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To cite this article: Sakine Serap Avgın, İrem Dertli & Arvīds Barševskis (2014): A review of Turkish saproxylic beetles from
the European Red List, Annales de la Société entomologique de France (N.S.): International Journal of Entomology
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Annales de la Société entomologique de France (N.S.), 2014
http://dx.doi.org/10.1080/00379271.2014.896099
Summary. The conservation of beetles that depend on dying or dead wood (i.e. saproxylic), has received a great deal of
attention in many parts of the world in recent years. Human activities such as urbanization and logging, and their results, e.g.
global warming, destroy natural ecosystems and threaten unusual species such as Lucanus cervus (L. 1758), Rosalia alpina (L.
1758), Limoniscus violaceus (Müller 1821) and Osmoderma eremita (Scopoli 1763). Attempts have been made to conserve
the habitats upon which these species depend. Although represented by a large number of species, saproxylic beetles in Turkey
have not received much attention. Although some Turkish studies have acknowledged their value, comprehensive investiga-
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tions of these species are very limited. The aim of this study is to make a list, from the literature, of the species that are
endangered in Europe and found in Turkey, to highlight the importance of these species. The research includes 151 saproxylic
beetle species belonging to the families Bostrichidae, Cerambycidae, Elateridae, Eucnemidae, Scarabaeidae, Euchiridae,
Lucanidae and Mycetophagidae. The list of Turkish saproxylic beetles was prepared using the European Red List of
Saproxylic Beetles. Information is provided about the IUCN Red List category in Europe, the locations of the species in
Turkey, and their global geographic range. We gathered all data about these species from the international literature.
Furthermore, we also include in the list certain species whose locations are not specified, but which inhabit Turkey. We
obtained their records from some catalogs in Turkey. In addition to the list, we prepared three distribution maps for Turkey.
The maps show distribution in Turkey of some species in the Endangered, Vulnerable and Near Threatened categories in the
European Red List. It is expected that this study will lead to comprehensive studies on saproxylic beetles from Turkey.
Résumé. Revue des coléoptèrs saproxyliques turques de la Liste Rouge Européenne. La conservation des coléoptères
qui dépendent du bois mort ou pourrissant (dits saproxyliques) a reçu beaucoup d'attention depuis quelques années. Ls
activités humaines comme l'urbanisation, l'exploitation forestière et le réchauffement global, en détruisant des écosystèmes
naturels, menacent ainsi jours après jours des espèces uniques telles que Lucanus cervus (L. 1758), Rosalia alpina (L.
1759), Limoniscus violaceus (Müller 1821), Osmoderma eremita (Scopoli 1763). De nombreux chercheurs d'Europe se sont
réunis afin de proposer des mesures de conservation des habitats dont ces espèces dépendent. Malgré qu'ils soient
représentés par un grand nombre d'espèces, les coléoptères saproxyliques de Turquie n'ont pas reçu beaucoup d'attention.
Même si des chercheurs turcs ont mentionné leur valeur dans plusieurs études, les recherches détaillées sont encore très
limitées. Le but de ce travail est de constituer, à partir de la littérature, une liste des espèces qui sont à la fois menacées en
Europe et présentes en Turquie. La recherche inclut 151 espèces de coléoptères saproxyliques de Turquie qui appartiennent
aux familles Bostrichidae, Cerambycidae, Elateridae, Eucnemidae, Scarabaeidae, Euchiridae, Lucanidae et
Mycetophagidae. La liste des coléoptères saproxyliques de Turquie a été préparée au départ de la Liste Rouge des
espèces d'Europe. Dans cette liste, nous avons donné pour chaque espèce des informations sur les catégories de menace
en Europe, les localisations en Turquie et dans le reste du monde. Nous avons rassemblé les données de ces espèces à partir
de la littérature internationale. Par ailleurs, certaines espèces dont les localisations ne sont pas précisées mais qui existent en
Turquie sont seulement énumérées. Nous présentons aussi trois cartes de distributions en Turquie, pour les espèces en
Danger, Vulnérables et Presque Menacées de la Liste Rouge des espèces d'Europe. Nous espérons que cette étude va
conduire à des recherches plus complètes des coléoptères saproxyliques de Turquie.
Keywords: European Red List of saproxylic beetles; forest biodiversity; diversity; Coleoptera; Middle Asia
Kaila et al. (1994) state that saproxylic beetles often saproxylic species native to Turkey that appear on the
exhibit a high degree of habitat specificity, and that vari- European Red List.
ables including tree species, diversity and types of fungus
species decaying the wood, degree of decay, and quality
and structure of the surrounding biotope may affect the Material and method
occurrence of saproxylic insect species. Because rotten The material of this work is the European Red List of
wood is inhabited by the larvae of saproxylic beetles, it Saproxylic Beetles published by International Union for
is difficult to examine their conditions of microhabitat in Conservation of Nature (IUCN 2012). A lot of saproxylic beetle
species are classified in different threatened categories in the
detail without destroying them. Therefore, often the adults
European Red List. These categories are presented in Figure 1.
of saproxylic beetles are studied. Knowledge about the When the categorization was made, assessment protocols were
habitat requirements of saproxylic beetles is less detailed used taking into account geographic range, data of population,
than for other insects such as butterflies (Ranius 2002). habitat preferences, etc. (Nieto & Alexander 2010; IUCN
Saproxylic beetles have important interactions with 2012). The research on saproxylic beetles is very limited; there-
fore the threat status of these beetles is unknown in Turkey.
other organisms, e.g. mites, nematodes, bacteria and
During this study, no specimens were collected from any
fungi, that are significant for ecosystem and economy. locations and no statistical processing for threatened status has
The beetles act as vectors, helping to disperse these organ- been used. First, we examined the European Red List of
isms by carrying them on or within their bodies. Another Saproxylic Beetles and we determined saproxylic beetle species
important role performed by some kinds of saproxylic living in Turkey from the list. For detecting these species we
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Figure 1. Red List categories after International Union for Conservation of Nature (IUCN) (from Nieto & Alexander 2010, with
authorization).
Figure 2. Map showing distribution of species in Turkey in the Endangered category in the European Red List of Saproxylic Beetles. 1,
Adelocera pygmaea (Baudi 1871); 2, Tetrigus cyprius Baudi 1871; 3, Callergates gaillardoti (Chevrolat 1854); 4, Limoniscus violaceus
(Müller 1821); 5, Osmoderma lassallei (Baraud & Tauzin 1991).
