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(INSECTA, COLEOPTERA)

03.00.09

2009

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..




..

8 2009 . 14
002.213.01 .
.. : 119071, , , 33
(495) 952-35-84
e-mail: admin@sevin.ru

: 119071, , , 33

2009 .

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..


.
Arthropoda ,
.

(Bourdeaux, 1979; Bitsch, 2002). ,

..

Crustacea Myriapoda, Hexapoda


(Apterygota). Pterygota, Coleoptera ,
, , .
, (Joshi, 1976),
(Crowson, 1938, 1944).
Coleoptera,

(Baehr, 1975, 1979; Beutel, 1992; Friedrich et al, 2006, 2009).

.
. -
Coleoptera
.
:
1)
, ;
;
;
2) ,
, ;
3)
;

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4) ,
-
Coleoptera;
5)

Coleoptera
;
6)
Coleoptera.
. 202 90
, .
,
. .
Coleoptera
,
Neuropteroidea. Coleoptera
Holometabola
.
.

.
.

Coleoptera.
,
, Coleoptera.
-
,
.
. : XIII

(,

2007);

XIII

European

Carabidologists Meeting, (Blagoevgrad, 2007),

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.
. 6 , 3
, 1 ; 1
2 .
. , 7 ,
. 189 , 4 196 .
160 , 145 .
.
.. ()
. .. ()
,
, ,
.. (), .. (), .. (), ..
(), .. (), .. (), .. (),
.. (), Killian A. (Wroclaw University, ), Pollock D.A. (Eastern New Mexico
University, ), lipiski S.A. (CSIRO Entomology, ), Horak J. (),
(-):
.., .., .. ..
.


. ,
.
1.
1.1.

Coleoptera

,
, , , ,
XIX (Eschscholtz, 1820; Baer, 1826; Audouin, 1828; Mac Leay, 1830; Faivre, 1861),
(Baehr, 1979; Ge et al, 2007).
.
(Berlese, 1909),

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. (Crampton, 1918),

Pterygota. ,
,
.
, Coleoptera,
- , (Speyer,
1922; Crome, 1957; Marvillet, 1960; Jaboulet, 1960; Barlet, 1992).
, ,
(Paulian, 1944), 14

.
, ,
Coleoptera. ,
,
.
, , ,

Coleoptera , .
1.2.
Coleoptera

Coleoptera,
(Chapius, Candze, 1853; Perris, 1877; Schidte, 1862-1883; Saalas, 1917, 1923;
Verhoeff, 1923).
,

, ..

,
,
(Bving, Craighead, 1931).

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Coleoptera
,
.

(Crowson, 1955; Rozen, 1960; Viedma, 1966; Doyen, Lawrence, 1979; Young 1976, 1985;
Lawrence & lipinski, 1999).

(
) (Bousquet, 1984; Kovarik & Passoa, 1993;
Makarov, 1996; Delgado, 1996; Solodovnikov, 2007) (Ruhnau,
1984; Beutel et al, 1993, 1998, 2000; Kiselyova, McHugh, 2006; Zaitsev, 2005, 2008).
, Coleopera
.

, ,
. , ,
,

(van Emden, 1957; Hinton, 1955, 1966).
Coleoptera,
, 80- XX .
,
Hydradephaga (Ruhnau, 1984).
.
(Beutel, 1988, 1991;
Beutel et al, 1999a, 1999b, 2000, 2006) Coleoptera,
.
; , ,
, - .

Coleoptera . ,
(Crowson, 1938)
.

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2.
202
90 Coleoptera Archostemata, Adephaga Polyphaga,
70% .
Ascalaphus sp. (Neuroptera, Ascalaphidae), Myrmeleon sp. (Neuroptera, Myrmeleontidae),
Osmylus sp. (Neuroptera, Osmylidae), Chrysopa sp. (Neuroptera, Chrysopidae), Inocellia sp.
(Rhaphidioptera, Inocelliidae), Sialis sp. (Megaloptera, Sialidae), Panorpa communis L.
(Mecoptera, Panorpidae), Tenthrediniidae gen. sp. (Hymenoptera), Allotrichia sp. (Trichoptera,
Hydroptilidae).
,
.
, NaOH
.
.
Leica MZ6.5 MZ16.5,
.
Corel Draw 12.
Statistica 8.0.

