Invertebrates, Second Edition

Richard C. Brusca and Gary J. Brusca

ABOUT THE BOOK

Invertebrates, Second Edition presents a modern survey of the 34 animal phyla (plus the Protista) and
serves as both a college course text and a reference on invertebrate biology. Thorough and up-to-date, it is
organized around the themes of bauplans (body plans) and evolution (phylogenetics). Each phylum is
organized in a standardized fashion, treating the systematics, bauplan (support and movement, feeding and
digestion, circulation and gas exchange, excretion and osmoregulation, nervous system, reproduction and
development), and phylogeny. Detailed classifications, phylogenetic trees, and references for all phyla are
provided. Tables summarize each phylums defining attributes. The text is accompanied by an abundance of
detailed line drawings andnew to this editioncolor photographs.

Other key changes from the First Edition (1990) include:

the incorporation of new developments in phylogenetics, developmental biology, and molecular
genetics
major changes at the highest levels among the invertebrates. Three phyla that appeared in the
original bookPentastomida, Pogonophora, and Vestimentiferano longer exist, and a new
phylum, Cycliophora, has been erected. Moreover, this edition discusses recent work in molecular
systematics that has shaken classic views on animal classification.
a large new section on Kingdom Protista (replacing Protozoa) containing new contemporary
views of these organisms (arranged in 18 phyla).

ABOUT THE AUTHORS

Richard C. Brusca is Director of Conservation and Research at the ArizonaSonora Desert Museum, and
also holds adjunct research positions at the University of Arizona and CIAD (Centro de Investigacin en
Alimentacin y Desarrollo), Mexico. He earned a B.S. at California State Polytechnic University, an M.Sc.
at California State University (Los Angeles), and a Ph.D. at the University of Arizona. The author of over
100 research publications, including six books, Dr. Brusca is a widely recognized invertebrate zoologist,
marine biologist, and Sea of Cortez and Sonoran Desert naturalist. He is a Fellow in both the American
Association for the Advancement of Science and the Linnean Society (London). Dr. Brusca has been the
recipient of research grants from numerous organizations, including the National Science Foundation, the
National Oceanic and Atmospheric Administration, and the National Geographic Society.

The late Gary J. Brusca was Professor Emeritus at Humboldt State University, retiring in 1998. Dr. Brusca
earned his B.S. at California State Polytechnic University, his M.A. at the University of the Pacific, and his
Ph.D. at the University of Southern California. For 35 years he focused his research on the ecology and
systematics of hyperiid amphipods, also writing on a variety of other subjects in journals and textbooks.
His main academic love, however, was teaching, for which he was recognized with numerous awards.

CONTENTS

1. Introduction
2. Classification, Systematics, Phylogeny
3. Animal Architecture and the Bauplan Concept
4. Animal Development, Life Histories, and Origins
5. The Protista
6. Phylum Porifera: The Sponges
7. Four Phyla of Uncertain Affinity
8. Phylum Cnidaria
9. Phylum Ctenophora: The Comb Jellies
10. Phylum Platyhelminthes
11. Phylum Nemertea: The Ribbon Worms
12. Blastocoelomates and Other Phyla
13. Phylum Annelida: The Segmented Worms
14. Sipuncula and Echiura
15. The Emergence of the Arthropods: Onychophorans, Tardigrades, and the Arthropod Bauplan
16. Phylum Arthropoda: The Crustacea
17. Phylum Arthropoda: The Hexapoda (Insects and Their Kin)
18. Phylum Arthropoda: The Myriapoda (Centipedes and Their Kin)
19. Phylum Arthropoda: The Chelicerates (Spiders and Their Kin)
20. Phylum Mollusca
21. The Lophophorate Phyla
22. Phylum Echinodermata
23. Other Deuterostomes: Chaetognatha, Hemichordata, Chordata
24. Perspectives on Invertebrate Phylogeny

December 2002
ISBN 0-87893-097-3, $109.95 casebound