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HOMEOSTASIS
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9.1 Concept of Homeostasis
1. Explain the concept of homeostasis.

2. Describe the homeostatic control system.


Concept of Homeostasis
(homeo = sameness; -stasis = standing still)

The condition in which the bodys internal


environment remains relatively constant,
within physiological limits.
Concept of Homeostasis

This equilibrium produced by the dynamic


state, an interplay between outside factors
and internal control mechanisms.

Outside factors tend to change the


internal environment

Internal control mechanisms that


oppose such changes
Concept of Homeostasis

OUTSIDE
INTERNAL CONTROL
FACTORS MECHANISM

E g : Thermoregulation

E g : Temperature

Activates
Changes

Reverses the change

Internal
environment
Concept of Homeostasis

The characterization of an animal with


regards to environmental variables is
called regulation.

A regulator uses mechanisms of


homeostasis to control internal changes
caused by external fluctuations.
Concept of Homeostasis

It involves receptors and adjusting the


composition of the body fluids accordingly,
carrying out excretion and osmoregulation.

Regulation by an organism is very


important so that physiological processes
can proceed at an optimum rate.
Homeostatic Control System

The body can regulate its internal
environment through a multitude of
feedback systems.

A feedback system is a cycle of events in


which the status of body condition is:

continually monitored,

evaluated,

changed,

re-monitored,

re-evaluated and so on.



Each monitored variable, such as

body temperature,

blood pressure,

blood glucose level


is termed a controlled condition.

Any disruption that changes a controlled


condition is called a stimulus.
Three basic components make
up a feedback system

RECEPTOR

CONTROL
SYSTEM
EFFECTOR
Three basic components
make up a feedback system
RECEPTOR

A body structure that


monitors changes in a
controlled condition and
sends input in the form of CONTROL
nerve impulses or chemical
signals to a control center. SYSTEM

EFFECTOR
Three basic components make
up a feedback system
RECEPTOR
CONTROL SYSTEM
Evaluates the input it receives
from receptors and generates
output commands when they
are needed.

Output from the control center


can occur in several forms:
nerve impulses, hormones , or
other chemical signals.
EFFECTOR
Three basic components make
up a feedback system
RECEPTOR

CONTROL
EFFECTOR
SYSTEM A body structure that receives
output from the control centre
and produces a response or effect
that changes the controlled
condition.

Either depressing it (negative


feedback) or enhancing it (positive
feedback)
Feedback system

Negative feedback
Positive feedback
A feedback mechanism in
Homeostatic mechanism which the response
that reduces the intensity of enhances the original
the original stimulus stimulus

Consequently causes a The output is used to


enhance the input.
change in a variable that is
opposite in direction to the
initial change

The output is used to reduce


input.
Negative Feedback System

A primary mechanism of homeostasis.

If there is a change in a physiological variable


that is being monitored,

it triggers a response that counteracts the


initial fluctuation.
(Campbell, 6th edition)
Negative Feedback

Examples of negative feedback:

Negative feedback in regulation of blood


glucose level

Negative feedback in mechanism of body


temperature regulation
Positive Feedback System

A physiological control mechanism.

A change in some variable..

triggers mechanisms that amplify that change.

(Campbell, 6th edition)


An example of positive feedback (during childbirth)
Steady-state
condition

Decrease Increase
Positive feedback Positive feedback

Further decrease Further increase

Corrective mechanism
Negative feedback

Steady-
state
condition
A generalized diagram of homeostasis
HOMEOSTASIS

Negative Feedback
Mechanism in
Controlling
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Blood Glucose Level


Learning Outcomes

Explain the negative feedback in controlling


blood glucose.
2 Fundamental Facts

A rise in blood glucose level stimulates the


secretion of insulin.

A fall in blood glucose level would instead


stimulate glucagon secretion.

-cells secrete insulin

-cells secrete glucagon


When glucose level in
the blood increases
When glucose level in
the blood decreases
Please take note

-cells, not beta cells.

