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GRAIN
Feed Industry Guide
FIRST
EDITION
1
INTRODUCTION
This technical guide on the use FEED BARLEY
of barley grain as poultry and
livestock feed is the first to be
BACKGROUND & MARKET
S.J. Meale1, M.L. He2 and T.A. McAllister1; 1Agriculture and Agri-Food Canada (smea4184@uni.
published by Alberta Barley. sydney.edu.au; tim.mcallister@agr.gc.ca); 2University of Saskatchewan (moro_he@yahoo.com)
3
FEED BARLEY protein and lower crude fibre than hulled barley, as the
hull accounts for a large proportion of the crude fibre
PRODUCTION & content of the kernel. It also contains higher levels of the
polysaccharide -glucan. This compound is considered
QUALITY GRADING undesirable for malting barley since it interferes with the
starch modification process. However, -glucan is highly
J. ODonovan Agriculture and Agri-Food Canada
desirable when barley is grown for food due to linkages
(john.odonovan@agr.gc.ca)
to positive health outcomes in humans. For this reason,
hulless varieties are being developed primarily for use as
TYPES OF BARLEY
human food. Studies have shown that hulless feed barley
Barley can be two-row or six-row and hulled or hulless.
has higher digestibility, protein and energy content, and
The barley type used can result in differences in feed
lower fibre than hulled barley. The high -glucan content
intake, feed efficiency, as well as malting quality. Two-row
may prove problematic, however, when feeding to swine
barley varieties produce fewer, but larger kernels per plant
and poultry, and enzymes are frequently used to improve
than six-row varieties, so two-row barley generally results
digestibility. Other constraints to growing hulless barley
in better feed efficiency in livestock and malting quality.
in Canada include lower yields, lack of a premium, and
These differences can, to some extent, be related to kernel
difficulty in segregating the product from hulled varieties
plumpness. Kernels of six-row varieties tend to be less
within the grain transportation system.
plump, especially when produced under relatively dry
conditions. This can result in reduced starch content with
TIMING OF BARLEY SEEDING
increased protein and fibre content. This is because starch
With the relatively short growing season in Canada, virtually
accounts for a lower portion of the total kernel weight.
all of the barley is seeded in spring. In general, barley tends
Fibrous carbohydrates are less digestible than starch,
to mature earlier than other major crops, including canola
and therefore yield less available energy to livestock
and wheat, so there is a tendency among growers to seed
and poultry. Two-row varieties have tended to produce
these crops before barley. Early seeding (early April to
lower overall grain yields than their six-row counterparts;
mid-May) favours higher yields compared to later seeding
however, this is changing with the development of high-
(late May to early June). A 10% increase in barley yield and
yielding two-row varieties for both the malting and feed
a 25% increase in net economic return with early seeding
markets.
has been recorded in eight regions across Western Canada
with the exception of the Peace River region of Northern
Approximately 99% of barley grown in Canada is of hulled
Alberta (Table 3). In the Peace region, later seeding resulted
varieties in which the hull or glume is retained during
in higher yields (7%) and net economic returns (30%; Smith
the threshing process. With hulless varieties, the hull is
et al. 2012). The effect of seeding time on barley kernel
removed during threshing. Hulless barley may also be
feed quality was less pronounced. Slight increases in kernel
referred to as naked barley. Hulless barley has a major
protein (11.5 to 11.9% DM), starch (61.2 to 61.3% DM), lysine
advantage over conventional barley in transportation,
(6.06 to 6.20% DM) and soluble fibre (4.07 to 4.20% DM),
processing, and storage. Removing the hull fraction
and a decrease in kernel plumpness (90 to 88%) occurred
increases the bulk density (weight-per-unit volume)
with late seeding. It is doubtful, however, if these small
compared to hulled barley by about 25%, thus cost savings
differences would markedly affect the feed value of barley
can be considerable. Hulless barley has higher crude
for livestock and poultry.
