BIO 1 FINALS REVIEWER PhyloCode Proposed by systematists; recogs only grps
CONCEPT OF SPECIES w/ common ancestor & all its descendants
Polytomy Branch from w/c more than 2 grps emerge Define species Comes from L word for “kind” or Homology vs analogy Homology = similarity in structure due to “appearance”; is a group of organisms shared ancestry whose mems can breed and prod fertile Analogy = similarity due to convergent evol offspring but who don’t prod fertile Convergent evol Occurs when similar envi pressures & nat’l offspring with mems from other grps selection prod similar adaptations in What is speciation? Emergence of new sp that increases organisms from diff evol lineages diversity of life Homoplasies Analogous strucs/mol seqs that evolved 2 events involved in Linear events = a sp changes over time independently evolutionary change Branching events = prod new sp & diversity Enumerate the ways to 1. Biological sp concept What is phylogeny? Is the evolutionary history of a sp or a grp define a species - Defines sp as pop’n/grps of pop’ns w/ of related sp mems w/ potential to interbreed & What is systematics? The discipline that classifies organisms and prod fertile offspring determines their evol rels.; uses fossils, - Involves gene flow bet pop’ns which molecular, & genetic data to infer evol rels holds them together, and is prevented Taxonomy Ordered div and naming of organisms by reproductive isolation w/c What is binomial Created by Carolus Linnaeus in the 18th cen, maintains separate species nomenclature? this is a sys of taxonomy based on - DOES NOT WORK IN ALL SITUATIONS resemblances. 2. Morphological sp concept It has 2 key feats: - Classifies organisms based on 1. 2-part names for species observable phenotypic traits - Can be applied to asexual organisms & - The 2-part scientific name if a sp is fossils when we don’t know about called binomial interbreeding - The 1st part is the genus, the 2nd partis - Involves subjectivity but used by most the specific epithet (w/c is unique for field biologists each sp w/in genus) 3. Ecological sp concept 2. Hierarchical classification - Defines a sp thru its eco role/niche - Focuses on unique adaptations - Intro’ed by Linnaeus, sys for grping sp in increasingly broad categories 4. Phylogenetic sp concept - Broad to narrow: domain, kingdom, - Defines sp as a set of organisms phylum, class, order, family, genus, sp. rep’ing a specific evol lineage In hier. class., what is a A taxonomic unit @ any level of hierarchy - used on organisms w/ common taxon? ancestor & can be distinguished from Describe the change Early taxonomists class’ed all sp as either others who don’t have same ancestor from 2 kingdoms to 5 plants/animals (the 1st 2 kingdoms). What are reproductive Repro barriers isolate a sp’s gene pool & kingdoms to 3 domains barriers & what are the prevent interbreeding Later, 5 k-doms were intro’ed: Plantae, 2 types? Animalia, Protista, Fungi, Monera 2 types: prezygotic (formed before (prokaryotes). mating/fertilization, prevent mating) & postzygotic (operate after hybrid zygotes More recently, the 3-domain sys was are formed) adopted: Bacteria, Archaea, Eukarya. Enumerate the pre- & PREZYGOTIC Describe the tree of life Based on rRna genes, the tree of life postzygotic barriers 1. Habitat isolation = 2 sp encounter evolved slowly & suggests that eukaryotes each other rarely or not at all b/c they & archaea are more closely related to each live in diff habitats even though not other than to bacteria. separated by phys barriers What is horizontal gene Mvmt of genes from 1 genome to another; 2. Temporal isolation = sp that breed @ transfer? part of the substantial interchanges of diff times of day/seasons/yrs can’t mix genes bet. Org’isms in diff domains & thus gametes complicates efforts to build a tree of life 3. Behavioral isolation = courtship rituals What are phylogenetic Branching trees by which systematists & other behavs unique to a sp are trees? depict hypothesis about evolutionary effective barriers relationships; consist of a branch point 4. Mechanical isolation = morphological (reps the divergence of 2 sp), taxa, sister diffs prevent successful mating taxa (grps w/ immediate common 5. Gametic isolation = sperm of one sp ancestor), and rooted tree (includes a cant fertilize other sp’s eggs branch to rep the last common ancestor of POSTZYGOTIC all taxa in the tree) 1. Reduced hybrid viability = genes of the diff parent sp may interact & impair hybrid’s dev’t; MOST HYBRID Ex. Mammals after dino extinction OFFSPRING DON’T SURVIVE MECHANISMS OF MICROEVOLUTION 2. Reduced hybrid fertility = hybrids may Microevolution Change in the relative freqs of alleles in a be vigorous but may be sterile gene pool over time 3. Hybrid breakdown = 1st -gen sp are Describe the 3 main 1. Nat’l selection fertile but when they breed w/ causes of evol - Alters allele freqs when inds differ in another, next-gen offspring are sterile change/allele freq their survival & repro success 2 types of speciation ALLOPATRIC = “other country”; pop’ns of alterations in a pop’n - Ex. Non-/webbed boobies same sp are geog’ly separated, so gene flow is reduced/separated/prevented 2. Genetic drift initially by geog barriers like: - Change in the gene pool of a pop’n 1. Mt. formation due to chance events w/c may lead to 2. Deep canyons loss of genetic diversity 3. Removal of land bridges bet - Leads to bottleneck effect (loss of gen continents div when a pop’n is greatly reduced) 4. Conti drift founder effect (when a few inds colonize a new habitat; the smaller the SYMPATRIC = “same country”; speciation grp, the more diif the gene pool of the happens in geog’ly overlapping pop’ns; new pop’n will be from the gene pool gene flow is reduced by: of the orgi pop’n) 1. Polyploidy = presence of extra chromosome set b/c of accidents 3. Gene flow during cell division - mvmt of inds/gametes.spores bet pop’ns - Autopolyploid = ind w/ more than 2 What is an ind’s fitness Contrib it makes to next gen’s gene pool chrom sets derived from 1 sp. Disclaimers abt nat sel Nat sel can’t fashion perfect organisms - Tetraploid plants = arise from a diploid parent & can form by self-ferti of 2n 1. Sel can only act on existing variations gametes where meiosis hasn’t 2. Evol is limited by historical constraints reduced the chrom set 3. Adaptations are often compromises 2. Habitat diff’tion 4. Chance, nat sel, & envi interact 3. Sexual selection 5 Evidences of 1. Fossils How can broad patterns 1. Using the fossil rec’d evolution - Provs strong evidence for evol in speciation be 2. Morphological data - Biased in favor of sp that (1) existed studied? 3. Molecular data for a long time, (2) were abundant & 2 patterns of speciation ANAGENESIS = accu of changes assoc’d w/ widespread, (3) had hard parts transformation of 1 sp to another - Taphonomy = sci study of fossil CLADOGENESIS = branching evol; budding formation of 1 or more new sp from a parent sp; - Paleontology = study of fossils, dev’d PROMOTES BIODIV BY INC’ING NO. OF SP. by Fr sci Georges Cuvier, who advoc’d 2 models of speciation GRADUALIST MODEL = isolated pop’ns catastrophism (that each boundary change slowly as their allel freqs shift bet strat reps a catastrophe) during adaptation by nat sel PUNCTUATED EQUILIBRIUM MODEL = by 2. Biogeography Niles Eldredge & Stephen Jay Gould; pd of - Geog’ic distribution of sp, suggested rapid evol change (punc) followed by long to Darwin that organisms evolve from pds of no change (equi/stasis) common ancestors What is genomics? Allows researchers to iden specific genes - James Hutton & Charles Lyell = that involved in some cases of speciation changes in earth’s surface result from MECHANISMS OF MACROEVOLUTION slow continuous actions still op’ing Macroevolution Cumulative effect of many speciation & today extinction events - Uniformitarianism = by Lyell, says that What played a major CONTINENTAL DRIFT = slow continuous mechs of change are constant over role in macroevol? mvmt of earth’s crustal plates on the hot time; strongly influenced Darwin’s mantle thinking Mass extinction When extinction rates increase dramatically 3. Comparative anatomy Ordovician - Comparison of body strucs of diff sps. Devonian Permian – caused by volcanism 4. Comparative embryology Triassic - Comparison of early stages of dev’t K-T – most dinos died among diff organisms Adaptive radiatons