-cells, not alpha cells.


insulin
Glucose Glycogen

glucagon
Glycogen Glucose
INsulin is released when there is an
INcrease in the blood glucose level..

AND

glucaGON is released when


all the glucose has GONE!
But heres some news you can use..

Source : Business Week,


14th January 2010.
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HOMEOSTASIS

Negative Feedback
in
Body Temperature
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Regulation
Learning Outcomes

Explain negative feedback in the regulation of


body temperature
How does the body regulate
temperature?
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The bodys temperature is monitored by
nerve cells in the skin,
hypothalamus and other parts of the body

Some nerve cells functions as


hot thermoreceptors that signal
the hypothalamus when the skin
or blood temperature increases.

Others functions as cold


thermoreceptors that signal the
hypothalamus when there is a
decrease in temperature of the
skin or blood.
Diagram :
Cross-section of skin
More heat is lost from the surface of the skin

If the temperature
rises, the blood
vessel dilates
(vasolidation).
Sweating

The
skin
vasoconstriction
Less heat is lost from the surface of the skin

If the temperature
falls, the blood
vessel constricts
(vasoconstriction).
If all else fails,

Thank you
LECTURE 2

9.3.1 Liver
Learning outcomes:

Students should be able to:

v
Describe the detailed structure of liver
v
Explain its homeostatic function
- Carbohydrate metabolism
- Detoxification
- Transamination & deamination of
amino acids
v
Illustrate Urea / Ornithine cycle
Liver

Largest internal organ


of the body (1.4 kg)

Located in the right-


upper quadrant of the
abdomen and under
the diaphragm
Liver

The gall bladder,


which stores bile
secreted by the
liver, lies just below
it and connected to
the duodenum by
the bile duct.
Structure of liver : Lobes
Consist of

Two major lobes Two minor lobes

Right lobe Left lobe Caudate lobe Quadrate lobe


Structure of liver : Lobes
Two major lobes, Two minor lobes,
right & left caudate & quadrate
Structure of liver : Lobes
Structure of liver: Lobes

The right lobe, is visible


on all liver surface and
separated from the
smaller left lobe by a
deep fissure

The posterior most lobe


is the caudate lobe and
quadrate lobe, which lies
inferior to the left lobe.
Structure of liver : Lobules

The liver consist


of thousands of
tiny polygonal
blocks of cells
called lobules

Each lobules
about 1 mm in
diameter
Structure of liver : Lobules
Structure of liver : Lobules

Lobules are formed from


radiating rows of vertical
cord of liver cells,
referred to as acini
(singular, acinus).

Between the lobules are


branches of the hepatic
artery, hepatic portal
vein and bile duct.
Simplified diagram of a liver acinus

Acinus
Structure of liver : Hepatocytes

A lobule consists of
specialized liver cells
called hepatocytes.

Hepatocytes are
arranged in irregular,
branching,
interconnected plates
around a central vein.
Structure of liver : Hepatocytes

Hepatocytes have
prominent nuclei
and Golgi apparatus,
many mitochondria
and lysosomes.

Also rich in glycogen


granules and fat
droplets.
Structure of liver : blood vessels
Connected by
Two blood vessels sinusoids One blood vessel
(tiny blood vessel)
supply blood to drains blood away
the liver cell from the liver cells

Hepatic artery Hepatic Hepatic vein


portal vein
Structure of liver : blood vessels
Hepatic Hepatic vein
Hepatic artery portal vein

Carries Sends back


oxygenated Supplies blood blood from the
blood from the gut & bring liver to the
along most of the heart
digested food into it
Structure of liver : Bile

The sinusoids alternate with bile canaliculi.