1A
5
In all regions, net economic returns were highest at 200 Fig. 2. EFFECT OF NITROGEN RATE ON BARLEY
to 300 seeds per m2 compared to lower or higher seeding QUALITY PARAMETERS
rates. On a precautionary note, the study also indicated
that seeding hulled barley above 300 seeds per m2 often 2A
resulted in lower yields and economic returns, and reduced
kernel plumpness. This loss in yield may reflect greater
lodging at higher seeding rates.
FERTILIZING BARLEY
The amount of fertilizer required for barley and other
crops grown in Canada is normally based on a soil test
recommendation. Nitrogen is usually the most limiting
nutrient in arable soils in Canada, and applications are 2C
required annually during the cropping season. The amount
required depends on a number of factors including the
amount of soil nitrate-nitrogen present, as well as the
mineralization potential and moisture content of the soil,
along with expected precipitation.
7
NUTRITIONAL COMPOSITION & FEED VALUE
T. Vasanthan1 and S.J. Meale2; 1University of Alberta (tv3@ualberta.ca); 2 Agriculture and Agri-Food Canada (smea4184@uni.sydney.edu.au)
The composition of starch components in the barley grainspecifically, the ratio of amylose to amylopectindetermines
its classification as either waxy or regular. Regular barley typically has a ratio of amylose to amylopectin of 1:3, whereas
barley with higher levels of amylopectin (up to 100%) is referred to as waxy barley. High-amylose barley is associated with
enzymatic resistance to digestion in swine and poultry, and thus slower glucose release and prolonged satiety. Increased
amylopectin is associated with faster digestion of starch to glucose, which may result in higher feed intake as a consequence
of rapid rises in insulin.
Water-soluble fibre in barley grains is composed primarily of non-starch polysaccharides, such as -glucan. Barley can range
widely in its -glucan content, with hulled varieties ranging from 1.2 to 6.7% and 4.6 to 7.3% DM for regular versus waxy
barley, and hulless varieties ranging from 2.8 to 7.3% DM (non-waxy) and 4.8 to 16.9% DM (waxy). High levels of water-
soluble dietary fibre can increase the viscosity of intestinal contents, slowing intestinal transit and delaying gastric emptying
(Webster, 1986), a characteristic that poses challenges if feed with high levels of soluble fiber are used as feed for poultry.
Conversely, -glucans have been shown to have hypocholesterolaemic properties (Naumann et al. 2006), a trait that is
desirable from a human nutrition perspective.
HULLESS
COMPONENT, % HULLED REGULAR HULLED WAXY HULLESS REGULAR CORN
WAXY
Starch 58.23 55.14 62.75 57.43 62.55
-glucan 4.34 5.77 4.77 7.69 -
ADF1 5.85 - 2.14 2.40 2.88
NDF2 18.49 - 10.28 8.10 9.11
Protein 12.44 12.79 14.41 15.24 8.24
Lipid 2.58 2.81 2.45 3.08 3.48
Acid detergent fibre
1
Neutral detergent fibre
2
9
PROTEIN & AMINO ACIDS Table 7. MACRONUTRIENTS IN BARLEY GRAIN
The value of protein in barley reflects its concentration,
amino acid composition and digestibility. The protein MINERAL % DRY MATTER
content of hulled non-waxy, hulled waxy, hulless non-waxy
P 0.27-0.49
and hulless waxy barley range from 7.3 to 18.2%, 10.5 to
16.1%, 9.6 to 21.9% and 11.3 to 21.6% DM, respectively. K 0.45-2.51
Barley protein has complex effects on grain quality, as high Ca 0.01-0.28
protein content results in reduced starch content, thereby Mg 0.10-0.24
decreasing overall carbohydrate content (Fox, 2010). When
S 0.14-0.19
compared to wheat, barley (hulled regular) has a lower
protein content (12.44%), but in terms of quality, barley Na 0.006-0.045
protein is comparable to wheat (Eggum, 1969). Barley has
a 4% higher protein content than corn grain (Table 5) and a VITAMINS & MINERALS
higher level of lysine (Table 6). The range in content of the main macronutrients present in
barley are presented in Table 7. Diets high in barley typically
Table 6. ESSENTIAL AMINO ACIDS IN BARLEY VS. NORMAL meet the phosphorus requirements of ruminants, but
CORN GRAIN AS % OF PROTEIN supplemental Ca in the diet is often required.
