Cat

The domestic cat[1][5] (Felis silvestris catus or Felis catus) is a small,

Domestic cat[1]
typically furry, carnivorous mammal. They are often called house cats[6]
when kept as indoor pets or simply cats when there is no need to distinguish
them from other felids and felines. They are often valued by humans for
companionship and for their ability to hunt vermin. There are more than
seventy cat breeds recognized by variouscat registries.

Cats are similar in anatomy to the other felids, with a strong flexible body,
quick reflexes, sharp retractable claws, and teeth adapted to killing small prey.
Cat senses fit a crepuscular and predatory ecological niche. Cats can hear
sounds too faint or too high in frequency for human ears, such as those made
by mice and other small animals. They can see in near darkness. Like most Various types of domestic cat
other mammals, cats have poorer color vision and a better sense of smell than Conservation status
humans. Cats, despite being solitary hunters, are a social species and cat
Domesticated
communication includes the use of a variety of vocalizations (mewing,
purring, trilling, hissing, growling, and grunting), as well as cat pheromones Scientific classification
and types of cat-specific body language.[7] Kingdom: Animalia
Cats have a high breeding rate.[8] Under controlled breeding, they can be bred Phylum: Chordata
and shown as registered pedigree pets, a hobby known as cat fancy. Failure to
Class: Mammalia
control the breeding of pet cats by neutering, as well as the abandonment of
former household pets, has resulted in large numbers of feral cats worldwide, Order: Carnivora
requiring population control.[9] In certain areas outside cats' native range, this Suborder: Feliformia
has contributed, along with habitat destruction and other factors, to the
Family: Felidae
extinction of many bird species. Cats have been known to extirpate a bird
species within specific regions and may have contributed to the extinction of Genus: Felis
isolated island populations.[10] Cats are thought to be primarily responsible Species: F. silvestris
for the extinction of 33 species of birds, and the presence of feral and free-
ranging cats makes some otherwise suitable locations unsuitable for attempted
Subspecies: F. s. catus
species reintroduction.[11] Trinomial name

Since cats were venerated in ancient Egypt, they were commonly believed to Felis silvestris catus
have been domesticated there,[12] but there may have been instances of Linnaeus , 1758 [2]
domestication as early as the Neolithic from around 9,500 years ago (7,500
Synonyms
BC).[13] A genetic study in 2007[14] concluded that all domestic cats are
descended from Near Eastern wildcats, having diverged around 8,000 BC in Felis catus (original combination)[3]
the Middle East.[12][15] A 2016 study found thatleopard cats were undergoing Felis catus domestica (invalid junior synonym)[4]
domestication independently in China around 5,500 BC, though this line of
partially domesticated cats leaves no trace in the domesticated populations of
today.[16][17] A 2017 study confirmed that domestic cats are descendants of those first domesticated by farmers in the Near East
around 9,000 years ago.[18][19]

As of a 2007 study, cats are the second most popular pet in the US by number of pets owned, behind freshwater fish.[20] In a 2010
[21]
study they were ranked the third most popular pet in the UK, after fish and dogs, with around 8 million being owned.
 Contents
Taxonomy and evolution
Nomenclature and etymology
Biology
Anatomy
Physiology
Nutrition
Senses
Health
Diseases
Poisoning
Genetics
Behavior
Sociability
Communication
Grooming
Fighting
Hunting and feeding
Play
Reproduction
Ecology
Habitats
Feral cats
Impact on prey species
Impact on birds
Interaction with humans
Cat show
Cat café
Ailurophobia
Cat bites
Infections transmitted from cats to humans
History and mythology
Superstitions and cat burning

See also
References
External links

Taxonomy and evolution

The domestic cat is believed to have evolved from the Near Eastern wildcat, whose range covers vast portions of the Middle East
westward to the Atlantic coast of Africa.[22][23] Between 70,000 and 100,000 years ago the animal gave rise to the genetic lineage
that eventually produced all domesticated cats,[24] having diverged from the Near Eastern wildcat around 8,000 BC in the Middle
East.[12][15]

The felids are a rapidly evolving family of mammals that share a common ancestor only 10–15 million years ago[25] and include
lions, tigers, cougars and many others. Within this family, domestic cats (Felis catus) are part of the genus Felis, which is a group of
small cats containing about seven species (depending upon classification scheme).[1][26] Members of the genus are found worldwide
and include the jungle cat (Felis chaus) of southeast Asia, European wildcat (F. silvestris silvestris), African wildcat (F. s. lybica), the
Chinese mountain cat (F. bieti), and the Arabian sand cat (F. margarita), among others.[27]
The domestic cat was first classified as Felis catus by Carl Linnaeus in the 10th edition of his Systema Naturae published in
1758.[1][2] Because of modern phylogenetics, domestic cats are usually regarded as another subspecies of the wildcat, F.
silvestris.[1][28][29] This has resulted in mixed usage of the terms, as the domestic cat can be called by its subspecies name, Felis
silvestris catus.[1][28][29] Wildcats have also been referred to as various subspecies of F. catus,[29] but in 2003, the International
Commission on Zoological Nomenclature fixed the name for wildcats as F. silvestris.[30] The most common name in use for the
domestic cat remains F. catus. Sometimes, the domestic cat has been called Felis domesticus[31] as proposed by German naturalist
J.C.P. Erxleben in 1777,[32] but these are not valid taxonomic names and have been used only rarely in scientific literature.[33] A
population of Transcaucasian black feral cats was once classified as Felis daemon (Satunin 1904) but now this population is considered
to be a part of domestic cat.[34]

All the cats in this genus share a common ancestor that is believed to have lived around 6–7 million years ago in the Near East (the
Middle East).[35] The exact relationships within the Felidae are close but still uncertain,[36][37] e.g. the Chinese mountain cat is
sometimes classified (under the name Felis silvestris bieti) as a subspecies of the wildcat, like the North African variety F. s.
lybica.[28][36]

In comparison to dogs, cats have not undergone major changes during the
domestication process, as the form and behavior of the domestic cat is not radically
different from those of wildcats and domestic cats are perfectly capable of surviving
in the wild.[38][39] Fully domesticated house cats often interbreed with feral F. catus
populations,[40] producing hybrids such as the Kellas cat. This limited evolution
during domestication means that hybridisation can occur with many other felids,
notably the Asian leopard cat.[41] Several natural behaviors and characteristics of
wildcats may have predisposed them for domestication as pets.[39] These traits
include their small size, social nature, obvious body language, love of play and
relatively high intelligence.[42]:12–17 Several small felid species may have an inborn
tendency towards tameness.[39]

Cats have either a mutualistic or commensal relationship with humans. Two main
theories are given about how cats were domesticated. In one, people deliberately
tamed cats in a process of artificial selection as they were useful predators of
vermin.[43] This has been criticized as implausible, because the reward for such an Ancient Egyptian sculpture of the cat
effort may have been too little; cats generally do not carry out commands and goddess Bastet. The earliest
although they do eat rodents, other species such as ferrets or terriers may be better at evidence of felines as Egyptian
deities comes from c. 3100 BC.
controlling these pests.[28] The alternative idea is that cats were simply tolerated by
people and gradually diverged from their wild relatives through natural selection, as
[28]
they adapted to hunting the vermin found around humans in towns and villages.