Bile canaliculi contains bile, made by the hepatocytes


which line them, and transports them into the
branches of the bile duct.
Structure of liver : Bile

Bile

A bitter- tasting greenish-


yellow alkaline fluid

Produced by liver cells

Stored in the gall bladder


and secreted into
duodenum

It assist the digestion and


absorption of fats.
Structure of liver : Kupffer cell

The sinusoids are


lined by numerous
phagocytic cells
known as Kupffer
cells (also known as
star cells/stellate
cells)

Kupffer cells are part


of the bodys defence
system
Functions of liver

Carbohydrate metabolism

Detoxification

Transamination & Deamination of


amino acids
Carbohydrate metabolism
Carbohydrate metabolism
Role of the liver in (the control) blood glucose level:
carbohydrate metabolism
Sugar enters the liver from the gut by the hepatic portal
vein.

When the blood glucose level is high

v
-cells from islet of Langerhans of pancreas secrete
insulin and stimulate the liver


The conversion of glucose to glycogen is known as
glycogenesis and is stimulated by the presence of
insulin
When blood glucose level is low

-cells of islet of Langerhans in pancreas secretes


glucagon

glucagon stimulates liver cells to convert glycogen


into glucose (glycogenolysis)

glucose can be synthesized from non-carbohydrate


precursors such amino acids, and glycerol
(gluconeogenesis)
Muscle: Uptake glucose by cell
Insulin secreted Liver: Negative feedback
glucose CO2 + H2O
glucose glycogen
Islet of langerhans (detector)
glucose fat

Fall in glucose
Rise in glucose level
level

Normal blood Normal blood


Glucose level Glucose level

Fall in glucose
level
Rise in glucose
level
Islet of langerhans (detector)

Liver:
glycogen Negative feedback
Glucagon secreted
glucose
protein glucose
Figure : summary diagram of control blood glucose level
Transamination of amino acid

Process of transferring the amino group from one


amino acid to form other amino acid

An amino group from amino acid is transffered to


organic acid or to an acid (keto acid) in the Krebs
cycle.

It means other non-essential amino acids can be


formed by the body.
Deamination of amino acid

The liver is the site of the


deamination of excess
amino acids

amino acids are broken


down and the amino group
(NH2) is removed

and used to make ammonia


NH3
Detoxification


From deamination, amino acid is broken
into a non-nitrogenous part and a
nitrogenous part.
H R R
deamination
2 N C COOH + O2 2 C COOH + 2NH3

H
H O
ammonia
Keto acid
Detoxification

The non-nitrogenous parts are


converted into glucose in the
liver and then stored as
glycogen

The nitrogenous part is


ammonia which is toxic even in
small quantities

Ammonia is converted into


urea which is less toxic
Formation of urea

Urea, CO(NH2)2 is
the nitrogenous
waste product of
humans and other
land-living mammals

Ammonia from
deamination is
converted to urea in
the ornithine cycle
(urea cycle).
Formation of urea

Two molecules ammonia are combined with one carbon


dioxide

And enter the ornithine cycle

2NH3 + CO2 NH2 + H2O


C=O

Ammonia
NH2
carbon dioxide urea water
Formation of urea

Urea cannot be made


directly by combining
carbon dioxide and
ammonia because this
is not a feasible
reaction.
Formation of urea

By using the four


molecules:

ornithine, citrulline,
arginosuccinate and
arginine as intermediates,

and the energy from ATP


molecules,

the synthesis of a molecule


of urea becomes feasible
arginosuccinate
A porter

BACK
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LECTURE 3

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9.3.2 Kidney
Structure of kidney and nephron
Learning outcomes

Describe the gross structure of kidney and


the detailed structure of nephron and the
associated blood vessels.

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Gross structure of kidney
Kidney

Is the major excretory and


osmoregulatory organ of
mammals

Function:

Removal of metabolic
waste products

Regulation of the water
content of body fluids

Regulation of pH of body
fluids

Regulation of the
chemical composition of
body fluids
Structure of kidney

Pair of bean-shaped
organ

Situated towards the back


of the lower part of the
abdominal cavity

The left kidney lies


slightly above the right

It is 10-20 cm long, 5-7.5


Structure of kidney

Receive blood from


the aorta via the renal
arteries

Renal vein returns


blood to the posterior
vena kava
Structure of kidney

Urine formed passes


by a pair of ureters
to the bladder where
it is stored until it is
released via urethra
Structure of kidney : transverse section
Two regions
Cortex (outer) Medulla (inner)

covered by fibrous
composed of tubular
connective tissue parts of the nephron and
blood vessels which

Contains glomerulus, together form renal


Bowmans capsule and pyramids
part of the nephron
A transverse section
Structure of nephron
Structure of nephron
Bowmans capsule

Each nephron begins as a


hollow, cup shaped component
(diameter 200m)
in the cortex of the kidney.