11
BARLEY IN RUMINANT DIETS
BEEF CATTLE
B. Lardner and G. Penner University of Saskatchewan (blardner.wbdc@pami.ca; greg.penner@usask.ca)
Barley grain is as an excellent energy source for beef cattle. For this reason, barley grain is generally fed to beef cows when
cows are grazing poor-quality pastures or consuming forages that do not have enough energy to meet requirements. While
barley can be used to increase the energy content of the diet, the extent of barley grain processing and level of feeding can
impact forage and total feed intake and forage utilization.
The processing index (PI) can be used as a practical tool to evaluate the extent of barley grain processing and is calculated
as follows:
Processing index (PI) = weight of processed sample / weight of unprocessed sample 100%
For both processed and unprocessed barley, the sample should be measured into a container so the same volume is used for
both weight measurements. It is recommended to process barley for beef cows to a similar extent as suggested for dairy cattle
and growing beef cattle, equating to a processing index ranging between 65 and 75% (Dehghan-banadaky, 2007; Table 9).
COW CALF
Barley grain is often provided to increase the energy intake of beef cattle. However, barley grain feeding also affects the
intake of forage as cattle preferentially consume barley over forage. Past studies have shown that as the level of barley
increases, the consumption of hay decreases (Fig. 3).
In addition, forage digestibility decreases with increasing rates of barley intake as the digestion of barley in the rumen may
decrease pH, reducing the digestion of fibre by shifting the bacterial species from a fibre-digesting community to one that is
more adept at starch digestion.
13
Fig. 3. The effect of level of barley grain supplementation on forage intake, total DMI and digestible organic matter intake.
No further increases in total DMI and only marginal increases in digestible organic matter were noted when barley exceeded
12.5% of total DMI (red line). Total DMI and only marginal increases for digestible organic matter intake. Data adapted from
Lardy et al. 2004.
As a consequence of reduced forage intake and forage digestibility, the benefit of barley supplementation on energy supply
may not be as great as expected. For example, research by Lardy et al. (2004) demonstrated that as barley supplementation
increased from 0.8 to 2.4 kg/d, there was no further improvement in digestible organic matter supply (Fig. 3). Thus, to
maximize the benefit from barley grain for forage-fed cattle, it is recommended barley grain be limited to approximately
12.5% of the total DM intake (DMI). If forage is not the main energy source in the diet, such as during a drought, higher
levels of barley can be utilized. Producers are encouraged to provide barley grain on a daily basis as infrequent provision
decreases the beneficial effects of cereal-based supplementation (Loy et al. 2008).
The period following weaning until the cattle enter a feedlot is known as the backgrounding or growing period. During this
time, cattle require relatively high dietary nutrient and energy contents to maximize growth. Typical backgrounding diets
contain feed grain and protein supplement together with ~50% of forage DM. Owing to its high energy content, barley is an
ideal grain for inclusion in the diets of backgrounding cattle. An additional benefit is that barley contains more protein than
corn, reducing the level of expensive supplemental protein that needs to be added to the diet.
Barley grain can be included in backgrounding diets at a maximum of 40 to 60% DM. To improve digestibility while
maintaining normal rumen function, barley grain should be properly processed before feeding to backgrounding beef
cattle. Whole barley is less digestible due to its resistant nature to ruminal microbes, whereas finely ground barley can be
rapidly utilized by ruminal bacteria for fermentation and may cause health issues, such as acidosis and bloat. Processing
barley via steam-rolling, temper-rolling, flaking or dry-rolling effectively reduces the risk of acidosis by modulating starch
degradation in the rumen (Stock and Britton, 1993). In Western Canada, dry-rolling is the major processing method used due
to its simplicity and cost-effective nature. The processing index should ideally be around 70 to 75% for a backgrounding diet
(Wang et al. 2003). An alternate processing method for maximizing barley utilization in backgrounding beef cattle diets is
pelleting. For example, pelleting barley blended with canola meal in a ratio of 85:15 improved feed efficiency compared to a
rolled barley diet on its own, with both diets containing ~30% of DM as barley.