Caused by mass extinctions; evol of diversely adapted sp from a common 5. Molecular bio ancestor upon intro to new envi opp’ities - Reveal evol rels thru comparisons of DNA & amino acid seqs What is radiometric Determines the absolute age of fossils; a ancestry; may give rise to ANALOGOUS dating parent isotope decays to a daughter traits (when grps independently adapt to isotope @ a constant rate, where each ist similar envis in similar ways) has a known half-life, the time req’d for half HUMAN EVOLUTION the parent iso to decay Describe primates Order of mammals w/ flexible hands & feet, Define pop’ns Grp of inds of same sp living in same place forward-looking eyes, enlarged brains @ same time; OR localized grp of inds that can interbreed & prod fertile offspring Lack strong specialization; related to Gene pool Total coll. Of genes in a pop’n @ any 1 time arboreal/tree-dwelling ancestry Darwin’s DWM & NS That sp showed evidence of DWM from common ancestors, and NS is the mech 3 main grps: behind DWM 1. Lemur, loris, potto Artificial selection From Darwin; that humans have modi’d sps 2. Tarsier by selecting & breeding inds w/ desired 3. Anthropoids (monkeys and apes) traits Primate classification 2 Suborders Darwin’s 4 observations Obs 1: Mems of a pop’n often vary greatly 1. Prosimians (lower primates, “before & 2 inferences in their traits ape”) Obs 2: Traits are inherited from parents to - Oldest primate lineage, all the way to offspring Paleocene Obs 3: All sps are capable of prod’ing more - Lemurs, lorises, tarsiers, tree shrews offspring than the envi can handle - Generally small (size of a mouse to a Obs 4: Due to lack of food/other resources, house cat) many of these offspring don’t survive - Arboreal, w/ 5 digits - Omnivorous Inf 1: Inds w/ inherited traits that give them - Large, forward-looking eyes a higher prob’ity of survival and repro, tend specialized for night vision to leave more offspring than others - Declined due to colder weather; Inf 2: This uneq ability of inds to survive & presently only in tropical regions repro lead to the accu of fav traits in the - pop’n over gens. 2. Anthropoids (higher primates) Scala naturae By gk philosopher Aristotle; arrangement of - Monkeys, apes, humans sp where sps are fixed - Evolved from a prosimian lineage Mutation = changes in the nucleotide seq of DNA; - Divided into 3 superfamilies ULTIMATE SOURCE OF NEW ALLELES; 2 superfamilies of 1. Cercopithecoidea (Old World Monkey) beneficial when envi is changing Anthropoids - Close-set, downward nostrils + Chromosomal Impt source of genetic variation; if a gene is grasping hands + nonprehensile tail duplication duplicated, new copy can undergo - In tropical regions of Africa & Asia mutation w/o affecting orig copy’s fcn Hardy-Weinberg eqn Used to test if a pop’n is evolving; 2. Ceboidea (New World Monkey) Hardy-Weinberg That allele & genotype freqs w/in a sexually - In Central & S. America principle repro’ing 2n pop’n will remain in equi - From African monkeys unless outside forces act to change those - Prehensile tail + widely separated freqs nostrils Hardy-Weinberg Allele & geno freqs won’t change unless - Howler, spider, squirrel monkeys equilibrium something acts to change the gene pool; must satisfy 5 conditions (w/c are rarely 3. Hominoidea (Great Ape) ever met): - 3 families: 1. Great Apes (chimps, orangutan, 1. Very large pop’n gorilla) 2. No gene flow bet pop’ns 2. Lesser Apes (gibbon, siamang) 3. No mutations 3. Hominids (humans, extinct 4. Random mating ancestors) 5. No nat sel Hominid Genus Australopithecus DWM vs NS DWM = explains life’s unity & diversity; Not the same as modern apes (w/c are not summarized Darwin’s perception of the bipedal, no large brain, don’t make tools) unity of life; that all organisms are related Paleoanthropology Study of human origins thru descent from an ancestor from the Hominins From Africa abt 6-7 mya; had small brain remote past but walked upright; formerly hominids, NS = cause of adaptive evolution more closely related to humans than Lamarck’s Theory of Sps evolve thru USE & DISUSE OF BODY chimps Evolution PARTS and INHERITANCE OF ACQUIRED Homo sapiens Included in the primate