Nomenclature and etymology

The origin of the English word cat (Old English catt) and its counterparts in other Germanic languages (such as German Katze),
descended from Proto-Germanic *kattōn-, is controversial. It has traditionally thought to be a borrowing from Late Latin cattus
"domestic cat", from catta (used around 75 AD by Martial),[44][45] compare also Byzantine Greek κάττα, Portuguese and Spanish
gato, French chat, Maltese qattus, Lithuanian katė, and Old Church Slavonic kotъ (kotka), among others.[46] The Late Latin word is
generally thought to originate from an Afro-Asiatic language, but every proposed source word has presented problems. Many
references refer to "Berber" (Kabyle) kaddîska "wildcat" and "Nubian kadīs" as possible sources or cognates, but M. Lionel Bender
says the Nubian term is a loan from Arabic ‫ ﻗِﻄﺔ‬qiṭṭa.[47] Jean-Paul Savignac suggests the Latin word is from an Egyptian precursor
of Coptic ϣ (šau) "tomcat" or its feminine form suffixed with -t,[48] but John Huehnergard says "the source [...] was clearly not
[47] Huehnergard opines it is "equally likely that the forms might derive from an
Egyptian itself, where no analogous form is attested."
ancient Germanic word, imported into Latin and thence to Greek and to Syriac and Arabic".
Guus Kroonen also considers the word to
be native to Germanic (due to morphological alternations) and Northern Europe, and suggests that it might ultimately be borrowed
from Uralic, cf. Northern Sami gađfe "female stoat" and Hungarian hölgy "stoat; lady, bride" from Proto-Uralic *käďwä "female (of a
fur animal)".[49] In any case, cat is a classic case of a Wanderwort.

An alternative word is English puss (extended as pussy and pussycat). Attested only from the 16th century, it may have been
introduced from Dutch poes or from Low German puuskatte, related to Swedish kattepus, or Norwegian pus, pusekatt. Similar forms
exist in Lithuanian puižė and Irish puisín or puiscín. The etymology of this word is unknown, but it may have simply arisen from a
sound used to attract a cat.[50][51]

A group of cats is referred to as a clowder or a glaring,[52] a male cat is called a tom or tomcat[53] (or a gib,[54] if neutered), an
unaltered female is called a queen,[55] and a juvenile cat is referred to as a kitten. The male progenitor of a cat, especially a pedigreed
cat, is its sire,[56] and its female progenitor is itsdam.[57] In Early Modern English, the word kitten was interchangeable with the now
obsolete word catling.[58]

A pedigreed cat is one whose ancestry is recorded by a cat fancier organization. A purebred cat is one whose ancestry contains only
individuals of the same breed. Many pedigreed and especially purebred cats are exhibited as show cats. Cats of unrecorded, mixed
ancestry are referred to as domestic short-haired or domestic long-haired cats, by coat type, or commonly as random-bred, moggies
(chiefly British), or (using terms borrowed fromdog breeding) mongrels or mutt-cats.

While the African wildcat is the ancestral subspecies from which domestic cats are descended, and wildcats and domestic cats can
completely interbreed (being subspecies of the same species), several intermediate stages occur between domestic pet and pedigree
cats on one hand and entirely wild animals on the other. The semiferal cat, a mostly outdoor cat, is not owned by any one individual,
but is generally friendly to people and may be fed by several households. Feral cats are associated with human habitation areas and
[40]
may be fed by people or forage for food, but are typically wary of human interaction.

Biology

Anatomy
Domestic cats are similar in size to the other members of
the genus Felis, typically weighing between 4 and 5 kg (9
and 10 lb).[36] Some breeds, such as the Maine Coon, can
occasionally exceed 11 kg (24 lb). Conversely, very small
cats, less than 2 kg (4 lb), have been reported.[59] The
world record for the largest cat is 21 kg (50 lb).[60] The
smallest adult cat ever officially recorded weighed around
1 kg (2 lb).[60] Feral cats tend to be lighter as they have
Diagram of the generalanatomy of a male
more limited access to food than house cats. In the Boston
area, the average feral adult male will weigh 4 kg (9 lb) and
average feral female 3 kg (7 lb).[61] Cats average about 23–25 cm (9–10 in) in height and 46 cm (18 in) in head/body length (males
being larger than females), with tails averaging 30cm (12 in) in length.[62]

Cats have seven cervical vertebrae, as do almost all mammals; 13 thoracic vertebrae (humans have 12); seven lumbar vertebrae
(humans have five); three sacral vertebrae like most mammals (humans have five); and a variable number of caudal vertebrae in the
tail (humans retain three to five caudal vertebrae, fused into an internal coccyx).[63]:11 The extra lumbar and thoracic vertebrae
account for the cat's spinal mobility and flexibility. Attached to the spine are 13 ribs, the shoulder, and the pelvis.[63] :16 Unlike
human arms, cat forelimbs are attached to the shoulder by free-floating clavicle bones which allow them to pass their body through
any space into which they can fit their head.[64]
 The cat skull is unusual among mammals in having very large eye sockets and a
powerful and specialized jaw.[65]:35 Within the jaw, cats have teeth adapted for
killing prey and tearing meat. When it overpowers its prey, a cat delivers a lethal
neck bite with its two long canine teeth, inserting them between two of the prey's
vertebrae and severing its spinal cord, causing irreversible paralysis and death.[66]
Compared to other felines, domestic cats have narrowly spaced canine teeth, which
is an adaptation to their preferred prey of small rodents, which have small
vertebrae.[66] The premolar and first molar together compose the carnassial pair on
Cat skull each side of the mouth, which efficiently shears meat into small pieces, like a pair of
scissors. These are vital in feeding, since cats' small molars cannot chew food
effectively, and cats are largely incapable of mastication.[65]:37 Although cats tend
to have better teeth than most humans, with decay generally less likely because of a thicker protective layer of enamel, a less
damaging saliva, less retention of food particles between teeth, and a diet mostly devoid of sugar, they are nonetheless subject to
occasional tooth loss and infection.[67]

Cats, like dogs, are digitigrades. They walk directly on their toes, with the bones of their feet making up the lower part of the visible
leg.[68] Cats are capable of walking very precisely because, like all felines, they directly register; that is, they place each hind paw
(almost) directly in the print of the corresponding fore paw, minimizing noise and visible tracks. This also provides sure footing for
their hind paws when they navigate rough terrain. Unlike most mammals, when cats walk, they use a "pacing" gait; that is, they move
the two legs on one side of the body before the legs on the other side. This trait is shared with camels and giraffes. As a walk speeds
up into a trot, a cat's gait changes to be a "diagonal" gait, similar to that of most other mammals (and many other land animals, such
.[69]
as lizards): the diagonally opposite hind and fore legs move simultaneously

Like almost all members of the Felidae, cats have protractable and retractable claws.[70] In their normal, relaxed position, the claws
are sheathed with the skin and fur around the paw's toe pads. This keeps the claws sharp by preventing wear from contact with the
ground and allows the silent stalking of prey. The claws on the fore feet are typically sharper than those on the hind feet.[71] Cats can
voluntarily extend their claws on one or more paws. They may extend their claws in hunting or self-defense, climbing, kneading, or
for extra traction on soft surfaces. Most cats have five claws on their front paws, and four on their rear paws.[72] The fifth front claw
(the dewclaw) is proximal to the other claws. More proximally is a protrusion which appears to be a sixth "finger". This special
feature of the front paws, on the inside of the wrists, is the carpal pad, also found on the paws of big cats and dogs. It has no function
in normal walking, but is thought to be an antiskidding device used while jumping. Some breeds of cats are prone to polydactyly
(extra toes and claws).[72] These are particularly common along the northeast coast of North America.
[73]