From here , the tubule runs


towards the center of the
kidney.

The cupped wall region and


blood vessel is a blood
filtering unit called a
glomerulus.
Structure of nephron
Proximal
convoluted tubule

Close to the capsule,


highly coiled and
diameter 60m.

Located in the cortex


of the kidney.
Structure of nephron
Loop of Henle

Hair-pin shaped.

Has descending
limb & ascending
limb.

Located in the
medulla of the
kidney.
Structure of nephron
Distal convoluted
tubule

Not as long as the


proximal convoluted
tubule.

Located in the
cortex of the kidney.
Structure of nephron
Collecting duct

End of kidney.

Located in the medulla of


the kidney.

Several nephrons feed


into the same collecting
duct.

The collecting ducts all


eventually drain into the
pelvis of the kidney, from
where the urine flows into
the ureter.
Nephron and the associated blood
vessels
Renal artery
Blood enters the kidney by the renal artery
which branches into the afferent arteriole.
Nephron and the associated blood
vessels

An afferent arteriole brings blood towards each


Bowmans capsule, dividing to form a network of
capillaries called glomerulus in the hollow part of
the capsule.
Nephron and the associated blood
vessels
The blood leaves the Bowmans capsule in an
efferent arteriole, which is narrower than the
afferent arteriole.
Nephron and the associated blood
vessels
The efferent arteriole is
divided to form a second
set of capillaries which
surround the:

proximal and distal


convoluted tubules

and the loop of Henle in


the medulla.
Nephron and the associated blood
vessels
The capillaries of the
peritubular capillaries
(vasa recta) run parallel
to the loop of Henle
and the collecting duct
in the medulla.

Blood leaves the


kidney through the
renal vein.
Summary of blood
circulation in the
kidney:

Renal artery afferent


arteriole glomerulus
efferent arteriole
peritubular capillary (vasa
recta) renal vein
9.3.2 Kidney
Urine formation

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Learning Outcomes

Explain the formation and concentration of


urine involving ultrafiltration, reabsorption
and secretion.
URINE FORMATION
ULTRAFILTRATION REABSORPTION SECRETION
Urine formation :
1) Ultrafiltration

Ultrafiltration occurs
from the capillaries of
the glomerulus into
the lumen of the renal
capsule.
Urine formation :Ultrafiltration

Filters out molecules


which are much smaller
than RBC and plasma
proteins.

Substance forced through


the capillary wall and
between the podocytes
by blood pressure enter
the lumen of Bowmans
capsule.
Filtrate

H2O

Salts (NaCl and


others)

HCO3-

H+

Urea

Glucose; amino
acids

Some drugs
Factors contributing to ultrafiltration
a) The hydrostatic blood
pressure in the glomerulus
is higher than in other
capillaries.

q
This high pressure is
mainly due to the high
resistance to outflow
presented by the efferent
arteriole, which is
smaller in diameter than
the afferent arteriole.
Factors contributing to ultrafiltration

b) Large amount of
glomerulus filtrate large
surface area for filtration
provided by the highly coiled
glomerulus capillaries.
Factors contributing to ultrafiltration
c) High permeability of the
glomerulus

q
The wall of Bowmans
capsule in contact with
capillaries consists of
specialized epithelial cell
called podocytes.