, U-
- ( - )
95% (, 2001).
3. Coleoptera
3.1.
Coleoptera
- ,
. Coleoptera
10 ,
.
(. 1).

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.
(aes1)
. (aem1),
.
Coleoptera ,
.
(apl1),
. .
,
. (f1) - ,
, .
( . 1 ),
- .
(ais1), - ,
, .
- . , ,
(
, ), .

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10

, , ,
. , ,
( ) .
(spina),
- - , ,
.
( , )
, .
.
3.2. Coleoptera

3.2.1. ( Calosoma auropunctatum Herbst)
Carabidae
. , ,
Calosoma auropunctatum Herbst
. ,
Calosoma ,
,
. ,
, - - .
3.2.2. ( Cerambycidae)
, ,
(Buprestidae, Curculionidae, Cerambycidae).

,
, .

.

, .

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11

,
(Oedeomeridae, Melandryidae, Peltidae)

, ,
. , ,
.
3.3.
3.3.1.
3-5 ,
. Coleoptera,

: ,
,
.

.
Calosoma auropunctatum Herbst., Ophonus
asureus Fabr., Carabus granulatus L., Anisodactylus binotatus Fabr., Platynus assimilis Payk.
(Carabidae) Cucujus haematodes Er. (Cucujidae). ,

apl1, aes2-3 f1-3 (2-3)
. , , ais1-3
, , -, . ,
, ,
.

: Carabus nemoralis
Muller, C. granulatus L. (Carabidae), Stenagostus undulatus Deg. (Elateridae), Dorcadion sp.
(Cerambycidae), Hylecoetus dermestoides L. (Lymexylidae), Schizotus pectinicornis L.
(Pyrochroidae), Calopus serraticornis L. (Oedeomeridae).
. ,

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12

Stenagostus Carabus ( )
. , ,
( ) ,
de novo ( , , Stenagostus)
(Carabus). f3 .
(.. ,

).
, , :
f1-2, f3
( ), . -,
,
.
,
, (
).
3.3.2. Holometabola
Coleoptera

Holometabola.
(Poyarkoff, 1914; Hinton, 1955; Sehnal et al, 1996). ,
() ,
, ,
. ,
Holometabola,
Coleoptera. ,
,
Holometabola.
,
, ,
(Lubbock, 1883; Berlese, 1913; , 1929; Henson, 1946, ,
1991).

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13



Hemi- Holometabola (. 2). ,
, Coleoptera
. ,
, ,
(Poyarkoff, 1914; Henson, 1946). ,
, .
(
), .
, ,
(, 1961) , ..
Hemimetabola.

4. Neuropteroidea, Mecoptera, Hymenoptera,


Lepidoptera Trichoptera


Neuropteroidea (Neuroptera, Rhaphidioptera Megaloptera) Coleoptera (
),
.
. ,
Sialis
Coleoptera.

: Hymenoptera
Lepidoptera, Mecoptera, ,
Trichoptera. Panorpa Allotrichia
; -

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14

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15

, aes2-3 sp3
. , Mecoptera
Trichoptera Neuropteroidea: ,
.
,
Coleoptera Neuropteroidea, Megaloptera,
(Crampton, 1914; Beutel, 2007).

5.


. , ,
.

, .

6. Coleoptera
6.1. Coleoptera
, Coleoptera
: 1)
, 2)
.
, ,
. , Schizotus, Dendroides, Pyrochroa
Pyrochroidae, ,
; Melandrya Orchesia
(Melandryidae) . , Coleoptera, ,
,
, . ,

Tenebrionoidea

Cucujoidea,

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16

.
,
, ,
. ,
,
( )
.
6.2. Coleoptera

, ,
. 8
, , ,
( 1).