Physiology
Cats are familiar and easily kept animals, and their physiology has been particularly well studied; it generally resembles those of
other carnivorous mammals, but displays several unusual features probably attributable to cats' descent from desert-dwelling
species.[31] For instance, cats are able to tolerate quite high temperatures: Humans generally start to feel uncomfortable when their
skin temperature passes about 38 °C (100 °F), but cats show no discomfort until their skin reaches around 52 °C (126 °F),[65]:46 and
.[74]
can tolerate temperatures of up to 56 °C (133 °F) if they have access to water

Normal physiological values[75]:330

Body temperature 38.6 °C (101.5 °F)
Heart rate 120–140 beats per minute
Breathing rate 16–40 breaths per minute

Cats conserve heat by reducing the flow of blood to their skin and lose heat by
Thermograph of various body parts evaporation through their mouths. Cats have minimal ability to sweat, with glands
of a cat
located primarily in their paw pads,[76] and pant for heat relief only at very high
temperatures[77] (but may also pant when stressed). A cat's body temperature does
not vary throughout the day; this is part of cats' general lack of circadian rhythms and may reflect their tendency to be active both
during the day and at night.[78]:1 Cats' feces are comparatively dry and their urine is highly concentrated, both of which are
adaptations to allow cats to retain as much water as possible.[31] Their kidneys are so efficient, they can survive on a diet consisting
only of meat, with no additional water,[79] and can even rehydrate by drinking seawater.[80][78]:29While domestic cats are able to
[81]
swim, they are generally reluctant to enter water as it quickly leads to exhaustion.

Nutrition
Cats are obligate carnivores: their physiology has evolved to efficiently process meat, and they have difficulty digesting plant
matter.[31] In contrast to omnivores such as rats, which only require about 4% protein in their diet, about 20% of a cat's diet must be
protein.[31] A cat's gastrointestinal tract is adapted to meat eating, being much shorter than that of omnivores and having low levels of
several of the digestive enzymes needed to digest carbohydrates.[82] These traits severely limit the cat's ability to digest and use
plant-derived nutrients, as well as certain fatty acids.[82] Despite the cat's meat-oriented physiology, several vegetarian or vegan cat
foods have been marketed that are supplemented with chemically synthesized taurine and other nutrients, in attempts to produce a
complete diet. However, some of these products still fail to provide all the nutrients cats require,[83] and diets containing no animal
products pose the risk of causing severe nutritional deficiencies.[84] However, veterinarians in the United States have expressed
concern that many domestic cats are overfed.[85]

Cats do eat grass occasionally. A proposed explanation is that cats use grass as a source of folic acid. Another is that it is used to
vomit.[86]
supply dietary fiber, helping the cat defecate more easily and expel parasites and other harmful material through feces and

Cats are unusually dependent on a constant supply of the amino acid arginine, and a diet lacking arginine causes marked weight loss
and can be rapidly fatal.[87] Arginine is an essential additive in cat food because cats have low levels of the enzymes
aminotransferase and pyrroline-5-carboxylate which are responsible for the synthesis of ornithine and citrulline in the small
intestine.[88] Citrulline would typically go on to the kidneys to make arginine, but because cats have a deficiency in the enzymes that
make it, citrulline is not produced in adequate quantities to make arginine. Arginine is essential in the urea cycle in order to convert
the toxic component ammonia into urea that can then be excreted in the urine. Because of its essential role, deficiency in arginine
results in a build up of toxic ammonia and leads to hyperammonemia.[88] The symptoms of hyperammonemia include lethargy,
vomiting, ataxia, hyperesthesia and can be serious enough to induce death andcoma in a matter of days if a cat is being fed a arginine
free diet. The quick onset of these symptoms is due to the fact that diets devoid in arginine will typically still contain all of the other
amino acids, which will continue to be catabolized by the body producing mass amounts of ammonia that very quickly build up with
no way of being excreted.

Another unusual feature is that the cat cannot produce taurine, with a deficiency in this essential amino acid causing macular
degeneration, wherein the cat's retina slowly breaks down, causing irreversible blindness.[31] This is due to the hepatic activity of
cystinesulfinic acid decarboxylase being low in cats.[89] This limits the ability of cats to biosynthesize the taurine they need from its
precursor amino acids ethionine and cysteine, which ultimately results in inadequate taurine production needed for normal
function.[89] Deficiencies in taurine result in compensated function of feline cardiovascular and reproductive systems.[89] These
abnormalities can also be accompanied by developmental issues in the central nervous system along with degeneration of the
retina.[89]

In order to produce the essential vitamin niacin for use in the cat, tryptophan is needed for conversion purposes. However, due to a
competing pathway withacetyl coenzyme A (acetyl-CoA), niacin can become deficient and requiresupplementation.[90] This process
occurs when an overactive enzyme, picolinic acid carboxylase, converts the vitamin B6 precursor picolinic acid into the alternate
compound acetyl-CoA, instead of converting quinolinate into nictotinic acid mononlucleotide (niacin).[91] Niacin is required in cats
as it supports enzyme function. If niacin is deficient in the diet, anorexia, weight loss and an increase in body temperature can
result.[92]

Preformed vitamin A is required in the cat for retinal and reproductive health. Vitamin A is considered to be a fat-soluble vitamin and
is seen as essential in a cat's diet. Normally, the conversion of beta-carotenes into vitamin A occurs in the intestine (more specifically
the mucosal layer) of species, however cats lack the ability to undergo this process.[90] Both the kidney and liver are contributors to
the use of vitamin A in the body of the majority of species while the cats liver does not produce the enzyme Beta-carotene 15,15'-
monooxygenase which converts the beta-carotene into retinol (vitamin A).[93] To summarize: cats do not have high levels of this
enzyme leading to the cleavage and oxidation ofcarotenoids not taking place.[91]

Vitamin D3 is a dietary requirement for cats as they lack the ability to synthesize vitamin D3 from sunlight.[94] Cats obtain high
levels of the enzyme 7-dehydrocholestrol delta 7 reductase which causes immediate conversion of vitamin D3 from sunlight to 7-
dehydrocholesterol.[95] This fat soluble vitamin is required in cats for bone formation through the promotion of calcium retention,
along with nerve and muscle control through absorption ofcalcium and phosphorus.[95]

Cats, like all mammals, need to get linoleic acid, an essential fatty acid, from their diet. Most mammals can convert linoleic acid to
arachidonic acid, as well as the omega 3 fatty acids (eicosapentaenoic acid and docosahexaenoic acid) through the activity of
enzymes, but this process is very limited in cats.[90] The Δ6-desaturase enzyme eventually converts linoleic acid, which is in its salt
form linoleate, to arachidonate (salt form of arachidonic acid) in the liver, but this enzyme has very little activity in cats.[90] This
means that arachidonic acid is an essential fatty acid for cats as they lack the ability to create required amounts of linoleic acid.
Deficiency of arachidonic acid in cats is related to problems in growth, can cause injury and inflammation to skin (e.g. around the
mouth) decreased platelet aggregation, fatty liver, increase in birth defects of kittens whose queens were deficient during pregnancy,
and reproductive failure in queens.[90] Arachidonic acid can also be metabolized to eicosanoids that create inflammatory responses
[96]
which are needed to stimulate proper growth and repair mechanisms in the cat.

The nutrient chart provides a list of the many nutrients cats require as well as the use of the nutrients in the body and thefects
ef of the
deficiency.

