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9 – CHUA, Jaydee Stephanie A.

1E - PH
IMPORTANT TERMS

Angiosperms The informal term for flowering plants; the largest division in the plant kingdom

Apical Meristem Neww cells are created in this area through mitosis and cytokinesis for the
growth of the stem

Axillary Bud A miniature shoot with a dormant apical meristem and several young leaves

Basal A broad group of most primitive flowering plants. They are mostly woody plants
Angiosperms that produce seeds and flowers. Examples are Waterlilies, magnolias, and
laurels

Bulbs Short shoots that have thick, fleshy leaves

Circular The most derived and strongest tracheary elements; all of the primary wall is
Bordered Pits underlain by secondary wall.

Collenchyma Unevenly thickened primary walls. Typically alive at maturity. Provide elastic
Cells support.
Companion Controls the sieve tube members.
Cells

Corms Vertical, thick stems that have thin, papery leaves

Cortex Interior to the epidermis that can be either simple and homogenous or very
complex and contains many specialized cells

Cuticle A layer made of cutin that makes the wall impermeable to water

Epidermis The outermost surface of an herbaceous stem consisting of a single layer of


living parenchyma cells

Eudicots Broadleaf plants such as roses, asters, maples and others.


Fibers Long; many types are dead, other types remain alive and are involved in
storage

Ground The equivalent stages of pith and cortex


Meristem

Guard Cells Enables the opening and closing of the stomatal pore

Internodes The region between nodes

Metaphloem The mature cells of vascular bundles that are closest to the metaxylem

Metaxylem The largest tracheary elements of all

Monocots Named for and recognized bythe single cotlyedon, or seed leaf, within the
seed. Examples are grasses, lilies, cattails, palms, philodendrons, bromeliads
and several others
Nodes Criticial areas from which leaves, branches, and aerial roots grow out from the
stem

Botany Homework#1
9 – CHUA, Jaydee Stephanie A.
1E - PH
Parenchyma Thin primary walls. Typically alive at maturity. Many functions.

Perforation A large hole digested through a particular site of the primary wall during the
final stages of differentiation

Phloem A type of vascular tissue that distributes sugar and minerals

Phyllotaxy The arrangement of the leaves on the stem

Pits Wall depositions that came from low depressions in the developing secondary
walls
Pit-membrane Between the set of primary walls and middle lamella

Pit-pair Formed when the pits of adjacent sclerenchyma cells meet

Pith A region of parenchyma similar to the cortex


Primary Growth The growth and tissue formation that results from apical meristem activity

Primary Tissues Arise from apical meristems. It includes the epidermis, xylem, phloem and the
ground tissue

Protoderm Epidermal cells that are in the early stages of differentiation.


Protophloem The mature exterior cells of vascular bundles

Protoxylem The first xylem to appear

Provascular Young cells of xylem and phloem.


Tissues

Rhizomes Fleshy horizontal stems that allow a plant to spread underground


Sclereids More or less isodiametric; often dead at maturity

Sclerenchyma Primary walls plus secondary walls. Many dead at maturity. Provide elastic
support and some are involved in water transport.

Sieve Areas Cluster of sieve pores

Sieve Cells A type of conducting cells in phloem that must remain alive in order to conduct

Sieve Plates End-wall sieve areas with large sieve pores

Sieve Pores Plasmodesmata that enlarges to a diameter of more than 1 micrometer

Sieve Tube Stacked end to end with their large sieve areas aligned

Sieve Tube Sieve areas on the sides of the cell that are rather small
Members

Stoma A constitution of stomatal pores

Tracheary Refers to both tracheids and vessel elements


Element

Botany Homework#1
9 – CHUA, Jaydee Stephanie A.
1E - PH
Tracheids Long and narrow with tapered ends; contain no perforations. Dead at maturity.
Found in all vascular plants.

Trichomes Epidermal cells that elongate outward; make it difficult for an animal to land on,
walk on or chew into a leaf.

Tubers Horizontal like rhizomes but grow for only a short period and mainly a means of
storing nutrients.

Vascular Xylem and phloem; located just interior to the cortex.


Bundles

Vessel An entire stack of vessel elements

Vessel Short and wide with rather perpendicular end walls; most contain one or two
Elements perforations. Dead at maturity. Found almost exclusively in flowering plants.
Among nonflowering plants, only a few ferns, horsetails, and gymnosperms
have vessels.