Figure 3. Map showing distribution of species in Turkey in the Vulnerable category in the European Red List of Saproxylic Beetles. 1,
Delagrangeus angustissimus Pic 1892; 2, Proetia mirifica (Mulsant 1842); 3, Stenopterus atricornis Pic 1891; 4, Ischnodes sanguini-
collis (Panzer 1793).
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Figure 4. Map showing distribution of some species in Turkey in the Near Threatened category in the European Red List of Saproxylic
Beetles. 1, Osmoderma eremita (Scopoli 1763); 2, Ampedus cardinalis (Schiödte 1865); 3, Ampedus elongatulus (Fabricius 1787); 4,
Calais parreysii (Steven 1830); 5, Ectamenogonus montandoni (Buysson 1888); 6, Elater ferrugineus (L. 1758); 7, Megapenthes lugens
(Redtenbacher 1842); 8, Epiphanis cornutus (Eschscholtz 1829); 9, Lacon lepidopterus (Panzer 1801); 10, Saperda punctata (L. 1767);
11, Propomacrus bimucronatus (Pallas 1781); 12, Lucanus cervus (L. 1758); 13, Farsus dubius (Piller & Mitterbacher 1783); 14,
Rhaesus serricollis (Motschulsky 1838); 15, Cerambyx welensii (Küster 1846); 16, Cerambyx nodulosus Germar 1817; 17, Cerambyx
miles Bonelli 1812; 18, Cerambyx dux (Faldermann 1837); 19, Cerambyx cerdo L. 1758 (19).
Global distribution. Widespread in large parts of Global distribution. Mediterranean, and up to Hungary,
southern tropical areas in Tunisia (Sadok & Gerini the Caspian Sea and Iran. Reported from all over Italy and
1988), Tanzania (Kilambo et al. 2005), the islands (Nardi & Ratti 1995). This species is probably
Mediterranean area (Avidov & Harpaz 1969; Halperin & found on Ficus carica L. or Laurus nobilis L. in Venice
Damoiseau 1980) and in southern Italy (Bonsignore (Ratti 2004).
2012).
Xyloperthella picea Olivier 1790 and Nurhak (Tatlar) (Özdikmen & Okutaner 2006).
Variety A. moschata var. thoracica Fischer von
Red List category in Europe. LC.
Waldheim 1824 is recorded in Isparta (Tauzin 2000),
Distribution in Turkey. Aydın province: Germencik, Mersin (Malmusi & Saltini 2005) central Antalya as A.
Buharkent, Nazilli, Bozdoğan (Akşit et al. 2005). moschata var. thoracica Fischer von Waldheim 1824
(Demelt & Alkan 1962; Demelt 1963) (Özdikmen 2011a,
Global distribution. Angola (Danielson 2003), South 2011b). This variety is currently considered a synonym of
Africa, Egypt, Madagascar, Mediterranean region, Aromia moschata ambrosiaca (Steven 1809). The second
Palestine, Sardinia, Seychelles, Syria, Cameroon, Sicily subspecies found in Turkey is nominative Aromia
(Caillol 1954), Italy, Sardinia (Taglianti 2003) and moschata moschata (L. 1758) (Sama & Lobl 2010).
Turkey (Akşit et al. 2005).
Global distribution. A. moschata moschata (L. 1758)
occupies most of Europe including Britain, France,
Family Cerambycidae Germany (Althoff & Danilevsky 1997), Poland,
Aegosoma scabricorne (Scopoli 1763) Scandinavian and Baltic States (Silfverberg 2010), Czech
Republic, Austria, Slovakia (Adlbauer 2001), Ukraine
Red List category in Europe. LC.
(Zahaykevych 1991; Bartenev 2004; Zamoroka 2007),
Distribution in Turkey. Antalya province: Taşkent; Hungary (Csóka & Kovács 1999), Moldova (Baban
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Konya province: Taşkent–Alanya road, Afşar town: 2006), Byelorussia, Russia, Romania, Bulgaria, Albany
Kayadibi Akçapınar district (Turgut & Özdikmen 2010); (Althoff & Danilevsky 1997), Serbia (Pil 2005), Slovenia,
Ankara province: Kayaş, Bayındır dam env.; northern Italy (Brelih et al. 2006), and northern Spain
Kahramanmaraş province: Andırın as Megopis (Pena 2002). It is also found in Caucasus, Kazakhstan and
scabricornis (Scopoli 1763) (Adlbauer 1992); Isparta Turkey. A. moschata ambrosiaca (Steven 1809) occupies
province: Eğirdir (Ağıl) (Tozlu et al. 2002); Osmaniye countries of the Mediterranean basin including Portugal,
province: Bahçe road and Zorkun road (Özdikmen & Spain (Pena 2002), Italy, Greece (Althoff & Danilevsky
Turgut 2009c; Özdikmen et al. 2009, 2010; Özdikmen 1997) and southern France (Brustel et al. 2002; Zamoroka
2011b); Kirklareli province: Kanli Dere, Koprusu (Maican & Panin 2011).
& Serafim 2009); Samsun province: Çarşamba (Özdikmen
& Demir 2006).
Axinopalpis gracilis (Krynicki 1832)
Global distribution. Europe (Albania, Austria, Belarus,
Bosnia and Herzegovina, Bulgaria, Corsica, Crete, Red List category in Europe. LC.
Croatia, Czech Republic, Turkey, France, Germany,
Distribution in Turkey. Isparta province: Yukarıgökdere,
Greece, Hungary, Italy, Macedonia, Romania, southern
Eğirdir. (Sama et al. 2011), Şarkikaraağaç (Sama 1982);
Russia, Slovakia, Slovenia, Switzerland, Ukraine), Near
İçel province: Erdemli (Adlbauer 1988) (Özdikmen 2011a,
East (Maican & Serafim 2009).
2011b).
Turgut 2009c; Özdikmen et al. 2010; Özdikmen 2011a). Distribution in Turkey. Denizli province (Schimitschek
1944), Sarayköy (Gül-Zümreoğlu 1975); Aydın province:
Global distribution. Turkey, Syria, Lebanon (Özdikmen Germencik (Demelt 1963; Gül-Zümreoğlu 1972); Turkey
2008). (Lodos 1998; Alkan 2000; Özdikmen et al. 2005); Manisa
province: Muradiye; İzmir province: Kemalpaşa
Callidium aeneum (Degeer 1775) (Armutlu) (Tezcan & Rejzek 2002); Ankara province:
Güdül, Beytepe (Özdikmen 2007, 2008; Özdikmen et al.