15
FINISHING BEEF CATTLE
J. McKinnon University of Saskatchewan (john.mckinnon@usask.ca)
Barley grain is the predominant cereal grain fed to finishing cattle in Western Canada. This is a function of supply, price
and nutrient content. Nutrient characteristics that enhance barleys reputation as a feed grain include its relatively high
net energy value for maintenance (2.03 Mcal/kg DM) and gain (1.37 Mcal/kg DM) and crude protein value (12.5% DM). In
contrast, respective net energy values for cracked corn grain are 2.09 and 1.42 Mcal/kg DM and 9.5% DM for crude protein
(Table 10).
Despite barleys slightly lower net energy content than corn, the performance of barley-fed cattle is excellent and often
comparable to corn-fed cattle. To understand why, it is necessary to examine the nature of the barley and corn kernel. Unlike
corn, the starch in the endosperm of barley is surrounded by readily digestible protein that is rapidly digested once the hull
and pericarp is breached. Barley is often sold on the basis of bushel weight, which may range from 36 to 55 lbs per bushel.
Although lower-bushel-weight barley contains less starch than high-bushel-weight barley, gains of cattle are usually not
influenced by bushel weight as cattle compensate for the lower starch by consuming more barley. Feed efficiency can start to
decline at bushel weights below 40 lbs. Care must be taken to match the degree of barley processing to bushel weight as lots
of barley with different bushel weights also vary in kernel plumpness.
Table 10. COMPARISON OF ENERGY AND PROTEIN CONTENT OF TYPICAL CEREAL GRAINS ON A DM BASIS1
With corn, starch in regions of the endosperm is tightly bound to protein and, as a result, steam-flaking is required to
optimize the utilization of starch in corn. There are also differences between the two grains in terms of starch content and
rate of rumen fermentability. The barley kernel has less starch than corn, but the starch that it has is fermented at a much
faster rate. This can lead to increased digestion of starch in the rumen, enhancing the growth of rumen microbes, but
processing needs to be carefully managed to avoid the occurrence of digestive upsets, such as acidosis and bloat, which
have been linked to rapid ruminal starch digestion.
As with backgrounding cattle, the most common processing methods for barley are dry- and temper-rolling. Extensive
processing such as steam-flaking has not proven to be economically viable as the goal is to crack the outer hull and break
the kernel into two to four pieces. Too many whole kernels will result in poor digestibility, elevated feed intakes and poor
feed conversions. In contrast, too high a degree of processing results in excessive fines (grain particles <2 mm in size), which
can lead to digestive disturbances. While there are no hard and fast guidelines as to an acceptable level of fines, values
greater than 5% can indicate that the grain has been over-processed. Additionally, barley is often processed to a PI of 75 to
80% for finishing cattle, with lower PI used with higher levels of forage in the diet. Lower values indicate a more aggressive
feeding program and the need to pay closer attention to bunk management. Higher values indicate a less aggressive
feeding program and potentially issues that can lead to poor feed efficiency if whole kernels pass through the digestive tract.
Further confirmation of optimal processing can be gained by examining the nature of the feces in terms of consistency
A second key to optimizing the performance of barley-fed cattle is an effective bunk management program. The goal is to keep
cattle eating at a high level on a consistent basis over an extended period. Due to the rapid fermentation of barley starch, this
can be a challenge. Keys to sound bunk management include a step-up feeding program that allows for a smooth transition from
forage to high-barley-grain diets. This transition is usually accomplished through the use of four to six step-up diets where the
level of forage is gradually decreased and the level of barley grain gradually increased. A well-designed bunk reading and feed
delivery program along with an effective grain processing and monitoring program are keys to successfully feeding barley. Most
importantly, one needs effective communication between all staff involved in the feeding of the cattle.