. ,
; .
. ,
,
,
. ,
. ( 19 ) ,
, apl1 aes1-3. , ,
( )
, .
,
, - ,
.
, -
Silphidae, Eulichadidae, Endomychidae
- Scarabaeoidae. ,

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17


f2-3 apl2-3 sp2-3 ais2-3

apl1

aes1-3

Trachypachydae, Carabidae, Hydrophilidae,


Erotylidae

Silphidae, Eulichadidae, Endomychidae,


Scarabaeoidae, Endecatomidae

Lampyridae

Coccinellidae

Nosodendridae

Dytiscidae, Gyrinidae, Lucanidae

Byrrhidae, Dascillidae, Scirtidae,


Psephenidae, Helophoridae, Passalidae, etc

Lymexylonidae, Cerophytidae

Staphylinidae, Scaphidiidae

Heteroceridae, Elmidae

Cantharidae, Callirhipidae

Elateridae, Drilidae, Lycidae, Ptinidae,


Dermestidae, Haliplidae

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ais1

18

Zopheridae, Oedemeridae, Pythidae,


Monotomidae, Monommatidae, Othnidae
Salpingidae, Synchroidae, Nitidulidae,
Leiodidae, Corylophidae, Pyrochroinae

Cryptocephalinae

Cleridae, Peltidae, Trogossitidae, Silvanidae

Noteridae, Rhysodidae, Anobiidae,


Malachidae, Cucujidae, Byturidae, Ciidae, etc

1. Coleoptera.
. ; .
,

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19


. ,
,
.
6.3.
Coleoptera
,

, (
) . ,
, ,
. , , ,
- .

Coleoptera ( 2).
, ,
, ,
. ,
, ,
.
-,
. -,


, /
. -,
,
.

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20

Lawrence, Newton, 1995

\. Carabidae (Bell, 1998);


Carabidae (Beutel, 1990, 1992);
Rhysodes

Rhysodidae, Adephaga (
Carabidae)

Adephaga (Ponomarenko, 1995);

Carabidae

Archostemata (Makarov,
2008)
Nodinus

Apateticinae, Staphylinidae

Apateticinae, Silphidae (Madge, 1980)

Apateticinae, Staphylinidae, Scaphidiidae

Drapetes

Lissominae, Elateridae

Lissomidae (Burakowski, 1976 )

Lissominae, Elateridae

Orphilus

Orphilinae, Dermestidae

Nosodendridae (Ivie, 1985, 2002)

Orphilinae, Dermestidae; Thylodriinae

Lagria

Lagriinae, Tenebrionidae

Lagriidae (Boving, 1931; Crowson, 1981)

Lagriinae, Tenebrionidae

Elacatis

Othniinae, Salpingidae

Othniidae (Boving, 1931); Elacatidae (Crowson, 1981)

Othniinae, Pythidae ( )

Inopeplus, Diagrypnodes

Inopeplinae, Salpingidae

Inopeplidae (Sato, Hatta, 1988)

Inopeplidae

Aglenus

Agleninae, Salpingidae

Agleninae, Colydiidae (Dajoz, 1977)

Agleninae, Salpingidae

Pedilus

Pedilinae, Pyrochroidae

Anthicidae (Abdullah, 1969); Pedilidae (Lacordaire, 1859)

Pedilinae, Pyrochroidae

Morpholycus, Temnopalpus

Pilipalpinae, Pyrochroidae

Pythidae (Watt, 1987);


Pilipalpidae
Pilipalpidae (Nikitsky, 1986)
Colydiidae, Zopheridae,
Monommatidae

Colydiidae, Zopheridae, Monommatidae

\. Zopheridae (Lawrence, lipinski, 1997)

Zopheridae (Colydiinae + Zopherinae), Monommatidae

Meryx, Ulodes

Ulodidae

Ulodinae, Zopheridae (Doyen, Lawrence, 1979)

Ulodidae, Archeocrypticidae Mycetophagidae

2. Coleoptera.
(Lawrence et al, 1995) ; .
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21

7. Coleoptera
Coleoptera
.
,
,
.
,

(, 1974, 1976).
() ()
.