Red List category in Europe. LC. 2009).
Distribution in Turkey. Rize province: Çamlıhemşin Global distribution. Croatia, Bosnia and Herzegovina,
(Alkan & Eroğlu 2001). Yugoslavia, Macedonia, Crete, Bulgaria, Malta
Global distribution. Europe, Caucasus with (Danilevsky 2007), Turkey (Özdikmen 2008; Özdikmen
Transcaucasia, Middle East, Turkey, Syria, northern Iran, et al. 2009).
Siberia, Mongolia, Far East, Japan, northern China (Sama
& Lobl 2010). This species has two subspecies. In Turkey
is distributed the nominative subspecies Callidium aeneum Cerambyx cerdo L. 1758
aeneum (Degeer 1775). The second subspecies Callidium Red List category in Europe. NT.
aeneum longipenne Villiers 1978 is distributed in
Azerbaijan, Georgia and southern Russia (Sama & Lobl Distribution in Turkey. Muğla province: Milas (Gül-
2010) and it is possible in Turkey. Zümreoğlu 1972); İzmir province: Bergama, Bornova
(Gül-Zümreoğlu 1975); Denizli province (Özdikmen &
Şahin 2006); İçel province: Bolkar Mountains
Callimoxys gracilis (Brullé 1832) (Bodemeyer 1906); Adana province: Pozantı (Ulusoy
et al. 1999); Osmaniye province: Mitis’s plateau
Red List category in Europe. LC. (Özdikmen & Demir 2006); Kahramanmaraş province:
Pazarcık (Bağdınısağır) (Özdikmen & Okutaner 2006);
Distribution in Turkey. İzmir province: Çamlık pass
Ankara province: Hacıkadın (Özdikmen et al. 2005);
(Adlbauer 1988; Özdikmen 2008).
Kayaş (Bayındır dam env.) (Özdikmen & Demir 2006);
Global distribution. Yugoslavia, Croatia, Bosnia and Sinop province: Ayancık (Schimitschek 1944); Bursa
Herzegovina, Slovenia, Greece (without Crete), province: near Soğukpınar (Çanakçıoğlu 1956); Sakarya
Macedonia, Bulgaria, Romania, Hungary, Crimea, province: Sapanca (Nizamlıoğlu 1957); Kocaeli province:
Slovakia (Danilevsky 2007). İzmit (Sapanca) (İren & Ahmed 1973); Artvin province:
Ardanuç (Tepedüzü village) (Alkan & Eroğlu 2001);
Kırklareli province: Demirköy, İslambeyli, Şırnak
Callimus angulatus (Schrank 1789) province: Central (Özdikmen 2007); Manisa province:
Kırkağaç (Tezcan & Can 2009; Özdikmen 2008, 2011b;
Red List category in Europe. LC. Özdikmen et al. 2009; Özdikmen & Turgut 2009a);
Annales de la Société entomologique de France (N.S.) 7
Mersin province: 30 km NW Gülnar: Köseçobanli, Aktaş villages (Özdikmen 2006); Erzincan province:
Tasdüstü, Kizilen (Sama et al. 2011). Kemaliye; Muğla province: Marmaris and Fethiye
It is also recorded as C. cerdo acuminatus (Özdikmen 2007, 2008, 2011b; Özdikmen et al. 2009;
(Motschulsky 1852) by Turkish researchers. Distribution Özdikmen & Turgut 2009a).
of C. cerdo acuminatus (Motschulsky 1852) in Turkey:
İzmir province: Kemalpaşa (Armutlu) (Tezcan & Rejzek Global distribution. East Mediterranean: Macedonia,
2002); Hatay province: Akbez (Fairmaire 1884); Antalya Bulgaria, Ukraine, central and south-east, Turkey, Syria,
province: Aspendos (Demelt 1963); İstanbul province: Lebanon, Jordan, Israel (Sama et al. 2010), Macedonia,
Üsküdar (Adlbauer 1988); Tunceli province: (Tauzin Bulgaria, Crimea (Danilevsky 2007).
2000; Özdikmen 2008, 2011b; Özdikmen & Turgut
2009a).
Cerambyx miles Bonelli 1812
Global distribution. Europe, North Africa, Caucasus,
Turkey, Iran, Syria, Jordan, Lebanon, Israel (Sama et al. Red List category in Europe. NT.
2010); Portugal, Spain, France (without Corsica), Italy, Distribution in Turkey. Denizli province: (Gül-
Sardinia, Sicilia, Malta, Latvia Slovenia, Croatia, Bosnia Zümreoğlu 1975); Uşak province: Ulubey (Ovacık
and Herzegovina, Yugoslavia, Macedonia, Greece village) (Özdikmen & Çağlar 2004); Afyon province:
(without Crete), Bulgaria, Romania, Hungary, Austria, Akkale hill (Özdikmen 2006). İçel province: Bolkar Mts.
Switzerland, Germany, Luxembourg, Belgium,
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2004), Tarsus (Bağlarbaşı) (Tozlu et al. 2002); Osmaniye Çamlıca (Özdikmen & Demir 2006); Adıyaman province:
province: Nurdağı pass (Adlbauer 1992); Isparta province: Karadut village env. (Rejzek & Hoskovec 1999); Turkey
Eğirdir (Tauzin 2000); Adana province: Seyhan; Antalya (Sama & Rapuzzi 2000; Sama 2002; Özdikmen 2011b;
province: Manavgat, Alanya (Çayarası plateau–Sarımut Özdikmen & Turgut 2009a).
bridge); Kahramanmaraş province: Pazarcık (Botaş) It is also recorded as C. velutinus Brullé 1832 in İçel
(Özdikmen & Okutaner 2006); İstanbul province: province: Bolkar Mts. (Bodemeyer 1906); Turkey (Erdem
Bosphorus region (Bahçeköy) (Schimitschek 1944); 1968; Çanakçıoğlu 1983; Lodos 1998); İstanbul province:
Erzurum province: (Özbek 1978); Bingöl province: Polonez village, Alem Mountain (Demelt 1963), Belgrad
Central; Mardin province: Hop pass (Adlbauer 1992); forest (Öymen 1987; Özdikmen 2011a; Özdikmen &
Adıyaman province: Karadut village env. (Rejzek & Turgut 2009a).