family Hominidae; CHARS (both unsupported by evidence) Large brain + bipedal locomotion; 200k y.o. Convergent evolution Independent evol of similar feats in diff Derived chars of - Upright posture, bipedalism evol branches; does NOT give info abt humans (vs apes) - Larger brains - Language, symbolic thought workers - Use of complex tools - Energy consumption increased - Shortened jaw - Medical advances reduced deaths - Shorter digestive tract - Age of spaceship & computers Why did bipedalism 1. Carrying behavior 4. Info Tech become the primary 2. Reduced overall heat stress = thru - Millennials adaptation of convection; only humans have sweat - Globalization hominids? glands that prod moisture to cool body - Soc med 3. Most energy-efficient way to travel - Multitasking long distances - Reduced personal interaction 4. Better vision in open envis + defensive ECOLOGY action by freeing hands What is ecology? - Comes from Gk word Cradle of mankind Olduvai George, E. Serengeti Plain, N. “oikos”=”household” & “logos”=”study Tanzania b/c of many hominid discoveries of” Laetoli, Tanzania Discovered by Mary Leakey; preserved - Dynamic interplay/complex hominid footprints in volcanic ash, possibly interactions between living & Australopithecus afarensis nonliving things Homo habilis Handy man Organismal ecology Study of how an organism’s struc, physio, Homo ergaster 1st fully bipedal, large-brained hominid and behavior meet envi challenges Homo erectus 1st hominin to leave Africa Limiting factors of Disease, food, predators, climate, space, Homo floresiensis Hobbit pop’ns mates Homo sapiens 1st to show evidence of symbolic, Carrying capacity Greatest no of inds a pop’n can sustain sophisticated thought Community Grp of pop’ns of diff sps that inhabit a 2 theories about origin 1. OUT-OF-AFRICA (replacement particular area of modern humans hypothesis) Ecosystem Community of organisms in an area & the - They arose from Africa, and H. sapiens phys envi they interact w/ replaced other human pop’ns All organisms in a comm’ity & abiotic envi - More reliable b/c fossils of modern- Components of Energy flow = energy passage thru ecosys like humans were found in ecosystem Chemical cycling =mat transfer w/in ecosys Africa/stone tools support African Describe producers & PRODUCERS = autotrophs origin/DNA suggests a founding pop’n consumers - Where sunlight is main energy source from Africa/supported by mol data for life on earth - Plants, algae (plant-like protists), bac 2. Multiregional hypothesis w/c use light/chem to make food - Pre-modern humans migrated from Africa to become modern humans in CONSUMERS = heterotrophs other parts of the world - Organisms that rely on others for their - Interbreeding among diff H. erectus energy & food supply grps and a common origin for H. - Carnivores/herbivores/omnivores/ sapiens on diff continents decomposers 3 milestones in man’s 1. Erect stance Photosynthesis Using light energy to convert CO2 & H2O cultural history 2. Brain enlargement & prolonged post- into 02 & Carbs (6CO2 + 6H2O = birth dev’t of skull contents 6O2+C6H12O6) 3. Evol of a prolonged childhood during Chemosynthesis Performed by bacteria; using chem energy w/c cultural info is passed bet gens to prod carbs Define culture Includes the accu knowledge, customs, Ex. Deep-sea vent in benthic zone; energy bliefs, passed bet gens; gives us power to from chems from magma change envi Food chain Series of steps in w/c organisms transfer Major stages of culture 1. Scavenging-hunting-gathering energy by eating & being eaten; shows the - Organization into communal grps path of energy from 1 living thing to 2. Agriculture (slash-&-burn) another; decomposers are needed here - Led to pop’n increases Trophic levels Each step in a food chain/web - Thru advanced plows & overgrazing, led to soil erosion 1. Level 1 – producers (autotrophs) - Changed the rel bet humans & 2. Level 2 – primary consumers (herbi) biosphere 3. Level 3 – secondary consumers (carni - Led to est of permanent settlements or omni) - Allowed mems of many cultures to 4. Level 4 – tertiary consumers (usu top specialize in other activities carnivores) 3. Machine age Ecological pyramids The relative