
O Other ways to make minor (3-6 d)
male sheds pollen at the time of silk emergence adjustments in pollen shed: use
from the female (nicking). variable fertilizer rates, growth
O A method to ensure successful nicking with regulators to stimulate or retard
differing maturities is the use of split planting development, retarding growth via
dates. clipping or flaming of one parent,
O Female planted on a different date than male so planting at differing soil depths, seed
that synchronous flowering occurs. coatings.
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Flowering
O Hybrid seed production requires the female parent
have its tassels removed or made infertile before its
silk emergence to avoid self-fertilization.
O 3 ways to do this:
(1) Manually or mechanically remove tassel - must be
done in 7-10 d between when tassel is ready to
emerge and pollination. Fields inspected to ensure
that no tassels were missed. Yield reductions of up to
10% due to detasseling. Yield reductions even
greater with mechanical removal.
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Most harvesting of corn for grain is with a
O
combine; for seed use, it is best to retain the
Drying
Because seed corn is
kernels on the cob and allow seed to dry further O
usually harvested on the
so that mechanical damage is minimized. ear and at high moisture
content, it is necessary to
O Seed should not be harvested at moisture reduce seed moisture
contents above 35-38% so it can be safely dried. content for safe storage.
O Usually done by squirrel-
O Major risk to seed quality is the possibility of cage or fan systems to
freezes, which reduce germination. draw fresh air through a
burner and force the
O Level of freeze damage is dependent on temp, heated air through bins
filled with seed on the ear.
developmental stage (moisture content),
genotype, and husk protection.
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O Most seed corn is
3 Operations in Conditioning Seed now marketed
according to size in
1. Scalping operation – using scalpers or air-screen bagged units of
cleaners to remove cob and kernels, husks, silks and
22.7 kg (50 lb) each
or providing bags
other debris. with 80,000 seeds
2. Sizing operation – separates seeds into uniform lots each which vary in
of size and shapes (width, thickness, length); width weight from 13.6 to
graders and aspirators used. 231.7 kg (30 to 70
3. Treating seed - with a fungicide-insecticide slurry lb) because of the
prior to bagging.
differences in seed
size.
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O Double-crosses were necessary because O Double-cross plants are more variable than
the first inbred lines had poor vigor and single or 3-way crosses.
performed poorly as female parents. O They are not all alike genetically and allow
O Thus, single cross hybrids had to be breeders to bring more different desirable
formed because they had enough vigor to characteristics together into one hybrid than
enable them to be good female parents. is possible in a single cross.
O This allowed hybrid seed production to be O The plants may be “buffered” more against
a practical, economical system that unfavorable situations, which frequently
produced adequate seed that could occur at one or more times during the
perform well for farmers. growing season.
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O Genetic uniformity of single-cross plants Three-Way Hybrids
offers a particular advantage for high-yield
environments, where they yield better than O Crosses can be made between a single-cross hybrid
the best double-crosses. (AxB), as the seed parent with an inbred line, C, as
O Only two inbred parents are involved, a the pollen parent to give the pedigree [(AxB)xC].
higher level of resistance to diseases, insects O Three-way hybrid seed is produced on single-cross
and unfavorable weather is evident plants so that yield and quality may be equal, or nearly
O If any component is adverse, it will affect all so, to double-cross seed.
of the single-cross plants, resulting in lower O The pollinator is an inbred parent, and this may add
performance. some cost to seed production
O Pollen shed occurs during a shorter period O More variable than single-crosses and less variable
since all the plants are genetically alike, with than double-crosses.
the potential for lower yields, especially O Advantages and disadvantages are likely to be
under stress conditions. between those double and single crosses.
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Kernel properties associated with poor
Conditioning (continued)
sweet corn emergence and seedling vigor
O Shelling, milling, sizing, sorting, treating and
packaging processes again modified for high- O Seed maturity at harvest
sugar sweet corn seed. O Pericarp injury and other damage during
O Electronic sorters remove discolored kernels harvest, drying
O Milling, sizing equipment modified to handle O Membrane damage due to rapid imbibition
seed ½ the weight and size of su1 O Solute leakage during germination
(Marshall & Tracy, 2003)
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Credits
O McDonald and Copeland: Seed Production
text
O ISU Report No. 48
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