Hoskovec 1999); Uşak province: Ulubey (Ovacık
Global distribution. Europe, southern Turkey, Near East
village, Gökgöz hill); Muğla province: Kemer (Ceylan
(Syria, Jordan, Lebanon, Israel) (Sama et al. 2010);
village); Konya province: Taşkent (İshaklı village, Gevne
Portugal, Spain, France (without Corsica), Italy (without
valley); Bitlis province: Nemrut Mountain (Özdikmen
Sardinia and Sicily), Sicilia, Malta, Slovenia, Croatia,
2007, 2008, 2011b; Özdikmen & Turgut 2009a).
Bosnia and Herzegovina, Yugoslavia, Greece, Crete,
It is also recorded as C. nodulosus nodicornis Küster
Bulgaria, Romania, Hungary, Ukraine, Crimea
1846 by Özdikmen & Turgut (2009c). Distribution of C.
(Danilevsky 2007).
nodulosus nodicornis Küster 1846 in Turkey: Asia Minor
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Slovakia, Poland, Estonia, Latvia, Lithuania, Belorussiya, province: Belen (Güzelyayla road) (Özdikmen &
Ukraine Crimea, Moldova, Russia, European Kazakhstan Demirel 2005); İçel province: exit of Atakent, Mut–
(Danilevsky 2007). Silifke road, Erdemli–Güzeloluk road Mersin–Gözne
road (Çukurkeklik), Silifke–Mut road (Göksu bridge)
(Özdikmen 2006); Kahramanmaraş province: Pazarcık
Chlorophorus sartor (Müller 1766) (Bağdınısağır), Aksu bridge, Sakarkaya village (Kısık),
Çağlayancerit (Bozlar), Kahramanmaraş–Andırın road
Red List category in Europe. LC.
(Körsülü bridge env.), Afşin (Çardak–Afşin road),
Distribution in Turkey. Muğla province: Bodrum Nurhak (Nurhak–Malatya road, Tatlar), Andırın
(Gümbet), İzmir province: Kemalpaşa, Tire, Çeşmealtı (Andırın–Çokak road, Çınar place, Parmaksız plateau);
(Gül-Zümreoğlu 1975), Selçuk (Adlbauer 1988); Manisa, Denizli province: Menderes Valley (Schimitschek 1944);
Aydın, Muğla, Denizli (Lodos 1998); Kahramanmaraş İzmir province: Torbalı (Tuatay et al. 1972), Bornova,
province: Pazarcık, Bağdınısağır, Göksun road, entry of Turgutlu (İren & Ahmed 1973), Kemalpaşa, Urla (Çıtlık
Tekir, Sakarkaya village, Andırın road, Çuhadırlı village village) Bornova, Menemen, Çeşmealtı (Gül-Zümreoğlu
(Özdikmen 2006; Özdikmen 2008; Özdikmen & Okutaner 1975); Manisa province: Demirci (Gül-Zümreoğlu 1972),
2006). Mersin province: 30 km NW Gülnar: Köseçobanli, Demirci, Turgutlu (Gül-Zümreoğlu 1975); Muğla
Tasdüstü (Sama et al. 2011). Gaziantep province: province: Köyceğiz, Fethiye (Kesikkapı) Dalaman
Fevzipaşa (Sama 1982); Osmaniye province: Nurdağı (Karaçalı), Marmaris (Gökova); Denizli province:
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pass (Adlbauer 1988), Yarpuz road forest store env., Sarayköy, Central, Çivril; Aydın province: Çine (Gül-
Zorkun road (Karacalar village), Arslantaş–Osmaniye Zümreoğlu 1975; Özdikmen 2008, 2011a, 2011b);
road (Kazmaca village), Cebel road (Çürükarmut Mersin province: Devrent (Sama et al. 2011).
plateau), Yarpuz road (Yukarı Haraz plateau), Yarpuz It is also recorded as C. varius damascenus Chevrolat
road, 8th km, Düziçi (Böcekli village) (Özdikmen et al. 1854 in Antalya province: Alanya (Demelt & Alkan 1962;
2010); Hatay province: Antakya (Lodos 1998), Hassa Demelt 1963; Turgut & Özdikmen 2010). Toprakkale
(Akbez, Geneluşağı village, Zeytinoba village, Aktepe) (Antakya road 1st km) (Özdikmen et al. 2010);
Kırıkhan (Taşoluk village), Belen (Güzelyayla road) Çağlayancerit (Ç. cerit–Düzbay road, Ç. cerit–Bozlar
(Özdikmen & Demirel 2005); Antalya province: road) (Özdikmen & Okutaner 2006) ; İzmir province:
Arapsuyu (Tozlu et al. 2002), Isparta province: Bornova (Demelt & Alkan 1962; Demelt 1963;
Yenişarbademli–Gedikli (near Beyşehir lake) (Özdikmen Özdikmen 2008).
& Çağlar 2004). Ankara province: Kızılcahamam
Global distribution. Europe, south-western Siberia
(Soğuksu National Park) (Özdikmen et al. 2005);
(Urals), Asia Minor, Caucasus, Transcaucasia, northern
Kızılcahamam, Beypazarı (Dereli) (Özdikmen 2006,
Iran, Middle East (Iraq, Jordan, Lebanon, Israel), western
2011a, 2011b; Özdikmen et al. 2009). Konya province:
part of North Africa (Egypt), Cyprus as C. varius
Taşkent–Alanya, Ilıcapınar town, Hadim–Alanya road,
damascenus Chevrolat 1854 (Sama et al. 2010).
Bozkır–Hadim road, Beyşehir: Üstünler env. (Turgut &
Özdikmen 2010).
Kavak (Hacılar pass) (Malmusi & Saltini 2005); Düzce Clytus tropicus Panzer 1795
province: Yığılca; Kastamonu province: Küre; Bolu
Red List category in Europe. LC.
province: Pazarköy env. (Özdikmen 2007; Özdikmen
et al. 2009; Özdikmen &Turgut 2009b). Distribution in Turkey. European Turkey (Althoff &
Danilevsky 1997); Turkey (Lodos 1998) (Özdikmen &
Global distribution. Portugal, Spain, France, Corsica, Italy Turgut 2009b).