Xylem A type of vascular tissue that conducts water and minerals


REVIEW QUESTIONS

# QUESTIONS ANSWERS

1 The body of an herb contains just three basic The body of an herb contains just three basic parts:
parts. What are they? leaves, stems and roots.

2 Stems of the first land plants functioned Aside from functioning primarily as transport and
primarily as transport and support structures. support structures, they also reproduce leaves and
Modern stems have these functions also, plus hold them in the sunlight, and during winter, they store
several more. Describe the additional sugars and other nutrients, such as the sugary sap of
functions. maples. Stems may also be a means of survival
(underground bulbs and corns) and as a means of
dispersal (runners, vines or breaking off of pieces).
3 Over the years, several different names have Over the years, several different names have been
been used for the flowering plants. Here they used for the flowering plants. Here they are called the
are called the division _____, but they are also division Magnoliophyta, but they are also known
known informally as _____. There are about informally as angiospersms There are about 297 000
_____ species of flowering plants. species of flowering plants.

4 Early angiosperms diversified into severalEarly angiosperms diversified into several groups that
groups that are known as _____and _____. are known as basal angiosperms (waterlilies,
Name three examples of plants within each magnolias, laurals), eudicots (broadleaf plants such as
line of evolution. roses, asters, maples) and monocots (grasses, lilies,
cattails, palus, philodendrons, bromeliads.
5 An herb has only a primary plant body. An herb has only a primary plant body. That means it
That means it has roots, stems, and has roots, stems, and leaves, but it never becomes
leaves, but it never becomes _____ and woody and covered with bark.
covered with _____.
6 In general, animals have ______ growth with a In general, animals have determinate growth with a
fixed size and number of organs, whereas fixed size and number of organs, whereas plants have
plants have _____ growth with no net size and indeterminate growth with no net size and any number
Botany Homework#1
9 – CHUA, Jaydee Stephanie A.
1E - PH
any number of organs. Which parts of our body of organ
grow continuously like plants? Which parts of a
. Our ears and noses grow continuously like plants.
plant have fixed size such that no matter how
well we take care of the plants those organs do Annual plants live for just one year while perrenial
not become larger? What is the name of the plants are those who live for just two years.
plants that live for just 1 year? For just 2
years?

7 What are the important differences between Parenchyma cells are primary walls that are typically
parenchyma, collenchyma and sclerenchyma alive at maturity. They have many functions such as
cells? involvement in photosynthesis, secretion of nectar,
fragments, mucilage, resins and oils, and many more.
Collenchyma cells are evenly thickened walls that are
also typically alive at maturity. They are the ones that
provide plastic support. The sclerenchyma consists of
primary and secondary walls. Many of them are dead
at maturity. They provide elastic support and some
(tracheary elements) are involved in water support.
8 Examine Figure 5-6A and 5-6B. Why do most Since nuclei are large cells and the section (slice)
parenchyma cells shown in figure 5-6A lack were cut so thin, most nuclei were cut away during the
nuclei? In the geranium for Figure 5-6A, did the preparation of the microscope slide.
real cells lack nuclei or did something happen
to the nuclei as the tissue was sliced to make
the microscope slide?
9 What is the special name of photosynthetic The photosynthetic parenchyma cell is called the
parenchyma cells? Which organelle is Chlorenchyma cells. Chloroplasts are especially
especially abundant in these? How does wall abundant in these. Wall thinness allows light and
thinness affect carbon dioxide and light? carbon dioxide to pass through in the chloroplast.
10 It was previously stated that parenchyma cells Little glucose is expended in constructing the cellulose
are relatively inexpensive to build. What does and hemicellulose of such thin walls. Most leaves are
that mean? How does this relate to most soft since they are composed almost entirely of
leaves being soft? parenchyma.
11 Like clay, walls of collenchyma exhibits Collenchyma, which is present in tips, makes the tips
plasticity. Does that mean it can or cannot be stronger and more resistant to breaking; however, the
stretched? If a tissue is supported by tips can still elongate because collenchyma can be
collenchyma, can it still grow? stretched. Yes, it can still grow.
12 How does collenchyma support a tissue? No, collenchyma and turgid parenchyma must work
Would it work if the tissue did not have together to support a tissue. Collenchyma must be
parenchyma also? What happens if the tissue confined by the turgid parenchyma. The tendency for
is not turgid (does not have enough water)? parenchyma to expand is counterbalanced by the
resistance of the collenchyma, and the stem becomes
rigid but able to grow. If a vine or other collenchyma-
rich tissue is cut off from its water supply, it wilts and
droops; the collenchyma is unable to hold up the
stem.