amt of energy or organisms - Industrial revolution in England contained in each trophic level of a food - From small-hand tools to large-scale chain/web machines Rule of 10 Only abt 10% of available energy w/in a - Complex machs reduced need for agri trophic level is passed to next higher level 1st law of That energy cannot be created or mimics a harmful one thermodynamics destroyed, only transformed - Mullerian mimicry = 2 unpalatable sps 2nd law of That every energy exchange increases the mimic each other thermodynamics universe’s entropy GLOBAL ECOLOGY Law of conservation of That matter cannot be created or Biosphere Global ecosys; sum of all planet’s ecosys; mass destroyed; chem elems are continually atmos, lakes, oceans, streams recycled w/in ecosystem Most impt abiotic - Solar energy Biogeochem cycles 1. WATER CYCLE factors - Temp - Environmental h2o cycle = - H2o atmospheric water vapor condenses & - Nutrients falls to earth’s surface; heated by sun - Other aquatic/terrestrial factors & reenters atmos by evap Rachel carson 2st to see dangers of pesticide abuse; - Organismic h2o cycle = surface h2o wrote concerns in 1962 book Silent Spring taken by plant roots, reenters atmos Aquatic biomes = 75% of earth’s surface thru transpiration from leaf surface - Wetlands - Lakes 2. CARBON CYCLE - Rivers, streams - Carbon enters atmos as CO2, w/c - Intertidal zones plants take thru photosyn & prod - Oceanic pelagic biome glucose - Coral reefs - Organisms eat plant bodies and store - Benthos carbon into their own tissue Terrestrial biomes - Tropical forest - Most organisms extract energy from - Chaparral food source & release CO2 as by - Temperate grassland product of respiration - Tundra - Desert 3. NITROGEN CYCLE - tropical, dry forest - N2 comprises 80% of atmos but it’s Niche = involves both an organism’s habitat & often the scarcest resource b/c it roles needs to be converted to NH4 or NO3 = Consists of: to be absorbed by plants thru nitrogen - role in ecosys (herbi, carni, etc.) fixation - tolerance limits (soil PH, humidity) - reqs for shelter (nesting sites, all 4. PHOSPHORUS CYCLE varying thru time) - Plants absorb phosphate ions from soil Kinds of biodiv 1. Ecosystem diversity & convert them to organic compounds - diff habitats, niche, sp interaction - Decomposers return phosphates to 2. Species diversity soil - Diff kinds of organisms, rels among sps 3. Genetic diversity Interspecific 1. Competition (-/-) - Diff genes & gene combos w/in pop’ns interactions - 2 organisms of same/diff sp try to use Megadiverse countries Grp of countries that harboer majority of a shared eco resource @ same time @ earth’s sps & are thus considered same place extremely biodiverse 2. Mutualism (+/+) Coral triangle = marine area in W. Pacific Ocean; includes - Both sps benefit waters of Indonesia, Malaysia, Ph, PNG, 3. Predation (+/-) Timor Leste, Solomon Islands - Where 1 sp kills & eats other for food = global center of marine biodiversity 4. Herbivory (+/-) = global priority for conservation - Where an herbivore eats plant/alga = Amazon of the seas - Led to evol of plant mech and chem Endemism When a sp lives only in a certain loc, such defenses & herbivore adaptations as a specific island/habitat type; found only 5. Commensalism w/in a specific region - Where 1 benefits & the other is Indigenous Found both w/in region & elsewhere neither helped/harmed Native species Plants/animals living in an area w/o being 6. Parasitism (+/-) brought there by human activity - 1 organism lives inside another & Biodiversity hotspot Biogeog area that is a significant reservoir harms it; parasite gets all/some of its of biodiv & is threatened w/ destruction nutri needs from host Causes of biodiversity 1. Direct = hunting, coll’n, persecution Symbiosis any rel where 2 sps live closely together loss 2. Indirect = habitat destruction & modi Adaptations - Introduced species Those that humans move from native locs - Cryptic coloration = canyon tree frog to new geog regions; may spread rapidly - Aposematic coloration = chem (w/o predators & pathogens), but may defense in form of bright warning disrupt their adopted community coloration; poison dart frog CLIMATE - Batesian mimicry = a harmless sp habitat & biodiv recovery; rebuild a Tradewinds Dry air descends & spreads back to equator functional ecosystem Restoration Biology To preserve & maintain existing habitat & Cooling tradewinds blow from E to W in biodiv; climate change mitigation; tropics; prevailing tradewinds blow from W preservation of endangered sps to E in temperate zones Restoration Biology To restore pop’ns to be able to persist & What causes seasons? The earth’s tilt evolve w/in larger communities over time Doldrums Area of calm & very light winds Conservation biology Seeks to preserve life and integrates fields Greenhouse effect Certain gases in atmos absorb heat & make of ecology, physio, mol bio, genetics, evol earth’s overall temp higher than it’d be if bio these gases weren’t present Pop’n fragmentation Harmful effect of habitat loss Greenhouse gases Allow short-wave radiation to pass thru & Splitting & isolation of Can lead to extinctions warm earth, but prevent long-wave portions of pop’ns radiation from escaping back to space Pop’n conservation Focuses on pop’n size, genetic diversity, and critical habitat 75% CO2, and 25% methane, nitrogen, 2 approaches to pop’n 1. Small pop’n approach oxides, ozone, and industrial GHGs conservation - Studies processes making small pop’ns Ozone Form of oxygen that protects life on earth extinct by shielding us from sun’s radiation - Extinction vortex = downward spiral CFCs Chlorofluorocarbons produced by human unique to mall pop’ns; driven by loss activity and w/c destroy atmos ozone of genetic diversity Ecological footprint Concept that describes out impact on - Min viable pop’n (MVP) size = smallest earth; uses idea of multiple constraints no. of inds needed to sustain a pop’n - Effective pop’n size = based on pop’ns Area of land & amt of H2O needed to inbreeding rate (4NfNm/(Nf + Nm)) support a human pop’n @ a particular - Pop’n viability analysis (PVA) = standard of living method of predicting if a sp will survive over time Dev’d by Wackernagel & Rees from Canada; world’s premier measure of our demand on 2. Declining pop’n approach nature - Proactive conservation strat for detecting, diagnosing, and halting Measurement of how much nat resources pop’n declines an ind uses in his lifestyle - More action-oriented Biocapacity How much land & water are needed to Landscape ecology Aims to make biodiv part of land-use produce a resource & absorb its wastes planning; consists of ecosys mgmt. using prevailing tech Nature reserves Bio div islands in a sea of habiata degraded Ecological overshock When humanity’s eco resource demands by human activity; consider disturbances as are higher than what nature can give a fcnal component of all ecosys Ecological debtor If a country doesn’t have enough eco Restoration ecology Sci study of repairing disturbed ecosys thru country resources in its own territory and there’s a human intervention; assumption is envi local ecological deficit damage is reversible Ecological creditor If country has ecological remainder 2 key strats: country 1. Bioremediation = use of living CONSERVATION BIOLOGY & RESTORATION ECOLOGY organisms to detoxify ecosys (prokaryotes, fungi, plants can take up Biodiversity = biological diversity toxic molecules) = describes the no., variety, variability of 2. Biological augmentation = uses living organisms organisms to add essential mats to a = syn of “life on earth” degraded ecosys = most commonly refers to “species o Ex. Nitro-fixing bacteria diversity” Sustainable dev’t = dev’t that meets the needs of people Species diversity Variety of sps in ecosys/thruout biosphere today w/o limiting the ability of future Endangered species In danger of becoming extinct throughout gens to meet their needs all/sig part of its range = ultimate goal Threatened species Likely to become endangered in future = improves human condition while Causes for sps 1. Destruction of habitat conserving biodiv becoming endangered 2. Wildlife trade 3. Overhunting IUCN Int’l Union for Conservation of Nature Biodiversity areas in 1. Zamboanga Peninsula Mindanao 2. Davao Peninsula 3. Cape San Agustin Restoration Ecology To leverage human intervention to prod