(without Sardinia and Sicily), Slovenia, Croatia, Bosnia and
Herzegovina, Yugoslavia, Macedonia, Greece (without Global distribution. Spain, France (without Corsica),
Crete), Bulgaria, Turkey, Romania, Hungary, Austria, Croatia, Bosnia and Herzegovina, Yugoslavia,
Switzerland, Luxembourg, Belgium, Netherlands, Macedonia, Greece (without Crete), Bulgaria, Turkey,
Denmark, Germany, Great Britain, Czechia, Slovakia, Romania, Hungary, Austria, Germany, Czechia, Slovakia,
Poland, Norway, Sweden, Finland, Estonia, Latvia, Poland, Belorussiya, Ukraine (without Crimea), Moldova,
Lithuania Belorussiya, Ukraine, Crimea, Moldova, Russia, Middle of European Russia (Danilevsky 2007).
European Kazakhstan (Danilevsky 2007).
Deilus fugax (Olivier 1790)
Clytus rhamni Germar 1817 Red List category in Europe. LC.
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Red List category in Europe. LC. Distribution in Turkey. Antalya province: Termessos
(Adlbauer 1992); Hatay province: Kırıkhan (Malmusi &
Saltini 2005); İzmir province: Kemalpaşa (Demelt &
Distribution in Turkey. Konya province: Hadim: Korualan
Alkan 1962), Efes (Adlbauer 1992); Denizli province:
town env. (Turgut & Özdikmen 2010); Ankara province:
Pamukkale; Manisa province: Turgutlu Çardağı
Kızılcahamam (Işık Mountain, Yukarı Çanlı) (Özdikmen &
(Domunludeve valley) (Özdikmen & Demirel 2005;
Demir 2006); Kızılcahamam (south of Dam, Güvem, Yasin
Özdikmen 2008, 2011b).
village, Yukarı Çanlı), Beypazarı (Dereli village) (Özdikmen
2006); İzmir province: near Central, Kemalpaşa, Efes, Global distribution. Europe eastward to the Urals and
Bergama (Demelt & Alkan 1962); Antalya province: near Ukraine, North Africa, Asia Minor, Caucasus, Cyprus,
Central, Belkıs (Aspendos, Cumali), Antitoros Mountains Near East (Sama et al. 2010).
(Bey Mountains, Korkuteli), Alanya (Güzelbağ) and
nearby, Kemer, Arapsuyu (Tozlu et al. 2002) Alanya
(Mahmutlar), Kemer (Olimpos Mt.) (Özdikmen & Çağlar Delagrangeus angustissimus Pic 1892
2004), Akseki: Murtiçi–Güzelsu, İbradı; Isparta province:
Eğirdir and near (Demelt & Alkan 1962); İçel province: Red List category in Europe. VU.
Silifke (Tuatay et al. 1972), Anamur, Silifke (Central, Distribution in Turkey. İçel province: Namrun (Demelt
Gülnar), Kuzucubelen, Tarsus (Çamlıyayla), Kanlıdivane; 1963; Holzschuh 1975), Silifke (Olba, Gülnar) (Adlbauer
Hatay province: Amanos Mountains (near Dörtyol) 1988); Turkey (Sama & Rapuzzi 1999, 2000). Hatay
(Öymen 1987), Erzin (Tozlu et al. 2002); Osmaniye province: Akbez (Özdikmen 2011b; Özdikmen et al.
province: Nurdağı pass, Zorkun plateau road (Ürün plateau, 2012); Mersin province: Çamlıyayla /Kizilen (Sama
Olukbaşı place), Yarpuz road (Karataş place) (Özdikmen & et al. 2011, 2012).
Demirel 2005), Kahramanmaraş province: Andırın; İçel
province: Silifke (Özdikmen et al. 2005), Erdemli– Global distribution. Europe (Greece: Rodos), Turkey,
Güzeloluk, Güzeloluk, from Tarsus to Çamlıyayla, from Cyprus, Lebanon (Özdikmen 2011b).
Ortagören to Mut (Malmusi & Saltini 2005), Uzuncaburç,
Mersin–Gözne road (entry of Yeniköy) (Özdikmen 2006,
2008, 2011a, 2011b; Özdikmen & Turgut 2009b), Mersin, Ergates faber (L. 1760)
Devrent 30 km N Erdemli (Sama et al. 2011).
It is also recorded as C. rhamni temesiensis Germar Red List category in Europe. LC.
1824 in İzmir (Adlbauer 1988); Gaziantep province: Distribution in Turkey. Antalya province: Sarısu Forest,
Fevzipaşa (Sama 1982); Adana province: Kozan Kaş (Sütleğen) (Alkan 2000), Central (Tosun 1975; Alkan
(Adlbauer 1988) (Özdikmen 2008, 2011b). 2000); Mediterranean Region (Çanakçıoğlu 1983;
Çanakçıoğlu 1993); Kahramanmaraş province: Başkonuş
Global distribution. Central and southern Europe, (Kanat 1998); Kızılcahamam (Çamkoru) (Özdikmen &
Caucasus, Transcaucasia, Asia Minor, Cyprus, Syria, Şahin 2006; Turkey (Bodenheimer 1958; Acatay 1948,
Lebanon, Israel (Sama et al. 2010). 1961, 1963, 1968; Danilevsky & Miroshnikov 1985;
Annales de la Société entomologique de France (N.S.) 11
Svacha & Danilevsky 1986; Önder et al. 1987); Düzce Turkey. Distribution of G. marmottani frischi Sama
province: Çiçekli plateau; Sinop province: Ayancık (Kepez 1995: Osmaniye province: Zorkun plateau (Adlbauer
forests); Bolu province: Abant (Sakarca plateau); Bursa 1992), Zorkun plateau (Sama 1995; Tauzin 2000);
province: western Black Sea region (Defne 1954), Orhaneli Turkey (Sama 1995; Lodos 1998; Sama & Löbl 2010);
(Karıncalı forest) (Çanakçıoğlu 1956); Antalya province: south-eastern Turkey (Sama 2002; Özdikmen 2011b).