q
These cells have numerous
cytoplasmic extension
called foot processes that
cover most of the
capillaries.
Factors contributing to ultrafiltration

q
Foot processes of
adjacent podocytes
are separated by
narrow gaps called
filtration slits.
q
The perforated walls of the
capillaries and the
podocytes form a filtration
membrane that permits fluid
and small solutes dissolved in

the plasma, to pass through


and become part of the
filtrate.

q
This filtrate membrane holds
back blood cells, platelets,
and most of the plasma
protein.
2) Reabsorption (proximal convoluted
tubule, loop of Henle, distal convoluted
tubule and collecting duct)

Nearly all sugars,


vitamins and other
organic nutrients
present in the initial
filtrate are reabsorbed.
i) Proximal tubule

Helps maintain a constant pH in


body fluid
By controlled secretion of
hydrogen ions and
By reabsorbing about 90% of
HCO3- from filtrate

)
Reabsorbing of NaCl (salt) and
water

Salt moves from the filtrate to
the interstitial fluid by active
transport

Water follows passively by
osmosis
Reabsorption and secretion at
proximal convoluted tubule

Reabsorption from
proximal tubule
into blood

Secretion from
blood into proximal
tubule
Reabsorption in the proximal convoluted
tubule

The wall is one cell thick

The cells of the tubule are


packed with mitochondria

And the surface facing into


the tubule has a brush
border of microvilli

Microvilli provide a large


surface area for the
movement of solute
molecules into the cell
i) Proximal tubule
(cont.)

)
Glucose , amino acids ,
vitamins and other
useful materials are
also reabsorbed into
the blood.

)
This is accomplished
by a combination of
active transport,
diffusion and osmosis.
ii. Decending limb of the loop
of Henle

Permeable to water but not to


salt

Water moves out of the tubule


by osmosis

Filtrate moves downwards and


continues to lose water

NaCl concentration inside the


loop of Henle increases
iii. Ascending limb of
the loop of henle

Consist of thin segment


and thick segment

The walls become more


permeable to NaCl

and less permeable to


water

Salt that was concentrated


now diffuse out of the thin
segment of ascending limb.
iii.Ascending limb of the
loop of henle

This contributes to the high


salt concentration in the
interstitial fluid in medulla

In the thick segment ,


salt is actively transported
out of the tubule

Filtrate becomes progressively


more dilute as it moves up to
the distal tubule
iv. Distal convoluted tubule

Here, NaCl and HCO3- are


also reabsorbed into the blood
by active transport.

Water molecules are


reabsorbed into the blood by
osmosis.
Reabsorption and secretion at
distal convoluted tubule

Reabsorption from
distal tubule into
blood

Secretion from
blood into distal
convoluted tubule
v. Collecting duct

Carries the filtrate in the


direction of the medulla
and renal pelvis

The special tissue


epithelium is permeable to
water but not to salt
v. Collecting duct (cont.)

The filtrate becomes more


and more concentrated as
water is lost to the interstitial
fluid

The bottom portion is


permeable to urea and
leakage of this solute into
the interstitial fluid
contributes to the high
osmolarity of the medulla
3) Secretion
The opposite direction of
reabsorption

K+ and H+ are among the


substances actively secreted
from blood into the filtrate

NH3 and certain drugs are


also removed from blood by
secretion

Secretion occurs mainly in


the regions of the distal
convoluted tubule.
Urine concentration by counter
current multiplier mechanism

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Definition:

1) Countercurrent
Fluids past each other in opposite directions.
Fluid flow down one side of the loop the
descending limb, and up the other side the
ascending limb

2) Multiplier
Fluid flow down the descending limb, high
concentration of NaCl ; hypertonic
Fluid flow up the ascending limb, less
concentration of NaCl ; hypotonic
counter current multiplier mechanism
The loop of Henle as a
countercurrent multiplier
The loop of Henle maintains the interstitial fluid
gradient of NaCl in medulla.
The filtrate concentration of this salt increases
because of the loss of water from the
descending limb.
Then the ascending limb leaks the salt into the
interstitial fluid.
Additional salt is actively transported out of the
thick segment of the ascending limb.
The loop of Henle as a countercurrent
multiplier
The high concentration of
salt in the medulla makes
water leave the
descending limb by
osmosis