Botany Homework#1
9 – CHUA, Jaydee Stephanie A.
1E - PH
13 Collenchyma is plastic, but sclerenchyma is Elastic means the ability to be deformed, but returns
elastic. What does “elastic” mean? If you to their original size and shape when the pressure or
stretch or deform an elastic object, will it keep tension is released. If you stretch or deform an elastic
its new shape or snap back to its original object, it will snap back to its original shape.
shape? As an organ grows, its shape changes, Deforming forces such as wind, animals or snow will
but after it has achieved its mature size and deform a mature organ. A mature organ should have
shape, what kind of things would deform it? elastic properties to maintain its shape (the branch
Should a mature organ have plastic properties should stay in its bent shape because going back to
or elastic ones (e.g., if a heavy load of snow its original shape or to the shape it grew before the
bends a branch down, should the branch stay snow would be optimal).
in its bent shape when the snow melts or
should it go back to the shape when the snow
melts or should it go back to the shape it grew
before the snow)?
14 What are the two types of mechanical, The two types of mechanical, nonconducting
nonconducting sclerenchyma (hint:Table 5-3)? sclerenchyma are the sclereids and fibers. The fibers
Which tends to be flexible and useful in wood? tend to be flexible and useful in wood. The sclereids
Which tends to be brittle and inflexible, useful tend to be brittle and inflexible; they are useful in “pits”
in “pits” and “stones” that protect seeds? and “stones” that protect seeds.
15 Fibers and sclereids have secondary walls that Fibers and sclereids grow from cells produced from
are so thick and tough that the cell cannot cell division and when newly formed, they are small
grow. If that is true, how do fibers and sclereids and have only a primary wall which means they are
grow to their mature size and shape? Do they parenchyma cells.
have a secondary wall even when they are
young?
16 Imagine a leaf of a palm tree. When it is fully Sclerenchyma cells originated from parenchyma cells
grown, it must have sclerenchyma to support which means that the tiny and growing tree doesn’t
its size and weight. Do you think it has any have any sclerenchyma yet. Parenchyma cells and
sclerenchyma when it is tiny and just starting to collenchyma cells are the only one that are present.
grow? Would you expect that it might have
collenchyma when it is medium sized and still
growing?
17 Technically, the _____ is an axis, whereas the Technically, the stem is an axis, whereas the shoot is
_____ is the _____ is the _____ plus any the stem plus any leaves, flowers or buds that might
leaves, flowers or buds that might be present. be present.
18 The point where a leaf is attached to a stem is The point where a leaf is attached to a stem is called a
called a ______. Just above this point is an node. Just above this point is the leaf axil which
_____ bud. contains the axillary bud.
19 Figure 5-13A shows two types of buds. What Two types of buds, namely bud scales and spines are
are the two types and how do they differ? present in the figure. Bud scales protect the delicate
organs inside while spines serves protection of the
plant.
20 After a leaf falls off a stem, it leaves a After a leaf falls off a stem, it leaves a leaf scar just
____________________ just below the axillary below the axillary bud.
bud.