Sarısu Forest, Kaş (Sütleğen), Central (Tosun 1975);
Kocaeli province: İzmit (Işıktepe) (Öymen 1987); Global distribution. Spain, France (without Corsica),
Kastamonu province: Yaralıgöz pass as Ergates faber Italy (without Sardinia and Sicily), Poland, Switzerland,
alkani Demelt 1968 (Adlbauer 1992); Trabzon province: Bulgaria, Austria, Germany, Czechia, Russia (south and
Maçka (Meryemana, Aksu); Artvin province: Şavşat central) (Danilevsky 2007), Turkey (Özdikmen 2011b).
(Veliköy, Karagöl Forests) (Yüksel 1996); European
Turkey (Althoff & Danilevsky 1997); western Black Sea
region, Mediterranean Region; Kahramanmaraş province: Glaphyra umbellatarum (Schreber 1759)
Başkonuş (Kanat 1998); Artvin province: Şavşat, Ardanuç Red List category in Europe. LC.
(Tosunlu), Trabzon province: Maçka; Sinop province:
Ayancık, (Malmusi & Saltini 2005); Bolu province: Abant, Distribution in Turkey. İzmir province: Çamlık pass
Orhaneli, Düzce (Central); Antalya province: Central, Kaş; (Adlbauer 1992; Özdikmen 2008).
Kocaeli province: İzmit (Alkan 2000; Özdikmen 2011a,
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2011b; Özdikmen et al. 2009; Özdikmen & Turgut 2009c). Global distribution. Spain, France (without Corsica), Italy
(without Sardinia and Sicily), Sicilia, Slovenia, Croatia,
Global distribution. Europe: Portugal, Spain, France, Bosnia and Herzegovina, Yugoslavia, Macedonia, Greece
Corsica, Italy, Sicily, Albania, Slovenia, Croatia, Bosnia (without Crete), Bulgaria, Turkey, Romania, Hungary,
and Herzegovina, Serbia, Greece, Bulgaria, European Austria, Germany, Luxembourg, Belgium, Netherlands,
Turkey, Hungary, Austria, Switzerland, Netherlands, Denmark, Great Britain, Czechia, Slovakia, Norway,
Germany, Czechia, Slovakia, Poland, Sweden, Latvia, Poland, Sweden, Estonia, Lithuania, Latvia, Belorussiya,
Lithuania, Belorussia, Ukraine, Crimea, European Moldova, Ukraine, Crimea, Russia (Danilevsky 2007).
Russia, North Africa (Algeria, Morocco), Caucasus,
Transcaucasia, Near East, Turkey, Syria (Özdikmen &
Turgut 2009c). Hesperophanes sericeus (Fabricius 1787)
Red List category in Europe. LC.
Glaphyra kiesenwetteri (Mulsant & Rey 1861) Distribution in Turkey. Isparta province: (Tozlu et al.
Red List category in Europe. DD. 2002); Turkey (İyriboz 1940; Sama 2002; Özdikmen &
Şahin 2006); İzmir province: Dikili, Makaron (Demelt
Distribution in Turkey. İçel province: Erdemli, Namrun 1963); Aydın province: Kuyucak (Öymen 1987); Denizli,
(Central, Çamlıyayla) as Molorchus kiesenwetteri anatolicus Erzincan (Özdikmen 2008, 2011b).
Adlbauer 1988 (Adlbauer 1988); Antalya province: Akseki
(Yarpuz, Cevizli) as M. kiesenwetteri anatolicus Adlbauer Global distribution. Mediterranean area from North
1988 (Adlbauer 1992); İçel province: Namrun (Erdemli) as Africa (including Egypt) and Iberian Peninsula to
M. kiesenwetteri hircus (Abeille de Perin 1881) (Tauzin southern France, Caucasus, Iran, southern Turkey, Iraq,
2000). İzmir province: Kemalpaşa (Armutlu) as M. Jordan, Israel, Cyprus (Sama et al. 2010).
kiesenwetteri hircus (Abeille de Perin 1881) (Tezcan &
Rejzek 2002; Özdikmen 2008, 2011b).
Hylotrupes bajulus (L. 1758)
Global distribution. Turkey, Armenia, Azerbajdzhan,
Red List category in Europe. LC.
northern Iran, Israel, Syria, Lebanon (Sama 1995) as G.
kiesenwetteri hircus (Abeille de Perin 1881; Sama et al. Distribution in Turkey. Hatay province: Akbez (Fairmaire
2010). 1884); Antalya province: Antitoros Mountains (Demelt &
Alkan 1962), Toros Mountains, on beach, Central, Sarısu
place (Erdem & Çanakçıoğlu 1977; Çanakçıoğlu 1983),
Glaphyra marmottani Brisout de Barneville 1863 Alanya (Adlbauer 1988), Kumluca, Serik; İçel province:
Red List category in Europe. DD. Silifke (Tuatay et al. 1972; Demelt 1963), Erdemli, Tarsus
(Tozlu et al. 2002), Silifke, Fındıkpınarı (Özdikmen et al.
Distribution in Turkey. This species has a subspecies 2005); Isparta province: (Tosun 1975), Central, Senirkent,
which is named G. marmottani frischi Sama 1995 in Sütçüler (Özdikmen et al. 2005); Kahramanmaraş,
12 S.S. Avgın et al.
Mediterranean Region (Kanat 1998), Pazarcık (Sama 1982) (Lodos 1998), Marmaris (Özdikmen et al.
(Bağdınısağır) (Özdikmen & Okutaner 2006); Adana 2005), Turgutlu Çardağı (Domunludeve valley)
province: Kozan (Bağtepe), Yumurtalık (Özdikmen et al. (Özdikmen & Demirel 2005); İzmir province:
2005); Hatay province: Central; Osmaniye province: Agamemnun, Bergama (Kozak), Kemalpaşa (Gül-
Zorkun road (Fenk plateau) (Özdikmen et al. 2010); İzmir Zümreoğlu 1975), as Callimellum adonis (Sama 1982),
province (Acatay 1943): Bornova, Karşıyaka (Gül- Selçuk (Meryemana) (Malmusi & Saltini 2005); Samsun
Zümreoğlu 1972), Söke (Gül-Zümreoğlu 1975); Denizli Mt. (Adlbauer 1988); Manisa province: Akhisar; Hatay
province (Acatay 1943): Menderes valley (Schimitschek province: Akbez (Fairmaire 1884), Amanos Mts.