Senses
Cats have excellent night vision and can see at only one-sixth the light level required
for human vision.[65]:43 This is partly the result of cat eyes having a tapetum
lucidum, which reflects any light that passes through the retina back into the eye,
thereby increasing the eye's sensitivity to dim light.[97] Another adaptation to dim
light is the large pupils of cats' eyes. Unlike some big cats, such as tigers, domestic
cats have slit pupils.[98] These slit pupils can focus bright light without chromatic
aberration, and are needed since the domestic cat's pupils are much larger, relative to
their eyes, than the pupils of the big cats.[98] At low light levels a cat's pupils will
Reflection of camera flash from the expand to cover most of the exposed surface of its eyes.[99] However, domestic cats
tapetum lucidum have rather poor color vision and (like most nonprimate mammals) have only two
types of cones, optimized for sensitivity to blue and yellowish green; they have
limited ability to distinguish between red and green.[100] A 1993 paper reported a
response to middle wavelengths from a system other than the rods which might be due to a third type of cone. However, this appears
to be an adaptation to low light levels rather than representing truetrichromatic vision.[101]

Cats have excellent hearing and can detect an extremely broad range of frequencies. They can hear higher-pitched sounds than either
dogs or humans, detecting frequencies from 55 Hz to 79,000 Hz, a range of 10.5 octaves, while humans and dogs both have ranges of
about 9 octaves.[102][103] Cats can hear ultrasound, which is important in hunting[104] because many species of rodents make
ultrasonic calls.[105] However, they do not communicate using ultrasound like rodents do. Cats' hearing is also sensitive and among
the best of any mammal,[102] being most acute in the range of 500 Hz to 32 kHz.[106] This sensitivity is further enhanced by the cat's
[104]
large movable outer ears (theirpinnae), which both amplify sounds and help detect the direction of a noise.

Cats have an acute sense of smell, due in part to their well-developed olfactory bulb and a large surface of olfactory mucosa, about
5.8 cm2 (0.90 in2) in area, which is about twice that of humans.[107] Cats are sensitive to pheromones such as 3-mercapto-3-
methylbutan-1-ol,[108] which they use to communicate through urine spraying and marking with scent glands.[109] Many cats also
respond strongly to plants that contain nepetalactone, especially catnip, as they can detect that substance at less than one part per
billion.[110] About 70–80% of cats are affected by nepetalactone.[111] This response is also produced by other plants, such as silver
vine (Actinidia polygama) and the herb valerian; it may be caused by the smell of these plants mimicking a pheromone and
stimulating cats' social or sexual behaviors.[112]

Cats have relatively few taste buds compared to humans (470 or so versus more than 9,000 on the human tongue).[113] Domestic and
wild cats share a gene mutation that keeps their sweet taste buds from binding to sugary molecules, leaving them with no ability to
taste sweetness.[114] Their taste buds instead respond to amino acids, bitter tastes, and acids.
[115] Cats and many other animals have a

Jacobson's organ located in their mouths that allows them to taste-smell certain aromas in a way of which humans have no
experience. Cats also have a distinct temperature preference for their food, preferring food with a temperature around 38 °C (100 °F)
which is similar to that of a fresh kill and routinely rejecting food presented cold or refrigerated (which would signal to the cat that
[113]
the "prey" item is long dead and therefore possibly toxic or decomposing).

To aid with navigation and sensation, cats have dozens of movable whiskers
(vibrissae) over their body, especially their faces. These provide information on the
width of gaps and on the location of objects in the dark, both by touching objects
directly and by sensing air currents; they also trigger protective blink reflexes to
protect the eyes from damage.[65]:47

Most breeds of cat have a noted fondness for settling in high places, or perching. In
the wild, a higher place may serve as a concealed site from which to hunt; domestic
cats may strike prey by pouncing from a perch such as a tree branch, as does a The whiskers of a cat are highly
leopard.[116] Another possible explanation is that height gives the cat a better sensitive to touch.
observation point, allowing it to survey its territory. During a fall from a high place,
a cat can reflexively twist its body and right itself using its acute sense of balance
and flexibility.[117] This is known as the cat righting reflex. An individual cat always rights itself in the same way, provided it has the
time to do so, during a fall. The height required for this to occur is around 90 cm (3.0 ft). Cats without a tail (e.g. Manx cats) also
have this ability, since a cat mostly moves its hind legs and relies on conservation of angular momentum to set up for landing, and the
tail is little used for this feat.[118]

Health
The average lifespan of pet cats has risen in recent years. In the early 1980s, it was about seven years,[119]:33[120] rising to 9.4 years
in 1995[119]:33 and 12–15 years in 2014.[121] However, cats have been reported as surviving into their 30s,[122] with the oldest
known cat, Creme Puff, dying at a verified age of 38.[123]

Spaying or neutering increases life expectancy: one study found neutered male cats live twice as long as intact males, while spayed
female cats live 62% longer than intact females.[119]:35 Having a cat neutered confers health benefits, because castrated males cannot
develop testicular cancer, spayed females cannot develop uterine or ovarian cancer, and both have a reduced risk of mammary
cancer.[124]

Despite widespread concern about the welfare of free-roaming cats, the lifespans of neutered feral cats in managed colonies compare
favorably with those of pet cats.[125]:45[126]:1358 [127][128][129][130]

Diseases
A wide range of health problems may affect cats, including infectious diseases, parasites, injuries, and chronic disease. Vaccinations
are available for many of these diseases, and domestic cats are regularly given treatments to eliminate parasites such as worms and
fleas.[131]

Poisoning
In addition to obvious dangers such as rodenticides, insecticides, and herbicides, cats may be poisoned by many chemicals usually
considered safe by their human guardians,[132] because their livers are less effective at some forms of detoxification than those of
many other animals, including humans and dogs.[31][133] Some of the most common causes of poisoning in cats are antifreeze and
rodent baits.[134] Cats may be particularly sensitive to environmental pollutants.[132][135] When a cat has a sudden or prolonged
serious illness without any obvious cause, it has possibly been exposed to a toxin.

Many human medicines should never be given to cats. For example, the painkiller paracetamol (or acetaminophen, sold as Tylenol
and Panadol) is extremely toxic to cats: even very small doses need immediate treatment and can be fatal.[136][137] Even aspirin,
which is sometimes used to treat arthritis in cats, is much more toxic to them than to humans[137] and must be administered
cautiously.[132] Similarly, application of minoxidil (Rogaine) to the skin of cats, either accidentally or by well-meaning guardians
attempting to counter loss of fur, has sometimes been fatal.[138] Essential oils can be toxic to cats and cases have been reported of
[139]
serious illnesses caused bytea tree oil, including flea treatments and shampoos containing it.

Other common household substances that should be used with caution around cats include mothballs and other naphthalene
products.[132] Phenol-based products (e.g. Pine-Sol, Dettol/Lysol or hexachlorophene)[132] are often used for cleaning and
disinfecting near cats' feeding areas or litter boxes, but these can sometimes be fatal.[140] Ethylene glycol, often used as an
automotive antifreeze, is particularly appealing to cats, and as little as a teaspoonful can be fatal.[141] Some human foods are toxic to
cats; for example chocolate can cause theobromine poisoning, although (unlike dogs) few cats will eat chocolate.[142] Large amounts
of onions or garlic are also poisonous to cats.[132] Many houseplants are also dangerous,[143] such as Philodendron species and the
[144]
leaves of the Easter lily (Lilium longiflorum), which can cause permanent and life-threatening kidney damage.