The counter current like


arrangement of
descending and ascending
limbs helps maintain the
high salt concentration in
the medulla
Under conditions in which the kidney
conserves as much water as possible,
urine can reach an osmolarity of about
1,200 mosm/L, considerably hyperosmotic
to blood (about 300 mosm/L)

This ability to excrete nitrogenous wastes


with a minimal loss of water is a key
terrestrial adaptation of mammals
Urine concentration by counter
current multiplier mechanism

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The loop of Henle acts as a
countercurrent multiplier.

Countercurrent refers to the fact that:

Fluids past each other in opposite directions.

Fluid flow down the descending limb, and up


the ascending limb.

There could be no osmotic gradient if
the filtrate in these adjacent tubes ran in
the same way.

Therefore there could be no exchange of


The term multiplier refers to the fact that:

Small differences of osmolarity at any one


point along the loop is multiplied along the
length of the loop.

This creates a large osmotic gradient.

Fluid flows down the descending limb, high


concentration of NaCl ; hypertonic.

Fluid flows up the ascending limb, less


concentration of NaCl ; hypotonic.
counter current multiplier mechanism
The loop of Henle as a
countercurrent multiplier
The loop of Henle maintains the
interstitial fluid gradient of NaCl in
medulla.
The filtrate concentration of this salt
increases because of the loss of water
from the descending limb.
Then the ascending limb leaks the salt
into the interstitial fluid.
Additional salt is actively transported out
of the thick segment of the ascending
limb.
The loop of Henle as a countercurrent
multiplier
The high concentration of
salt in the medulla makes
water leave the
descending limb by
osmosis

The counter current like


arrangement of
descending and ascending
limbs helps maintain the
high salt concentration in
the medulla
Under conditions in which the kidney
conserves as much water as possible,
urine can reach an osmolarity of about
1,200 mosm/L, considerably hyperosmotic
to blood (about 300 mosm/L)

This ability to excrete nitrogenous wastes


with a minimal loss of water is a key
terrestrial adaptation of mammals
9.3.2 kidney (continued)

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Learning Outcomes

Students should be able to explain:

-
water regulation by ADH

-
The action of aldosterone on Na+
reabsorption

-
pH regulation of the tissue fluid
Water regulation by ADH
Concepts


The kidneys play central role in the
regulation of the water content of the
body or osmoregulation.


The water content of the blood is
monitored by osmoreceptor cells in the
hypothalamus and they stimulate the
pituitary gland to secrete antidiuretic
hormone, ADH.
What happens if you havent
had a drink for a while?

1. Your blood will have a low water


potential (it becomes more
concentrated).

2. This is detected by osmoreceptors in


your hypotalamus, which stimulates
your pituitary gland to release
antidiuretic hormone (ADH).
The release of ADH into the
bloodstream brings about the
following:

ADH makes the distal


convoluted tubule and
collecting duct more
permeable to water.

This allows more water to be


reabsorbed into blood.

This produces a smaller


volume of more concentrated
urine.
In addition to stimulating
the release of ADH
when the blood is
concentrated, the
hypotalamus also
activates your thirst
centre in the brain.

As a result, you feel


thirsty and take in
additional water to dilute
the blood.
What happen if you drink lots of
fluids?
Your blood will have a
high water potential (it
becomes more dilute).

This is detected by the


osmoreceptors in your
hypotalamus and
results in a decrease in
the amount of ADH
secreted by the
pituitary.
The decrease in the amount
of ADH in the bloodstream
results in the following:

Lack of ADH makes the


distal convoluted tubule and
collecting duct less
permeable to water.
Less water is reabsorbed
into the blood.
Larger quantities of dilute
urine are produced.
The action of aldosterone on Na+
reabsorption
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Sodium reabsorption is regulated
by the hormone Aldosterone
Sodium is the most
abundant extracellular
ion.