1944), Çamlık forest (Tosun 1975), Central (Gül- (Nurdağı pass) (Villiers 1967), İskenderun (Güzelyayla
Zümreoğlu 1975); Uşak province: (Acatay 1943), road, entry of Kurtbağı village, Kozaklı village), Hassa
Mollamusa village; Aydın province: Ortaklar (Gül- (Akbez, Zeytinoba village) (Özdikmen & Demirel 2005);
Zümreoğlu 1975; Özdikmen et al. 2005); Denizli, İzmir, İçel province: Burna, Toros Mts. (Bolkar Mts.) as
Aydın Uşak (Erdem & Çanakçıoğlu 1977; Çanakçıoğlu Callimus adonis (Sama 1982) (Villiers 1959), Alata
1983); Aegean Region (Kanat 1998). Ankara province: (Villiers 1967), Erdemli (Öymen 1987), Anamur, Silifke
(Özdikmen et al. 2005), Elmadağ (Villiers 1967; Öymen (Gülnar), Erdemli, Kuzucubelen, Tarsus (Çamlıyayla)
1987); Elmadağ, Çamlıdere (Tozlu et al. 2002); Adlbauer 1988; Antalya province: Toros Mts.
Kızılcahamam (Çileklitepe) (Özdikmen 2006, 2008, 2011a, (Bodemeyer 1900), Central as Callimellum adonis (Sama
2011b; Özdikmen et al. 2009). 1982) (Demelt & Alkan 1962; Demelt 1963), Kemer, Yeni
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and Israel, North Africa, occasionally imported in North Sweden, Estonia, Latvia, Lithuania, Belorussia, Ukraine,
America (Sama et al. 2010). Crimea, Moldavia, European Russia, European
Kazakhstan), Siberia, Caucasus, Transcaucasia, Turkey,
Iran, Syria (Özdikmen & Turgut 2009d).
Phymatodes testaceus (L. 1758)
Red List category in Europe. LC.
Poecilium alni (L. 1767)
Distribution in Turkey. İçel province: Bolkar Mts. as P. Red List category in Europe. LC.
testaceus var. analis (Bodemeyer 1906); Hatay province:
Dörtyol (Öymen 1987); Osmaniye province: Nurdağı pass Distribution in Turkey. Osmaniye province: Nurdağı
(Adlbauer 1988); Antalya province: Central (Tozlu et al. pass (Adlbauer 1988), Kuşçubeli pass (Gaziantep border)
2002); Ankara province: Kızılcahamam (Soğuksu National (Özdikmen et al. 2010; Özdikmen 2011b).
Park), Beypazarı (Dereli village) (Özdikmen 2006, 2011b;
Özdikmen et al. 2009); Isparta province: Yukarıgökdere; Global distribution. Portugal, Spain, France (without
Burdur province: Gölhisar (Sama et al 2011). Corsica), Italy, Sicilia, Slovenia, Croatia, Bosnia and
Herzegovina, Yugoslavia, Greece (without Crete)
Global distribution. Europe, North Africa, Turkey, Bulgaria, Turkey, Romania, Hungary, Austria,
Caucasus, Middle East (Syria, Israel), Japan, Siberia, Switzerland, Luxembourg, Belgium, Netherlands,
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introduced in North America (Sama et al. 2010). Denmark, Germany, Great Britain, Czechia, Slovakia,
Poland, Norway, Sweden, Finland, Estonia, Latvia,
Lithuania, Belorussiya, Ukraine, Crimea, Moldova,
Plagionotus arcuatus (L. 1758) Russia, European Kazakhstan (Danilevsky 2007).
Red List category in Europe. LC.
Distribution in Turkey. Isparta province: Eğirdir (Tuatay Poecilium lividum (Rossi 1794)
et al. 1972), Eğirdir (Özdikmen et al. 2005), Red List category in Europe. DD.
Yukarıgökdere (Sama et al. 2011); Osmaniye province:
Central (Tozlu et al. 2002; Özdikmen 2011b). Distribution in Turkey. İçel province: Erdemli (Adlbauer
1988; Özdikmen 2011b); Mersin province: 30 km NW
Global distribution. Europe (Portugal, Spain, France, Gülnar: Köseçobanli, Tasdüstü, Devrent (Sama et al. 2011).
Italy, Sicily, Sardinia, Albania, Slovenia, Croatia, Bosnia
and Herzegovina, Serbia, Greece, Crete, Bulgaria, Global distribution. Europe, North Africa, Caucasus,
European Turkey, Romania, Hungary, Austria, Transcaucasia, Turkey, Syria, Lebanon, Israel (Sama
Switzerland, Belgium, Netherlands, Denmark, Germany, et al. 2010).
Luxembourg, Great Britain, Czechia Slovakia, Norway,
Poland, Sweden, Finland, Estonia, Latvia, Lithuania,
Belorussia, Ukraine, Crimea, Moldavia, European Poecilium fasciatum (Villers 1789)
Russia, European Kazakhstan), North Africa (Algeria, Red List category in Europe. LC.
Morocco), Caucasus, Iran, Transcaucasia, Turkey, Syria
(Özdikmen & Turgut 2009d). Distribution in Turkey. P. fasciatum (Villers 1789) has
not been recorded from Turkey from any exact locality,
although Sama (2002) and Sama and Löbl (2010)
Plagionotus detritus (L. 1758) mentioned that the species occurs in southern Turkey
(Cebeci et al. 2011).
Red List category in Europe. LC.
Global distribution. Central and southern Europe,
Distribution in Turkey. Manisa province: Muradiye southern Turkey, Cyprus (Sama 2002; Sama et al. 2010).
(Tezcan & Rejzek 2002); Adana, Hatay, Antalya (Lodos
1998); Kahramanmaraş province: Central (Tozlu et al.
2002; Özdikmen 2008, 2011b). Poecilium pusillum (Fabricius 1787)
Global distribution. Europe (Portugal, Spain, France, Red List category in Europe. LC.
Corsica, Italy, Albania, Croatia, Bosnia and Herzegovina,
Serbia, Greece, Crete, Bulgaria, European Turkey, Distribution in Turkey. Istanbul and Konya provinces
Romania, Hungary, Austria, Switzerland, Belgium, (Cebeci et al. 2011), Tunceli province: 15 km N of
Netherlands, Germany, Czechia, Slovakia, Poland, Pülümür (Sama et al. 2012).