Genetics
The domesticated cat and its closest wild ancestor are both diploid organisms that possess 38 chromosomes[145] and roughly 20,000
genes.[146] About 250 heritable genetic disorders have been identified in cats, many similar to human inborn errors.[147] The high
level of similarity among the metabolism of mammals allows many of these feline diseases to be diagnosed using genetic tests that
were originally developed for use in humans, as well as the use of cats asanimal models in the study of the human diseases.[148][149]

Behavior
Outdoor cats are active both day and night, although they tend to be slightly more
active at night.[150][151] The timing of cats' activity is quite flexible and varied,
which means house cats may be more active in the morning and evening, as a
response to greater human activity at these times.[152] Although they spend the
majority of their time in the vicinity of their home, housecats can range many
hundreds of meters from this central point, and are known to establish territories that
vary considerably in size, in one study ranging from 7 to 28 hectares (17–69
acres).[151]

Cats conserve energy by sleeping more than most animals, especially as they grow
older. The daily duration of sleep varies, usually between 12 and 16 hours, with 13
and 14 being the average. Some cats can sleep as much as 20 hours. The term "cat
nap" for a short rest refers to the cat's tendency to fall asleep (lightly) for a brief A cat on a fence.
period. While asleep, cats experience short periods of rapid eye movement sleep
often accompanied by muscle twitches, which suggests they aredreaming.[153]

Sociability
Although wildcats are solitary, the social behavior of domestic cats is much more variable and ranges from widely dispersed
individuals to feral cat colonies that form around a food source, based on groups of co-operating females.[154][155] Within such
groups, one cat is usually dominant over the others.[33] Each cat in a colony holds a distinct territory, with sexually active males
 having the largest territories, which are about 10 times larger than those of female
cats and may overlap with several females' territories.[109] These territories are
marked by urine spraying, by rubbing objects at head height with secretions from
facial glands, and by defecation.[109] Between these territories are neutral areas
where cats watch and greet one another without territorial conflicts. Outside these
neutral areas, territory holders usually chase away stranger cats, at first by staring,
hissing, and growling, and if that does not work, by short but noisy and violent
attacks. Despite some cats cohabiting in colonies, they do not have a social survival
Social grooming
strategy, or a pack mentality, and always hunt alone.[156]

However, some pet cats are poorly

socialized. In particular, older cats may show aggressiveness towards newly arrived
kittens, which may include biting and scratching; this type of behavior is known as
feline asocial aggression.[157]

Though cats and dogs are often characterized as natural enemies, they can live
together if correctly socialized.[158]

Life in proximity to humans and other domestic animals has led to a symbiotic social
adaptation in cats, and cats may express great affection toward humans or other
animals. Ethologically, the human keeper of a cat may function as a sort of surrogate
for the cat's mother,[159] and adult housecats live their lives in a kind of extended
kittenhood,[160] a form of behavioral neoteny. The high-pitched sounds housecats
make to solicit food may mimic the cries of a hungry human infant, making them
Cat with an Alaskan Malamute dog
particularly hard for humans to ignore.[161]

Communication
Domestic cats use many vocalizations for communication, including purring, trilling, hissing, growling/snarling, grunting, and
several different forms of meowing.[7] (By contrast, feral cats are generally silent.)[162]:208 Their types of body language, including
position of ears and tail, relaxation of the whole body, and kneading of the paws, are all indicators of mood. The tail and ears are
particularly important social signal mechanisms in cats;[163][164] for example, a raised tail acts as a friendly greeting, and flattened
ears indicates hostility. Tail-raising also indicates the cat's position in the group's social hierarchy, with dominant individuals raising
their tails less often than subordinate animals.[164] Nose-to-nose touching is also a common greeting and may be followed by social
[155]
grooming, which is solicited by one of the cats raising and tilting its head.

Purring may have developed as an evolutionary advantage as a signalling mechanism of reassurance between mother cats and nursing
kittens. Post-nursing cats often purr as a sign of contentment: when being petted, becoming relaxed,[165][166] or eating. The
mechanism by which cats purr is elusive. The cat has no unique anatomical feature that is clearly responsible for the sound.[167] It
was, until recent times, believed that only the cats of the Felis genus could purr. However, felids of the Panthera genus (tiger, lion,
[168]
jaguar, and leopard) also produce sounds similar to purring, but only when exhaling.

Grooming
Cats are known for spending considerable amounts of time licking their coat to keep it clean.[169] The cat's tongue has backwards-
facing spines about 500μm long, which are calledpapillae. These contain keratin which makes them rigid[170] so the papillae act like
a hairbrush. Some cats, particularly longhaired cats, occasionally regurgitate hairballs of fur that have collected in their stomachs
from grooming. These clumps of fur are usually sausage-shaped and about 2–3 cm (0.8–1.2 in) long. Hairballs can be prevented with
f brush.[169]
remedies that ease elimination of the hair through thegut, as well as regular grooming of the coat with a comb or stif
Fighting
Among domestic cats, males are more likely to fight than females.[171] Among feral
cats, the most common reason for cat fighting is competition between two males to
mate with a female. In such cases, most fights are won by the heavier male.[172]
Another common reason for fighting in domestic cats is the difficulty of establishing
territories within a small home.[171] Female cats also fight over territory or to defend
their kittens. Neutering will decrease or eliminate this behavior in many cases,
suggesting that the behavior is linked tosex hormones.[173] The hooked papillae on a cat's
tongue act like a hairbrush to help
When cats become aggressive, they try to make themselves appear larger and more clean and detangle fur.
threatening by raising their fur, arching their backs, turning sideways and hissing or
spitting.[163] Often, the ears are pointed down and back to avoid damage to the inner
ear and potentially listen for any changes behind them while focused forward. They
may also vocalize loudly and bare their teeth in an effort to further intimidate their
opponent. Fights usually consist of grappling and delivering powerful slaps to the
face and body with the forepaws as well as bites. Cats also throw themselves to the
ground in a defensive posture to rake their opponent's belly with their powerful hind
legs.[174]
Play media
Serious damage is rare, as the fights are usually short in duration, with the loser A tabby housecat uses its brush-like
running away with little more than a few scratches to the face and ears. However, tongue to groom itself, licking its fur
fights for mating rights are typically more severe and injuries may include deep to straighten it.
puncture wounds and lacerations. Normally, serious injuries from fighting are
limited to infections of scratches and bites, though these can occasionally kill cats if
untreated. In addition, bites are probably the main route of transmission of feline
immunodeficiency virus.[175] Sexually active males are usually involved in many
fights during their lives, and often have decidedly battered faces with obvious scars
and cuts to their ears and nose.[176]

Hunting and feeding

A cat that is playing with a caught
mouse. Cats play with their prey to
weaken or exhaust them before
making a kill.
Cats hunt small prey, primarily
birds and rodents,[177] and are An arched back, raised fur, and an
often used as a form of pest open-mouthed hiss can all be signs
control.[178][179] Domestic cats are of aggression in a domestic cat.
a major predator of wildlife in the
United States, killing an estimated
1.4–3.7 billion birds and 6.9–20.7 billion mammals annually.[180][181] The bulk of
predation in the United States is done by 80 million feral and stray cats. Effective
measures to reduce this population are elusive, meeting opposition from cat
enthusiasts.[180][181] In the case of free-ranging pets, equipping cats with bells and
[177]
not letting them out at night will reduce wildlife predation.