Its concentration is
carefully regulated by
the hormone,
aldosterone.
Sodium reabsorption is regulated
by the hormone Aldosterone
Aldosterone secretion can be
stimulated by a decrease in
blood pressure.

When blood pressure fall,


certain cells (juxtaglomerular
apparatus) near the
glomerulus secrete the
enzyme renin and activates
the renin-angiotensin
pathway.
Sodium reabsorption is regulated
by the hormone Aldosterone
Renin acts on a plasma
protein, converting it to
angiotensin, which
stimulates aldosterone
secretion.

Aldosterone is secreted by
the cortex of the adrenal
glands.
Sodium reabsorption is regulated
by the hormone Aldosterone
This hormone
stimulates the distal
tubules and collecting
duct to increase
sodium reabsorption.

Then the blood


volume and pressure
will increase.
pH regulation of the tissue
fluid
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pH regulation of the tissue fluid

pH is a measure of the concentration of


hydrogen ions and the acidity of the blood.

The nephron regulates the pH of the


blood, maintaining it at pH 7.4.

pH < 7.3 : acidosis

pH > 7.5 : alkalosis

One factor which tends to increase acidity


is production of CO2 during cell
Carbon dioxide reacts with water to produce
carbonic acid:

CO2 + H2O H2CO3

Carbonic acid dissociates to produce hydrogen


ions and hydrogen carbonate ions:

H2CO3 H+ + HCO3-
Process

In the distal
convoluted tubule
longer-term
adjustments in the
ion balance of the
blood are made.

If the blood pH is
tending to fall below
7.4, H+ ions from the
plasma are secreted
If the blood pH rises
above 7.4, OH- ions
from plasma are
secreted in the distal
convoluted tubule.
9.4 Water regulation in plants
Objectives :
a) Describe the role of stomata in regulation
of water loss
b) Explain the importance of transpiration
c) Describe the various types of plant
adaptation to the environment in relation
to morphology, anatomy and physiology
The role of stomata in
regulation of water loss
On a short term basis,
closing of the stomata can
control water loss

This occurs in many


plants when they are
subjected to water stress

About 90% of the water a


plant loses escapes
The role of stomata in regulation of
water loss
These pores account for only 1-2% of the
external leaf surface

The waxy cuticle limits water loss through


the remaining surface of the leaf

Guard cells control the diameter of the


stoma by changing shape

The amount of water lost by a leaf



The importance of transpiration

Transpiration is the loss of


water vapour from the leaves
and stems of plant

About 90% of the water is


lost through transpiration.

Transpiration is an important
process to plants, because it
provides the force to pull
water and mineral ions up
the xylem vessels
The importance of transpiration

Transpiration has a cooling


effect, countering the heating
effect of the sun on the whole
shoot system

Transpiration can lower the


temperature of a leaf in
comparison with the
surrounding air

This prevents the leaf from


reaching temperatures that
Plant adaptation of habitats


Xerophytes

Hydrophytes

Mesophytes

Halophytes
Xerophytes
Live in short water supply
Problem: loss of water death
Features to reduce water loss

Waxy cuticle

Smaller leaves

Presence of trichomes

Leaf rolled up

Succulent

Stomata
Close in the day
Open in the night

Photosynthesis - CAM
Hydrophytes
Live in water
Adaptation:

Xylem vessel is poorly
developed

Large leaves

Lack sclerenchyma tissue

Stem and leaves have
paerenchyma tissue

Large air spaces
Mesophytes
Live with an adequate
water supply
Most angiosperm plants
No special adaptation
Hot & dry:

close stomata

Thin layer cuticle

Stomata under leaves

Variable leaf shape


cold season:

Seeds dormant
Halophytes
Live in areas of high salinity
Problem:

absorption of water from salty
water

High rate of transpiration
Adaptive:

Root accumulate minerals by
active absorption

Thick layer of cuticle

Succulent leaves

Reduced stomata and hairs

Root submerged in salt
water/knee root
The End
Thank you and till we meet again

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