Botany Homework#1
9 – CHUA, Jaydee Stephanie A.
1E - PH
21 What is phyllotaxy? Corn and irises have two Phyllotaxy is the arrangement of leaves on the stem.
rows of leaves. This is known as Distichous phyllotaxy is present in corn and irises.
____________________ phyllotaxy
22 When you look at a head of cabbage or When we look at a head of a cabbage or lettuce, the
lettuce, what are you seeing? Where is the only thing we see are leaves. When we look at an
stem? What about when you look at an onion? onion, we see the fleshy onion leaves and the short
What are you seeing and where is the onion’s stem is within the leaves.
stem?
23 Describe each of the following types of Modification of leaves, internodes, orientation of
specialized shoots. Be certain to account for growth
modifications of the leaves, internodes, and
Stolon’s internodes are especially long and thin, and
orientation of growth: stolon, rhizome, tuber,
their leaves do not expand. They extend greatly but do
bulb, corm, and tendril. Each provides a plant
not use much of the plant’s nutrient reserves. After the
with a selective advantage. What is the
stolon encounters a suitable microhabitat, subsequent
adaptive value of each type of specialized
growth is by shorter, vertical internodes and fully
shoot?
expanded leaves; new roots are established, and the
end of the stolon resembles a new plant.
Rhizomes are fleshy horizontal stems that allow a
plant to spread underground.Tuber are horizontal like
rhizomes, but they grow for only a short period and
are mainly a means of storing nutrients.
Bulbs are short roots that have thick, fleshy leaves.
Corms are vertical, thick stems that have thin papery
leaves. There is no obvious selective advantage of
one over the other; the type seems to depend on
whether mutations affecting the stem or leaf happen to
occur first.
Tendril’s leaves can twine around. Their internodes
are elongated. Their orientation of growth is vertical.
Tendrils provide support and protection using its
modified leaves and lateral branches that are capable
of twinning around small objects.

24 Many scientific words are also ordinary English As long as you know the features that make them
words. If you read that a plant has a bulb or a distinct from others, you can always be certain what
tuber, can you always be certain exactly what the botanical structure is.
the botanical structure is?
25 What is the outermost surface of an The outermost surface of an herbaceous stem is the
herbaceous stem? The outer walls of this layer epidermis. The cuticle makes the wall impermeable to
are encrusted with a chemical made up of fatty water.
substance that makes the wall impermeable to
water. What is the name of the substance and
what is the name of the layer?

Botany Homework#1
9 – CHUA, Jaydee Stephanie A.
1E - PH
26 A stoma allows carbon dioxide to pass through The guard cells control the opening and closing of a
the epidermis. What is the name of the two stoma. The stomatal pore is where the carbon dioxide
cells that control the opening and closing of a passes through.
stoma? What is the name of the hole itself that
carbon dioxide passes through?
27 What is the technical term for a plant hair? Trichomes is the technical term for a plant hair. They
Plant hairs make it difficult for animals to do make it difficult for an animal to land on, walk on, or
certain things. Name three activities that are chew into a leaf.
more difficult for an animal because of a hairy
leaf.
28 What is the name of the region of cells The cortex is between a stem’s epidermis and its set
between a stem’s epidermis and its set of of vascular bundles. In some aquatic angiosperms,
vascular bundles? This is usually a compact they have large cortical air chambers that provide
parenchyma tissue, but in some aquatic buoyancy. The stems have a strong tendency to float
angiosperms that live submerged in lakes or that no sclerenchyma is necessary for support. I would
oceans, this region has what type of special hypothesize that this region is especially thin in desert
modification? Would you hypothesize that this plants such as cacti or plants that have succulent
region is especially thick or thin in desert plants water storage tissues.
such as cacti or plants that have succulent
water storage tissues (you must think about
this yourself, as the answer is not in the text).
29 All flowering plants have two types of vascular All flowering plants have two types of vascular tissues,
tissues, ______, which conducts water and xylem which conducts water and minerals, and
minerals, and ______, which distributes sugars phloem which distributes sugars and minerals.
and minerals. Are vascular systems of plants Vascular systems of plants are quite different from
similar to or different from those of animals? those of animals. Animals have a circulatory system
Explain the similarity or differences. while the vascular systems of plants are not a circular
one. Water and minerals enter xylem in the roots and
are conducted upward to leaves and stems. Xylem
sap travels through dead, hollow cells, not through in
tubes composed of living cells like our blood vessels.
Phloem pick up sugar from areas where it is abundant
and transport it to areas where sugar is needed.
30 As a young xylem cell matures into a tracheary As a young xylem cell matures into a tracheary
element, it first must enter ______ and stop element, it first must enter cell cycle arrest and stop
______. It is initially a small ______ cell, but dividing. It is initially a small parenchyma cell, but after
after it reaches its full size and shape, it it reaches its full size and shape, it deposits a
deposits a ______ wall that reinforces the secondary wall that reinforces the primary wall. The
______ wall. The xylem cell ______ and its xylem cell then dies and its protoplasm degenerates,
protoplasm ______, leaving a ______ wall. leaving a hollow tubular wall.
31 List the five types of secondary wall deposition There are five types of secondary wall deposition that
that can occur in the tracheary elements. can occur in the tracheary elements: annular
Which two types are most characteristic of thickening, helical thickening, scalariform thickening,
protoxylem? What is the selective advantage reticulate thickening, and circular bordered pits.
of vessel elements over tracheids? Protoxylem cells have annularand helical thickenings.
Vessel elements provide a way to move water with