Free-fed feral cats and house cats tend to consume many small meals in a single day,
although the frequency and size of meals varies between individuals.[156] Cats use two hunting strategies, either stalking prey
actively, or waiting in ambush until an animal comes close enough to be captured.[182] Although it is not certain, the strategy used
[183]:153
may depend on the prey species in the area, with cats waiting in ambush outside burrows, but tending to actively stalk birds.
Perhaps the best known element of cats' hunting behavior, which is commonly misunderstood and often appalls cat owners because it
looks like torture, is that cats often appear to "play" with prey by releasing it after capture. This behavior is due to an instinctive
imperative to ensure that the prey is weak enough to be killed without endangering the cat.[184] This behavior is referred to in the
idiom "cat-and-mouse game" or simply "cat and mouse".

Another poorly understood element of cat hunting behavior is the presentation of prey to human guardians. Ethologist Paul
Leyhausen proposed that cats adopt humans into their social group and share excess kill with others in the group according to the
dominance hierarchy, in which humans are reacted to as if they are at, or near, the top.[185] Anthropologist and zoologist Desmond
Morris, in his 1986 book Catwatching, suggests, when cats bring home mice or birds, they are attempting to teach their human to
hunt, or trying to help their human as if feeding "an elderly cat, or an inept kitten".[186][187] Morris's hypothesis is inconsistent with
[183]:153
the fact that male cats also bring home prey,despite males having negligible involvement with raising kittens.

Domestic cats select food based on its temperature, smell and texture; they dislike chilled foods and respond most strongly to moist
foods rich in amino acids, which are similar to meat.[84][156] Cats may reject novel flavors (a response termed neophobia) and learn
quickly to avoid foods that have tasted unpleasant in the past.[156] They may also avoid sugary foods and milk. Most adult cats are
lactose intolerant; the sugars in milk are not easily digested and may cause soft stools or diarrhea.[156][188] They can also develop
odd eating habits. Some cats like to eat or chew on other things, most commonly wool, but also plastic, cables, paper, string,
aluminum foil, or even coal. This condition, pica, can threaten their health, depending on the amount and toxicity of the items
eaten.[189][190]

Though cats usually prey on animals less than half their size, a feral cat in Australia has been photographed killing an adult
pademelon of around the cat's weight at 4 kg (8.8 lb).[191]

Since cats lack lips[192] to create suction, they use a lapping method with the tongue to draw liquid upwards into their mouths.
Lapping at a rate of four times a second, the cat touches the smooth tip of its tongue to the surface of the water
, and quickly retracts it,
drawing water upwards.[193]

Play
Domestic cats, especially young kittens, are known for their love of play. This
behavior mimics hunting and is important in helping kittens learn to stalk, capture,
and kill prey.[194] Cats also engage in play fighting, with each other and with
humans. This behavior may be a way for cats to practice the skills needed for real
combat, and might also reduce any fear they associate with launching attacks on
other animals.[195]
Play media
Owing to the close similarity between play and hunting, cats prefer to play with
Play fight between kittens, age 14
objects that resemble prey, such as small furry toys that move rapidly, but rapidly weeks
lose interest (they become habituated) in a toy they have played with before.[196]
Cats also tend to play with toys more when they are hungry.[197] String is often used
as a toy, but if it is eaten, it can become caught at the base of the cat's tongue and then move into the intestines, a medical emergency
which can cause serious illness, even death.[198] Owing to the risks posed by cats eating string, it is sometimes replaced with a laser
pointer's dot, which cats may chase.[199]

Reproduction
Female cats are seasonally polyestrous, which means they may have many periods of heat over the course of a year, the season
beginning in spring and ending in late autumn. Heat periods occur about every two weeks and last about 4 to 7 days.[200] Multiple
males will be attracted to a female in heat. The males will fight over her, and the victor wins the right to mate. At first, the female
rejects the male, but eventually the female allows the male to mate. The female utters a loud yowl as the male pulls out of her
because a male cat's penis has a band of about 120–150 backwards-pointing penile spines, which are about 1 mm long; upon
withdrawal of the penis, the spines rake the walls of the female's vagina, which acts
to induce ovulation. This act also occurs to clear the vagina of other sperm in the
context of a second (or more) mating, thus giving the later males a larger chance of
conception.[201]

After mating, the female washes her vulva thoroughly. If a male attempts to mate
with her at this point, the female will attack him. After about 20 to 30 minutes, once
[200]
the female is finished grooming, the cycle will repeat.

Because ovulation is not always triggered by a single mating, females may not be When cats mate, the tomcat (male)
impregnated by the first male with which they mate.[202] Furthermore, cats are bites the scruff of the female's neck
superfecund; that is, a female may mate with more than one male when she is in as she assumes a position conducive
heat, with the result that different kittens in a litter may have different fathers.[200] to mating known as lordosis
behavior.
At 124 hours after conception, the morula forms. At 148 hours, early blastocysts
form. At 10–12 days, implantation
occurs.[203][204]

The gestation period for cats is

between 64 and 67 days, with an
average of 66 days.[205] The size of
a litter usually is three to five
kittens, with the first litter usually
smaller than subsequent litters.
A newborn kitten Kittens are weaned between six and
seven weeks old, and cats normally
reach sexual maturity at 5–10
months (females) and to 5–7 months (males), although this can vary depending on
breed.[200] Females can have two to three litters per year, so may produce up to 150
kittens in their breeding span of around ten years.[200]

Cats are ready to go to new homes at about 12 weeks of age,[206] when they are
ready to leave their mother. They can be surgically sterilized (spayed or castrated) as
Radiography of a pregnant cat (about
early as 7 weeks to limit unwanted reproduction.[207] This surgery also prevents one month and a half)
undesirable sex-related behavior, such as aggression, territory marking (spraying
urine) in males and yowling (calling) in females. Traditionally, this surgery was
performed at around six to nine months of age, but it is increasingly being performed prior to puberty, at about three to six
months.[208] In the US, about 80% of household cats are neutered.
[209]

Ecology

Habitats
Cats are a cosmopolitan species and are found across much of the world.[38] Geneticist Stephen James O'Brien, of the National
Cancer Institute in Frederick, Maryland, remarked on how successful cats have been in evolutionary terms: "Cats are one of
evolution's most charismatic creatures. They can live on the highest mountains and in the hottest deserts."[210] They are extremely
adaptable and are now present on all continents except Antarctica, and on 118 of the 131 main groups of islands—even on isolated
islands such as the Kerguelen Islands.[211][212]
Feral cats can live in forests, grasslands, tundra, coastal areas, agricultural land,
scrublands, urban areas, and wetlands.[213] Their habitats even include small
oceanic islands with no human inhabitants.[214] Further, the close relatives of
domestic cats, the African wildcat (Felis silvestris lybica) and the Arabian sand cat
(Felis margarita) both inhabit desert environments,[28] and domestic cats still show
similar adaptations and behaviors.[31] The cat's ability to thrive in almost any
terrestrial habitat has led to its designation as one of the world's worst invasive
species.[215]
A cat in snowy weather
As domestic cats are little altered from wildcats, they can readily interbreed. This
hybridization poses a danger to the genetic distinctiveness of some wildcat
populations, particularly inScotland and Hungary and possibly also theIberian Peninsula.[41]

Feral cats
Feral cats are domestic cats that were born in or have reverted to a wild state. They are
unfamiliar with and wary of humans and roam freely in urban and rural areas.[9] The numbers
of feral cats is not known, but estimates of the US feral population range from 25 to
60 million.[9] Feral cats may live alone, but most are found in large colonies, which occupy a
specific territory and are usually associated with a source of food.[216] Famous feral cat
colonies are found in Rome around theColosseum and Forum Romanum, with cats at some of
[217]
these sites being fed and given medical attention by volunteers.