In summary, barley grain can be used as the sole cereal grain for finishing cattle with excellent results in terms of performance,
carcass and meat quality. Comparative studies have shown that barley-fed cattle produce meat of equal quality to those fed corn.
Optimizing the performance of barley-fed cattle is a function of sound management including knowledge of nutrient content and
rumen fermentability in the design and implementation of the feed program.
DAIRY COWS
M. Oba University of Alberta (masahito.oba@ualberta.ca)
Barley grain is widely used as a primary energy source in diets of dairy cows in Canada. Understanding its nutrient and
digestive characteristics is very important for formulating diets for lactating dairy cows that will maximize their productivity.
The level of inclusion of barley grain in dairy cattle diets is dependent on stage of lactation, body weight and other factors.
Consideration also needs to be given to the nature and level of forage included, as higher forage diets and longer chopped
forage or forage with a slower rate of fermentation can allow for higher rates of barley to be included. As with other cattle,
whole unprocessed barley grain is poorly digested, and processing is necessary to make the starch accessible to microbes
in the rumen of dairy cows. Christen et al. (1996) reported an increase in milk yield of 1.8 kg/d when temper-rolled barley
replaced dry-rolled barley in a diet of dairy cows, due to greater whole tract digestibility. Contrarily, feeding tempered barley
without rolling decreased milk yield by 2 kg/d compared to dry-rolled barley, so tempered barley must be rolled prior to
feeding. Intensive dry-rolling of barley can also generate the fine particles that can lead to a reduction in milk production as
a result of acidosis.
17
Table 11. PRODUCTIVITY AND TOTAL TRACT STARCH DIGESTIBILITY OF COWS FED STEAM-ROLLED BARLEY GRAIN VARYING
IN EXTENT OF PROCESSING1
P LINEAR QUADRATIC
COARSE MEDIUM M-FLAT FLAT
VALUE CONTRASTS CONTRASTS
DMI, kg/d 18.7 21.4 21.7 20.1 0.60 0.12 < 0.01
Milk yield, kg/d 25.6 28.1 30.8 29.0 0.40 < 0.01 < 0.01
Milk fat, % 3.93 3.89 3.78 3.9 0.06 0.5 0.25
Milk protein, % 3.15 3.30 3.29 3.34 0.02 < 0.01 < 0.05
Total tract starch digestibility, % 78.0 84.1 93.6 92.9 1.70 < 0.01 0.10
Barley grain is often steam-rolled prior to feeding to dairy cattle as it is frequently processed in large commercial feed
mills that possess this equipment. As with beef cattle, it is important to define the optimal degree of processing because
maximizing fermentation of barley in the rumen does not necessarily maximize milk yield. Yang et al. (2000) fed barley grain,
steam-rolled to coarse, medium, medium-flat and flat (PI = 81.0, 72.5, 64.0, and 55.5%, respectively) in diets of lactating dairy
cows. Total tract starch digestibility and milk yield increased linearly as the PI decreased from 81.0 to 64.0% (Table 11). However,
further processing, indicated by reduction in the PI from 64.0 to 55.5%, did not increase total tract starch digestibility, but
decreased DMI and milk yield. As a result, it was concluded that the optimum PI for barley grain fed to lactating dairy cows
was 64% as it maximized milk yield.
However, the optimum extent of processing is expected to differ depending on the quality of barley grain prior to
processing. In a similar study (McGregor et al. 2007), barley grain was steam-rolled to either 82.5 or 68.7% PI, and fed to
lactating dairy cows.
Dry matter intake and milk yield were not affected by the extent of processing in this study, a discrepancy that might be
attributed to differences in physical and chemical characteristics of barley. Barley grain used by Yang (2000) had a bushel
weight of 44.4 lb and 26.5% NDF while McGregors (2007) had a bushel weight of 53.1 lb and 16.8% NDF. Barley grain with
low NDF content might require less processing, while highly fibrous barley grain may need more extensive processing to
optimize rumen fermentation and digestibility.