, Adephaga, Staphylinoidea Hydrophyloidea,


,
.
,
. , , -
Buprestidae, Cerambycidae, Curculionidae,
. , ,
, .

, .

Coleoptera. ,

, .
Megaloptera
(Crampton 1918; Wheeler et al, 2001, , 2006), ,
Coleoptera .
Archostemata
, ,
, (Beutel, Hrnschemeyer, 2008).

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22


. ,
,
, ,
. Adephaga ( Trachypachidae
Gyrinidae) Hydrophyloidea Polyphaga (, 1969, 1995;
Grimaldi, Engel, 2005). ,
Trachypachidae (Geadephaga), Dytiscidae, Gyrinidae (Hydradephaga) Hydrophilidae
Polyphaga .
7.1. Coleoptera

.
, ,
,
.
Hydrophiloidea Elateroidea.
,
. ( )
Coleoptera ,

-, - , .
Polyphaga ( Cleroidea, Drilidae,
Elateridae Tenebrionidae)
, .
, Adephaga. ,

, .
- ,
. , ,
Buprestidae, Cerambycidae Curculionidae.

Coleoptera

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23

.
, ,
. ,
,
.
,
(, 1996). ,

.

.
,
,
, . , , Cicindelini
Carabidae, Clytrini Chrysomelidae, Callirhipidae, Eulichadidae .
,
, .
,
Coleoptera ,
. ,

(, 1929, 1939).
,
.
. ,
. ,

. ,
,
, , , ,
.
, (, 1939; , 1991),
Holometabola .

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24

1. Coleoptera
,
.
2.
,
.
.
,
.
3. Coleoptera
, ,

.
, Coleoptera , -, Holometabola,
,
, Hemimetabola
4. 8
,
,
5.

/
.
.
,


6.

Coleoptera,

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Nodynus

25

(Staphylinidae, Apateticinae); Lagria (Tenebrionidae, Lagriinae); Inopeplus, Diagrypnodes


(Inopeplidae);

Elacatis

(Pythidae,

Othniinae);

Pedilus

(Pyrochroidae,

Pedilinae);

Morpholycus, Temnopalpus (Pilipalpidae) .


7. , -,
Carabidae, Trachypachidae, Dytiscidae, Gyrinidae (Adephaga), Hydrophilidae (Polyphaga).
Sialis,
Coleoptera Megaloptera.
8.
, .

.

, , .

.. - (Coleoptera,
Pyrochroidae) / .. // 2005. - 14 (1). .
15-20.
Zaitsev A.A. The endoskeleton of thoracic segments of Carabid larvae (Coleoptera,
Carabidae) / .. // Back to the Roots and Back to the Future? Towards a New
Synthesis amongst Taxonomic, Ecological and Biogeographical Approaches in Carabidology.
- 2007. - Abstract Volume - p. 68.
.. Coleoptera / .. //
XIII , . 2007. .
117-118.
Zaitsev A.A. Description of larval and imaginal stages of new species from the genera
Pseudacroleptus Pic, 1911 Ceratoprion Gorham, 1880 (Coleoptera: Lycidae: Leptolycinae) /
S.V. Kazantsev, A.A. Zaitsev // Russian Entomol. J. 2008. - 17 (3). . 283-292.
Zaitsev A.A. Thoracic endoskeleton of carabid larvae (Coleoptera, Carabidae) / A.A. Zaitsev
// Back to the Roots and Back to the Future? Towards a New Synthesis amongst Taxonomic,
Ecological and Biogeographical Approaches in Carabidology Proceedings of the XIII

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26

European Carabidologists Meeting, Blagoevgrad. 2008. . 159-171.


.. Tenebrionoidea (Insecta, Coleoptera)
/ .. //
2009. 8 (2). - . 189-204.

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