Public attitudes towards feral cats vary widely, ranging from seeing them as free-ranging pets,
to regarding them as vermin.[218] One common approach to reducing the feral cat population
is termed 'trap-neuter-return', where the cats are trapped, neutered, immunized against diseases Feral farm cat
such as rabies and the feline Panleukopenia and Leukemia viruses, and then released.[219]
Before releasing them back into their feral colonies, the attending veterinarian often nips the
tip off one ear to mark it as neutered and inoculated, since these cats may be trapped again. Volunteers continue to feed and give care
to these cats throughout their lives. Given this support, their lifespans are increased, and behavior and nuisance problems caused by
competition for food are reduced.[216]

Impact on prey species

To date, little scientific data is available to assess the impact of cat predation on prey
populations outside of agricultural situations. Even well-fed domestic cats may hunt
and kill, mainly catching small mammals, but also birds, amphibians, reptiles, fish,
and invertebrates.[177][220] Hunting by domestic cats may be contributing to the
decline in the numbers of birds in urban areas, although the importance of this effect
remains controversial.[221] In the wild, the introduction of feral cats during human
settlement can threaten native species with extinction.[214] In many cases,
controlling or eliminating the populations of non-native cats can produce a rapid
recovery in native animals.[222] However, the ecological role of introduced cats can
be more complicated. For example, cats can control the numbers of rats, which also
Carrying half of a rabbit
prey on birds' eggs and young, so a cat population can protect an endangered bird
species by suppressingmesopredators.[223]

In isolated landmasses, such as Australasia, there are often no other native, medium-sized quadrupedal predators (including other
feline species); this tends to exacerbate the impact of feral cats on small native animals.[224] Native species such as the New Zealand
kakapo and the Australian bettong, for example, tend to be more ecologically vulnerable and behaviorally "naive", when faced with
predation by cats.[225] Feral cats have had a major impact on these native species and have played a leading role in the endangerment
and extinction of many animals.[226]

Even in places with ancient and numerous cat populations, such as Western Europe, cats appear to be growing in number and
independently of their environments' carrying capacity (such as the numbers of prey available).[227][228] This may be explained, at
least in part, by an abundance of food, from sources including feeding by pet owners and scavenging. For instance, research in
Britain suggests that a high proportion of cats hunt only "recreationally".[228] And in South Sweden, where research in 1982 found
[227]
that the population density of cats was as high as 2,000 per square kilometre (5,200/sq mi).

In agricultural settings, cats can be effective at keeping mouse and rat populations low, but only if rodent harborage locations are kept
under control.[229][230] While cats are effective at preventing rodent population explosions, they are not effective for eliminating pre-
exisiting severe infestations.[231]

Impact on birds
The domestic cat is a significant predator of birds. UK assessments indicate they
may be accountable for an estimated 64.8 million bird deaths each year.[177] A 2012
study suggests feral cats may kill several billion birds each year in the United
States.[232] Certain species appear more susceptible than others; for example, 30%
of house sparrow mortality is linked to the domestic cat.[233] In the recovery of
ringed robins (Erithacus rubecula) and dunnocks (Prunella modularis), 31% of
deaths were a result of cat predation.[234] In parts of North America, the presence of
larger carnivores such as coyotes which prey on cats and other small predators
A black cat eating a house sparrow
reduces the effect of predation by cats and other small predators such as opossums
and raccoons on bird numbers and variety.[235] The proposal that cat populations
will increase when the numbers of these top predators decline is called the mesopredator release hypothesis.

On islands, birds can contribute as much as 60% of a cat's diet.[236] In nearly all cases, however, the cat cannot be identified as the
sole cause for reducing the numbers of island birds, and in some instances, eradication of cats has caused a 'mesopredator release'
effect;[237] where the suppression of top carnivores creates an abundance of smaller predators that cause a severe decline in their
shared prey. Domestic cats are, however, known to be a contributing factor to the decline of many species, a factor that has ultimately
led, in some cases, to extinction. The South Island piopio, Chatham rail,[234] the New Zealand merganser,[238] and the common
diving petrel[239] are a few from a long list, with the most extreme case being the flightless Lyall's wren, which was driven to
.[240][241]
extinction only a few years after its discovery

Some of the same factors that have promoted adaptive radiation of island avifauna over evolutionary time appear to promote
vulnerability to non-native species in modern time. The susceptibility of many island birds is undoubtedly due to evolution in the
absence of mainland predators, competitors, diseases, and parasites, in addition to lower reproductive rates and extended incubation
periods.[242] The loss of flight, or reduced flying ability is also characteristic of many island endemics.[243] These biological aspects
have increased vulnerability to extinction in the presence of introduced species, such as the domestic cat.[244] Equally, behavioral
traits exhibited by island species, such as "predatory naivety"[245] and ground-nesting,[242] have also contributed to their
susceptibility.

Interaction with humans

Cats are common pets throughout the world, and their worldwide population exceeds 500 million.[12] Although cat guardianship has
commonly been associated with women,[246] a 2007 Gallup poll reported that men and women in the United States of America were
equally likely to own a cat.[247]
As well as being kept as pets, cats are also used in the international fur[248] and
leather industries for making coats, hats, blankets and stuffed toys;[249] and shoes,
gloves and musical instruments respectively[250] (about 24 cats are needed to make
a cat fur coat).[251] This use has now been outlawed in the United States, Australia,
and the European Union.[252] Cat pelts have been used for superstitious purposes as
part of the practise of witchcraft,[253] and are still made into blankets in Switzerland
as folk remedies believed to help rheumatism.[254] In the Western intellectual
tradition, the idea of cats as everyday objects have served to illustrate problems of
Cats and people
quantum mechanics in the Schrödinger's cat thought experiment.

A few attempts to build a cat census have been made over the years, both through
associations or national and international organizations (such as the Canadian Federation of Humane Societies's one[255] ) and over
the net,[256][257] but such a task does not seem simple to achieve. General estimates for the global population of domestic cats range
[258][259][260][261][262][263]
widely from anywhere between 200 million to 600 million.

Cat show
A cat show is a judged event where the owners of cats compete to win titles in various cat registering organizations by entering their
cats to be judged after a breed standard. Bothpedigreed and companion (or moggy) cats are admissible, although the rules dif
fer from
organization to organization. Cats are compared to a breed standard, and the owners of those judged to be closest to it are awarded a
prize. Moggy's are judged based on their temperament. Often, at the end of the year, all of the points accrued at various shows are
added up and more national and regional titles are awarded.

Cat café
A cat café is a theme café whose attraction is cats that can be watched and played with. Patrons pay a cover fee, generally hourly and
thus cat cafés can be seen as a form of supervised indoorpet rental.

Ailurophobia
[264]
Ailurophobia is a human phobia of cats; however, the term is often associated with humans that have a hatred of cats.