Schlau et al. (2013) conducted a similar study evaluating two lots of barley grain that also differed in starch content (Table 12),
but reported that milk yield was not affected by grain treatment. One possible reason for the discrepancy between the two
studies may be related to milk production level; Silveira et al. (2007) used cows in peak lactation producing more than 40
kg/d of milk, while Schlau et al. (2013) used late-lactating cows producing less than 30 kg/d of milk.
Maximum milk production of cows at peak lactation requires higher energy intakes, which may have been satisfied by high-
starch barley leading to increased milk production. Cows in the study of Schlau et al. (2013) may have consumed sufficient
energy with the moderate-starch barley grain, and thus the use of higher-starch barley did not result in a further increase in
milk production.
In summary, processed barley grain is highly fermentable in the rumen and an important energy source in dairy diets.
Feeding value of barley grain varies greatly among lots, and can affect productivity of high-producing dairy cows, in which
energy intake limits maximum milk production. Processing method and the extent of processing also affect milk production
of dairy cows, particularly if poor-quality barley grain is used. Feeding highly fermentable grains, such as barley, can enhance
milk production by increasing energy intake and metabolizable protein supply if diets are formulated properly, but may
decrease feed intake and milk production if it leads to subclinical or clinical acidosis. The feeding value of barley grain and its
optimum utilization are greatly affected by how it is processed and how it is incorporated into the diet.
19
OTHER RUMINANTS
K. Stanford Alberta Agriculture Food and Rural Development (kim.stanford@gov.ab.ca)
Barley is a common and valuable feedstuff for lambs, sheep and goats. As small ruminants chew whole grains more
thoroughly than cattle, the costs associated with processing are generally not recovered through improvements in the
efficiency of sheep and goats (Morgan et al. 1991; McGregor & Whiting, 2013). Feeding whole barley to small ruminants also
avoids the digestive disturbances that can occur with highly processed or pelleted barley diets.
Lambs
The extent to which barley grain is fed to lambs depends on the marketing strategy that is
being employed. Concentrate diets for finishing lambs promote rapid growth (Sormunen-
Cristian 2013), but may result in undesirable carcass fatness (Priolo et al. 2002). Accordingly,
Hatfield et al. (1997) determined that higher levels of whole barley (up to 90% of the diet)
resulted in fatter lamb carcasses as compared to diets that contained 50 or 70% whole
barley. However, as with cattle, the rapid and thorough rumen fermentation of barley grain
results in greater microbial protein synthesis than in corn-based diets, and the supplemental
protein required as outlined by the National Research Council (NRC) to meet the crude
protein requirement for lambs is higher for corn- than barley-based diets. Accordingly,
Stanford et al. (1995) determined that rapidly growing lambs fed diets consisting of 76%
barley required 32% less crude protein than NRC estimates.
Goats
Although barley grain is highly palatable to goats, kids and mature goats are less prone to
overeating than lambs and sheep (Fedele et al. 2002) and excess fat is uncommon in goat
carcasses. High-producing dairy goats and rapidly growing kids are commonly fed large
amounts of concentrate to meet their energy demands. Despite the lower pH associated
with increased volatile fatty acids production in the rumen, subacute ruminal acidosis was
not detected in growing kids fed up to 60% whole barley (Klevenhusen et al. 2013). Barley
grain has been found to be equivalent to corn with regard to milk yield and composition
of milk produced by dairy goats. Given a choice, dairy goats preferred barley grain over
corn, although proportion of hay consumed by does also increased relative to that of
concentrates (Avondo et al. 2013).
23
BREEDING GILTS & SOWS
Gestating sows are fed restrictively to prevent excess weight gain and therefore can be fed high-fibre diets that also mitigate
the incidence of undesirable chewing behaviours in sows housed in stalls. Gestating gilts and sows can thus be fed barley-
based diets with minimal supplemental protein as canola meal, dried distillers grains or soybean meal. Lactating sows can
also be fed barley-based diets, but it is usually mixed with wheat grain to further increase the energy content of the diet
(Table 15).