Botany Homework#1
9 – CHUA, Jaydee Stephanie A.
1E - PH
less friction than tracheids.
32 Consider tracheary elements with annular The annular thickening is weaker than those with
thickening (annular secondary walls) and those pitted walls. It also has a large percentage of its
with pitted walls. Which is weaker? Which has primary wall free of secondary wall and available for
a large percentage of its primary wall free of water movement. None of them is always more
secondary wall and available for water selectively advantageous than the other since each
movement? Is one always more selectively has its own weaknesses and strengths.
advantageous than the other? Under all
conditions? Explain.
33 Tracheids obtain water from ______ below Tracheids obtain water from other tracheids below
them and pass it on to ______ above. As water them and pass it on to those above. As water passes
passes into or out of tracheid, it passes into or out of tracheid, it passes through a pit of the
through a ______. The set of primary walls and adjacent tracheids. The set of primary walls and
middle lamella in the ______-______ is called middle lamella in the pit-pair is called a pit membrane.
a ______.
34 If each tracheid is 1 mm long, how many pit- If each tracheid is 1 mm long, water molecule will pass
pairs and pit membranes will a water molecule through a plant 1 m long by a thousand mm. The pit
pass through in a plant 1 m tall? Is this a pairs and pit membranes will also have the same
significant amount of friction? amount. As the water molecules pass through, the
friction will add up.
35 What is a perforation in a vessel element? How Perforation in a vessel element is the hole where both
does it reduce friction as water passes from primary and secondary walls are missing; thus, only
one vessel element to another? When a water little friction is created. When a water molecule arrives
molecule arrives at the end of a vessel, how at the end of a vessel, pit-pairs serve as the only
does it pass into another vessel (this is asking passage of water from one vessel to another. If
about vessels, not vessel elements). In tracheids were assumed to be 1 mm long, there are 3
Question 34, tracheids were assumed to be 1 020 vessel elemnts in an average vessel of American
mm long. If we assume that vessel elements beech. An American beech this long would not fit into
are also 1 mm long (they are usually shorter, a 1 m long. If a plant 1 m tall had vessels 1 m long, 1
but 1 mm is an easy number to work with), how 000 pit membranes had to be passed through by
many vessel elements are there in an average water molecules from root to tip. This is the same as
vessel of American beech (Table 5-7, the that of the answer for question 34.
lengths in the table are in centimeters)? The
plant in Question 34 was said to be 1 m tall.
Would an average vessel of American beech
even fit into a plant this tall? If a plant 1 m tall
had a vessels 1 m long, how many pit
membranes would a water molecule needed to
pass through as it traveled from root to shoot
tip? Is this more or less than the answer for
Question 34?
36 Which evolved more recently, tracheids or Vessel elements evolved more recently. Yes, they do.
vessel elements? Do angiosperms such as Yes, angiosperms both have vessel elements and
conifers and ferns have vessel elements? Do tracheids; their tracheids occur mostly in the fine veins
flowering plants (angiosperms) have of leaves.
tracheids? Vessel elements?