Cat bites
Cats may bite humans when provoked, during play or when aggressive. Complications from cat bites can develop.[265] A cat bite
differs from the bites of other pets. This is because the teeth of a cat are sharp and pointed causing deep punctures. Skin usually
closes rapidly over the bite and trapsmicroorganisms that cause infection.[266][265]

Infections transmitted from cats to humans

Cats can be infected or infested with viruses, bacteria, fungus, protozoans, arthropods or worms that can transmit diseases to
humans.[267] In some cases, the cat exhibits no symptoms of the disease,[268] However, the same disease can then become evident in
a human. The likelihood that a person will become diseased depends on the age and immune status of the person. Humans who have
cats living in their home or in close association are more likely to become infected, however, those who do not keep cats as pets
might also acquire infections from cat feces and parasites exiting the cat's body.[267][269] Some of the infections of most concern
include salmonella, cat scratch disease and toxoplasmosis.[268]

History and mythology

Traditionally, historians tended to think ancient Egypt was the site of cat
domestication, owing to the clear depictions of house cats in Egyptian paintings
about 3,600 years old.[28] However, in 2004, a Neolithic grave excavated in
Shillourokambos, Cyprus, contained the skeletons, laid close to one another, of both
a human and a cat. The grave is estimated to be 9,500 years old, pushing back the
earliest known feline–human association significantly.[15][271][272] The cat
specimen is large and closely resembles the African wildcat, rather than present-day
domestic cats. This discovery, combined with genetic studies, suggests cats were The ancient Egyptians mummified
probably domesticated in the Middle East, in the Fertile Crescent around the time of dead cats out of respect in the same
the development of agriculture, and then were brought to Cyprus and Egypt.[14][19] way that they mummified people.[270]

Direct evidence for the domestication of cats 5,300 years ago in Quanhucun, China
has been published by archaeologists and paleontologists from the University of
Washington and Chinese Academy of Sciences. The cats are believed to have been
attracted to the village by rodents, which in turn were attracted by grain cultivated
and stored by humans.[273]

In ancient Egypt, cats weresacred animals, with the goddess Bastet often depicted in
cat form, sometimes taking on the war-like aspect of a lioness.[274]:220 Killing a cat
was absolutely forbidden[270] and the Greek historian Herodotus reports that,
whenever a household cat died, the entire family would mourn and shave their
eyebrows.[270] Families took their dead cats to the sacred city of Bubastis,[270]
[270]
where they were embalmed and buried in sacred repositories.
Ancient Roman mosaic of a cat
Domestic cats were probably first introduced to Greece and southern Italy in the killing a partridge from the House of
fifth century BC by the Phoenicians.[275] The earliest unmistakable evidence of the the Faun in Pompeii
Greeks having domestic cats comes from two coins from Magna Graecia dating to
the mid-fifth century BC showing Iokastos and Phalanthos, the legendary founders
of Rhegion and Taras respectively, playing with their pet cats.[276]:57-58[277]

Housecats seem to have been extremely rare among the ancient Greeks and
Romans;[277] Herodotus expressed astonishment at the domestic cats in Egypt,
because he had only ever seen wildcats.[277] Even during later times, weasels were
far more commonly kept as pets[277] and weasels, not cats, were seen as the ideal
rodent-killers.[277] The usual ancient Greek word for "cat" was ailouros, meaning
"thing with the waving tail",[276]:57[277] but this word could also be applied to any A 19th century drawing of a tabby cat

of the "various long-tailed carnivores kept for catching mice".[277] Cats are rarely
mentioned in ancient Greek literature,[277] but Aristotle does remark in his History
of Animals that "female cats are naturallylecherous."[276]:74[277]

The Greeks later syncretized their own goddess Artemis with the Egyptian goddess Bastet, adopting Bastet's associations with cats
and ascribing them to Artemis.[276]:77-79 In Ovid's Metamorphoses, when the gods flee to Egypt and take animal forms, the goddess
Diana (the Roman equivalent of Artemis) turns into a cat.[276]:79 Cats eventually displaced ferrets as the pest control of choice
[278]
because they were more pleasant to have around the house and were more enthusiastic hunters of mice.

During the Middle Ages, many of Artemis's associations with cats were grafted onto the Virgin Mary.[278] Cats are often shown in
icons of Annunciation and of the Holy Family[278] and, according to Italian folklore, on the same night that Mary gave birth to Jesus,
a virgin cat in Bethlehem gave birth to a kitten.[278] Domestic cats were spread throughout much of the rest of the world during the
[274]:223
Age of Discovery, as ships' cats were carried on sailing ships to control shipboard rodents and as good-luck charms.

Several ancient religions believed cats are exalted souls, companions or guides for humans, that are all-knowing but mute so they
cannot influence decisions made by humans. In Japan, themaneki neko cat is a symbol of good fortune.[279]
Although no species are sacred in Islam, cats are revered by Muslims. Some Western writers have stated Muhammad had a favorite
cat, Muezza.[280] He is reported to have loved cats so much, "he would do without his cloak rather than disturb one that was sleeping
on it".[281] The story has no origin in early Muslim writers, and seems to confuse a story of a later Sufi saint, Ahmed ar-Rifa'i,
centuries after Muhammad.[282]

Freyja, the goddess of love, beauty, and fertility in Norse mythology, is depicted as riding a chariot drawn by cats.[283]

Superstitions and cat burning

Many cultures have negative superstitions about cats. An example would be the belief that a black cat "crossing one's path" leads to
bad luck, or that cats are witches' familiars used to augment a witch's powers and skills. The killing of cats in Medieval Ypres,
Belgium, is commemorated in the innocuous present-dayKattenstoet (cat parade).[284]

In medieval France, cats would be burnt alive as a form of entertainment. According to Norman Davies, the assembled people
carbonized".[285]
"shrieked with laughter as the animals, howling with pain, were singed, roasted, and finally

"It was the custom to burn a basket, barrel, or sack full of live cats, which was hung from a tall mast in the midst of the bonfire;
sometimes a fox was burned. The people collected the embers and ashes of the fire and took them home, believing that they brought
good luck. The French kings often witnessed these spectacles and even lit the bonfire with their own hands. In 1648 Louis XIV,
crowned with a wreath of roses and carrying a bunch of roses in his hand, kindled the fire, danced at it and partook of the banquet
afterwards in the town hall. But this was the last occasion when a monarch presided at the midsummer bonfire in Paris. At Metz
midsummer fires were lighted with great pomp on the esplanade, and a dozen cats, enclosed in wicker cages, were burned alive in
them, to the amusement of the people. Similarly at Gap, in the department of the Hautes-Alpes, cats used to be roasted over the
midsummer bonfire."[286]

According to a myth in many cultures, cats have multiple lives. In many countries, they are believed to have nine lives, but in Italy,
Germany, Greece, Brazil and some Spanish-speaking regions, they are said to have seven lives,[287][288] while in Turkish and Arabic
traditions, the number of lives is six.[289] The myth is attributed to the natural suppleness and swiftness cats exhibit to escape life-
threatening situations. Also lending credence to this myth is the fact that falling cats often land on their feet, using an instinctive
[290]
righting reflex to twist their bodies around. Nonetheless, cats can still be injured or killed by a high fall.

See also
Animal testing on cats List of cat breeds
Animal track List of cats
Cat burning List of cat documentaries
Cancer in cats List of fictional cats and felines
Cat and mouse (cat-and-mouse game) Pet door including cat flap
Cat lady Pet first aid
Cats and the Internet Popular cat names
Cat years Trap-neuter-return
Dried cat

Cats by location

Cats in ancient Egypt

Cats in Australia
Cats in New Zealand
Cats in the United States

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External links
The dictionary definition ofcat at Wiktionary
Data related to Cat at Wikispecies
Media related to Cat at Wikimedia Commons
Animal Care at Wikibooks
Quotations related to Cat at Wikiquote
"Cat, Domestic, The". Encyclopedia Americana. 1920.
High-Resolution Images of the Cat Brain
Biodiversity Heritage Library bibliographyfor Felis catus
Catpert. The Cat Expert– Cat articles
View the cat genome in Ensembl

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