Table 15. PROCESSING METHODS AND MAXIMUM INCLUSION RATES OF BARLEY GRAIN IN SWINE, OTHER LIVESTOCK AND
AQUACULTURE DIETS
INTRODUCTION
Barley is suitable for inclusion in the diet of all types and gut fill, then poultry should be able to consume enough
ages of poultry (Table 15). The primary limitation to its of a barley-based diet to meet their dietary requirements.
inclusion, as with any dietary ingredient, is the cost per However, as a result of the hull, feed conversion (g feed to g
kilogram of animal protein (meat or eggs) produced. This gain) will be ~15% higher; thus, to be economically feasible,
means it may be more profitable to have poorer feed barley costs need to be comparably lower than alternative
conversion using a cheaper feed. The costs associated with grains with a lower fibre content.
barley are based on nutrient level and availability, which
vary with the source of barley. With poultry, however, The energy levels of barley are primarily dependent on the
the cost associated with reducing antinutritional factors level, digestibility and utilization of starch and lipids.
also needs to be considered. The majority of the barley Scott et al. (1998) determined that the energy content of
available as poultry feed is hulled, but the lower fibre 14 cultivars of barley (hulled and hulless) in young broilers
content of hulless barley is especially advantageous for ranged from 2,800 to 3,320 kcal/kg when no fibrolytic
poultry, making it competitive with corn or wheat as an -glucanase enzymes were present, but increased to 3,240
energy source. to 3,570 kcal/kg with enzyme addition, resulting in an
overall increase in AME of 14%. As expected, this response
NUTRIENT LEVEL & AVAILABILITY was higher for hulless barley, due to higher levels of soluble
The primary considerations in formulating poultry diets non-starch polysaccharides (i.e., -glucan). The Australian
are metabolizable energy content (AME; kcal/kg) and the Premium Grains for Livestock Program (Black, 2008)
availability of amino acids, particularly lysine, to produce measured the energy from 38 barley samples (without
poultry protein, expressed per kilogram of barley. Nutrient enzyme) in broilers (2,360 to 2,940 kcal/kg) and layers (2,630
levels in barley are diluted by the hull, which accounts for to 3,530 kcal/kg) and noted that overall digestibility was
~15% of the grains weight, adding considerable bulk to a higher in layers than broilers, possibly due to the longer
diet. Theoretically, if there are no limits on intake due to residency time of digesta in the digestive tract of layers.
The recommendations are to always use barley enzyme Combinations of feed enzymes, often referred to as enzyme
supplements for broilers and young turkeys. However, cocktails, are commercially available. Selection should be
there is less digestive upset for older birds and benefits based on the specific enzymes needed and their minimum
should be evaluated prior to use in these situations. Scott et activity levels, but generally, little advantage has been
al. (2001) demonstrated that if intake of barley-based diets observed with the addition of enzyme cocktails that contain
without enzymes was slightly restricted in young broilers, proteases (which break up protein) or cellulases (which
-glucanases offered less benefit, although digesta viscosity break down fibre). Feed processing also affects enzyme
of non-supplemented diets was high. This suggests that efficacy as grinding, conditioning and pelleting result in
viscosity is more of a problem when birds are eating to higher solubilization of non-starch polysaccharides in feed
gut capacity, as would be the case in broilers and young ingredients, and heating can destroy both dietary and
turkeys. In layers, the higher moisture in the digesta with supplemental enzymes.
27
FEEDING WHOLE BARLEY GRAIN OTHER CONSIDERATIONS WHEN FEEDING
If implemented correctly, feeding whole barley grains can A HIGH-BARLEY DIET
reduce costs associated with feed processing. In broilers, Similarly to the effect in swine, high-barley diets can
the practice requires a gradual introduction of whole increase the whiteness and hardness of fat as compared to
grain and careful attention to proper mixing, so that all corn. This is also likely to result in paler yolks in eggs from
birds receive feed of uniform nutritive value that meets layers. This can be overcome by supplementing pigments in
their requirements. the diet; however, the cost should be rationalized with the
value of the finished product and consumer preference.