Botany Homework#1
9 – CHUA, Jaydee Stephanie A.
1E - PH
37 Like xylem, phloem has two types of Like xylem, phloem has two types of conducting cells,
conducting cells, _____ and _____ The term sieve cells and sieve tube members The term “sieve
“_____” refers to either one. Do these die like element” refers to either one. They need to remain
tracheary elements or do they need to remain alive in order to conduct.
alive in order to conduct?
38 What is the name of the holes that interconnect Sieve pores are the name of holes that interconnect
conducting cells in phloem, and what are conducting cells in phloem. The group of these holes
groups of these holes called? are called the sieve areas.
39 Sieve tube members differ from sieve cells by Sieve plates are the name of the end-wall sieve areas
having very large sieve pores on their end with big sieve pores.
walls, much larger than the ones on their side
walls. What is the name of these end-wall
sieve areas with big sieve pores?
40 Sieve elements (both sieve cells and sieve Sieve cells are associated with albuminous cells. The
tube members) lose their nuclei during one associated with sieve tube members are
development, but they must remain alive. What controlled by companion cells.
is the name of the cell associated with sieve
cells? The one associated with sieve tube
members?
41 During differentiation of a young cell into a During differentiation of a young cell into a sieve tube
sieve tube member, what are some changes member, the amount of cytoplasm in the pores and
that occur in the cell wall and the cytoplasm? the permission of rapid bulk movements increase. The
How long do most sieve tube members live sieve pores of adjacent cells should also be aligned so
after they become mature? the phloem sap can pass through. After they mature,
most sieve tube members live as companion cells.
42 Describe plasmodesmata, pits, perforations, Plasmodesmata are a thin thread of cytoplasm that
and sieve pores. Which connect living cells? pass through the walls of the neighbouring plant cells.
Which connect non-living cells? Which occur in Pits are channels where water passes through to
secondary walls? different parts of the plant cell. Perforations are form
of holes in which they can digest by a particular site of
the primary cell wall. Pits and perforations connect
living cells while plasmodesmata connect non-living
ones and occur in secondary walls.
43 All vascular bundles are _____. That is, All vascular bundles are collateral. That is, contains
contains both xylem and phloem. The xylem of both xylem and phloem. The xylem of a vascular
a vascular bundle is _____ xylem because it is bundle is primary xylem because it is part of the
part of the primary plant body, and the phloem primary plant body, and the phloem is primary phloem.
is _____ phloem
44 Stems grow longer by creating new cells at Stems grow longer by creating new cells at their tips,
their tips, in regions known as _____. Below in regions known as shoot apical meristems. Below
this region, in the subapical meristem, the very this region, in the subapical meristem, the very first
first primary xylem to appear is called _____, primary xylem to appear is called protoxylem, and
and cells that differentiate into xylem a little cells that differentiate into xylem a little later, after they
later, after they have grown larger, are called have grown larger, are called metaxylem.
_____.

Botany Homework#1
9 – CHUA, Jaydee Stephanie A.
1E - PH
45 In the apical and subapical region, there are a. Protoderm are the epidermal cells that are still
cells that will later give rise to epidermis and meristematic.
other tissues, but they are not yet mature b. Provascular tissues are the young cells of
enough to call them epidermis, and so on. xylem and phloem.
What are the terms for the following cells? c. Ground meristem are the young cells of pith
and cortex.
a. Epidermal cells that are still meristematic
b. Young cells of xylem and phloem
c. Young cells of pith and cortex
46 Describe the arrangement of tissues seen in a In the stem cross-secrion, the xylem consists mostly
stem cross-section; consider monocots of certain rows of vessels. These rows of vessels are
separately from the other angiosperms. Is the alternated with the rows of xylem parenchyma.
arrangement different in a stolon than in a However, the phloem mainly includes companion cells
rhizome, tuber or corm? and sieve tube members. Even they are huge
sclerenchyma cap in phloem fibers. The arrement is
different in a stolon than in a rhizome, tuber or corm.
47 Look at Figure 5-42. The top three rows are The five tissues are the epidermis, primary xylem,
labeled “Primary growth” because these are primary phloem, pith and cortex.
found in the primary plant body, the body of an
herb. The third row of the figure has five boxes
with the five types of mature tissues found in
the primary plant body. What are the five
tissues?
48 If you live to be 100 years old, how many sets A human will only have one set of organs during his
of organs will you have during your lifetime? lifetime cosisting of one of each organ (excluding
How many hearts, stomachs, and livers? If a those who come in pairs). If a plant lives to be 100
plant lives to be 100 years old, how many sets years old, it will have the same number of leaves. If it
of leaves will it probably have? If it begins begins blooming when it is 10 years old, it will also
blooming when it is 10 years old, how many have the same set of flowers. Yes, I will still depend
sets of flowers? When you are 70 years old, on my digestive sustem to supply myself with energy
you will still depend on your digestive system for another 30 years; my digestive system and I both
to supply you with energy for another 30 years, have the same number of years/same age. In a 70-
but how old is that set of organs? In a 70-year- year old tree, the cells of the leaves that supply it with
old tree, how old are the cells of the leaves that energy is also 70 years old.
supply it with energy?

Botany Homework#1

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