Bennett et al. (2002a) reported that whole barley grain Small flock owners may also consider steeping barley
could be included in broiler diets at up to 35% DM and at (approximately one part by weight feed to 1.0 or 1.2 parts
least 20% DM for growing turkeys (Bennett et al. 2002b). water) for 15 to 30 min just before feeding to form a
Whole-grain feeding for laying hens and broiler breeders feed with a porridge-like consistency. This approach has
(either in the diet or spread on the litter) is associated significantly increased intake and growth of broilers and
with reduced behaviour problems, such as feather minimized the need to supplement with enzymes. Care
pecking and cannibalism. must be taken to ensure the feed is fed immediately after
the steeping process to avoid mould growth.
INTRODUCTION
Barley grain can be included in the diets of horses, rabbits and fish to provide energy and nutrients. The level of inclusion
needs to consider the digestive physiology of the animal and its ability to digest fibre. Barley is recommended for inclusion
in the diet of high-performance horses due to its high energy content. However, care needs to be taken in order to properly
process the grain and to formulate a well-balanced diet that includes sufficient forage. For rabbits, barley can be the primary
dietary component due to its moderate protein and fibre content as compared with other major grains. Barley grain can also
be included in fish diets as it provides starch as an energy source, but requires extrusion and pelleting to form pellets that float.
Concentration of barley protein has also been used as a partial replacement for fish meal and soybean meal in fish diets.
HORSES
Under certain production (e.g., growth, pregnancy, and lactation) and performance conditions, nutrient requirements
of horses far exceed those provided by forage. Barley grain is a good source of protein and energy; however, enzymatic
hydrolysis of starch in whole grains is limited in the small intestine of horses. As such, processing of the barley (Table 15) is
required to increase the absorption of starch in the small intestine. Barley starch that reaches the large intestine is rapidly
fermented by resident microbes and the acid produced can lead to intestinal disturbances such as colic and laminitis.
It has been recommended that barley grain be fed to horses at a maximum 0.2% of body weight as total non-structural
carbohydrates (TNC), when included with alfalfa cubes to prevent negative effects on horse health and digestion (Hussein et al.
2004). Barley grain can also be fed to horses in combination with oats and corn.
Digestion of starch and sugar in grains provides the main substrate for muscle glycogen synthesis after exercise.
Hydrolyzable carbohydrate from rolled barley grain has a glycemic index of approximately 60 compared to 100 in pure
glucose (Jose-Cunilleras et al. 2004). Consequently, barley is a good substrate for muscle glycogen synthesis and thus a
suitable feed for high-performance horses.
RABBITS
Barley is a viable grain source for all aspects of rabbit production. As barley contains more fibre than wheat or corn, it may
be a more desirable grain source for rabbits (Acedo-Rico et al. 2010). Barley grain can be included in the diets of growing
rabbits at 10 to 25% DM (de Blas et al. 2010) with a maximum level of 40 to 45% DM (Seroux, 1984; Table 15). For breeding
rabbits, recommended inclusion rates of barley are 25 to 40% DM (Pascual et al. 1998) with a maximum rate of 64% DM
(Prasad et al. 1998). Flaking or grinding of barley has been reported to have no added benefit for the nutritional value of
barley for rabbits and thus, as with sheep, it can be fed to rabbits whole.
AQUACULTURE
Barley grain is included in fish diets as a starch source for energy (Table 15). Starch in extruded pellets also helps contribute
to their buoyancy. Energy and nutrient digestibility of barley grains (30% DM) with various starch types (waxy vs. normal)
were compared as feed for rainbow trout. Results indicated that waxy barley had higher digestible energy than non-waxy
barley (Gaylord et al. 2009). Barley meal (50% DM) has also been included in diets of common carp and tilapia (Degani et
al. 1997a, b). Barley meal provides 6.69 and 14.36 kJ digestible energy per g of barley to common carp and adult tilapia,
respectively. Barley grain has lower protein and amino acid availability than fish meal for rainbow trout (Gaylord et al. 2010).
Development of a barley protein concentrate has shown promise as a protein source in pilot feeding studies with salmon
and trout (Durham, 2010).
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