Annales de la Société entomologique de France (N.S.) 15
Global distribution. Northern and central Europe, P. Original combination. Prionus besicanus Fairmaire 1855.
pusillum barbipes (Küster 1847) known from Czeck
Republic (Moravia), Slovakia and Greece, P. pusillum Distribution in Turkey. Hatay province: Akbez as
(Villers 1789) was also recorded from NW Turkey Prionus besicanus (Pic 1897); Asia Minor as Prionus
(Rapuzzi & Sama 2009). besicanus (Semenov 1900); European Turkey and Asia
Minor (Winkler 1924–1932); Turkey (İyriboz 1938,
1940; Bodenheimer 1958; Svacha & Danilevsky 1986;
Poecilium rufipes (Fabricius 1776) Sama & Rapuzzi 2000); İstanbul province: Polonez
village, İzmir province: Dikili, Makaron (Demelt 1963),
Red List category in Europe. LC.
Ödemiş (Bozdağ), Bornova, Dikili (Gül-Zümreoğlu
Distribution in Turkey. The nominotypical subspecies is 1972), Bergama, Makaron, Urla, Bornova, Çeşme (Gül-
known from Bolu and Samsun, the subspecies syriacum Zümreoğlu 1975); Uşak, Antalya province: Kaş (Demelt
from Antalya, Hatay (type locality), Icel (Mersin), Niğde 1963), Manavgat (Demirciler village); western Anatolia
and Osmaniye (Özdikmen 2011a); Hatay province: 4 km (İren & Ahmed 1973); Denizli province: Çal, Hançalar
south of Şenköy (Sama et al. 2012). (Gül-Zümreoğlu 1975); Kütahya province: Simav;
Çanakkale province: İntepe (Sama 1982); Nevşehir
Global distribution. The nominotypical subspecies is province: Göreme as Prionus besicanus (Adlbauer
distributed from Europe to northern Turkey; P. r. 1988); European Turkey (Althoff & Danilevsky 1997);
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syriacum (Pic 1891) in known in south-eastern Turkey, Aegean Region (Lodos 1998); Adana, Antalya, İçel,
Syria, Lebanon and Israel (Sama et al. 2010). Bilecik, Burdur, Çanakkale, Erzurum, Kayseri, Kilis,
Konya, Muğla, İstanbul, İzmir, Kütahya, Nevşehir (Tozlu
et al. 2002); Burdur province: Bucak (Özdikmen & Şahin
Prinobius myardi Mulsant 1842 2006), Uludağ (Malmusi & Saltini 2005); Erzincan
Red List category in Europe. LC. province: Kemaliye; Bursa province: Çalı village
(Özdikmen 2006); Kırıkkale province: Sulakyurt
Distribution in Turkey. İzmir province: Kuşadası as (Özdere); Ankara province: Kalecik (Yeşildere)
Macrotoma scutellaris (Germar 1817) (Demelt 1963; (Özdikmen & Demir 2006; Özdikmen et al. 2009;
Öymen 1987), Bornova (Gül-Zümreoğlu 1972); Aydın Özdikmen & Turgut 2009c).
province: Kuşadası as Macrotoma scutellaris (Germar
1817) (Gül-Zümreoğlu 1972), Dilek (Alkan 2000); Muğla Global distribution. Europe (Albania, Croatia and Bosnia
province: Marmaris as Prinobius scutellaris (Germar 1817) and Herzegovina, Serbia, Macedonia, Greece, Bulgaria,
(Adlbauer 1992), Central (Tozlu et al. 2002), Marmaris, European Turkey), Cyprus, Middle East (Syria, Lebanon,
Aktur (Malmusi & Saltini 2005); Adana province: Dörtyol Jordan), Turkey (Özdikmen & Turgut 2009c).
as Macrotoma scutellaris (Germar 1817) (Bodenheimer
1958; Gül-Zümreoğlu 1975; Öymen 1987), Karataş as
Prinobius scutellaris (Germar 1817) (Adlbauer 1992; Prionus coriarius (L. 1758)
Lodos 1998); Burdur province: Bucak; Antalya province:
Red List category in Europe. LC.
Kemer, Kaş–Gürsu, Çakırlar (Ekici 1971; Öymen 1987;
Tosun 1975; Yüksel 1996; Alkan 2000), Beldibi, Distribution in Turkey. Aydın province: Dilek Peninsula
Manavgat, Serik, Alanya (Çayarası plateau–Sarımut National Forest (Öymen 1987; Çanakçıoğlu & Mol 1998);
bridge) (Özdikmen & Çağlar 2004); İçel province: Erdemli Burdur province: Bucak (Ekici 1971); Antalya province:
(Tozlu et al. 2002; Özdikmen 2008; Özdikmen 2011b); Kemer (Beldibi), Kaş (Gürsu), Çakırlar forest (Tosun
Isparta: Yukarıgökdere, Mersin: 30 km NW Gülnar: 1975; Erdem & Çanakçıoğlu 1977; Çanakçıoğlu 1983),
Köseçobanli, Tasdüstü (Sama et al. 2011). Kaş (Sinekçi village, Sinekçi Beli) (Özdikmen & Çağlar
Global distribution. Europe (Portugal, Spain, France, 2004), Akseki (Turgut & Özdikmen 2010);
Corsica, Italy, Sardinia, Sicily, Croatia, Bosnia and Kahramanmaraş province: (Kanat 1998), Pazarcık
Herzegovina, Serbia, Greece, Crete, Crimea), North (Bağdınısağır Mahallesi) (Özdikmen & Okutaner 2006);
Africa (Algeria, Morocco, Libya, Iran, Tunisia), Hatay province: Hassa (Söğütler) (Özdikmen & Demirel
Caucasus, Near East, Turkey, Syria, Lebanon, Jordan 2005); Konya province: Çayarası–Alanya (Özdikmen &
(Özdikmen & Turgut 2009c). Turgut 2010); Osmaniye province: Çiftmazı, Zorkun road
and Mitisin plateau (Özdikmen et al. 2010); Ankara
province: Kızılcahamam (Çamkoru) (Özdikmen & Şahin
Mesoprionus besicanus (Fairmaire 1855) 2006); Artvin province: Hopa, Şavşat (Yayla,
Kocabeyplace), Şavşat (Veliköy, Karagöl Forests)
Red List category in Europe. DD. (Yüksel 1996); Balıkesir province: Erdek, Kocaeli
16 S.S. Avgın et al.
province: Kerpe, İzmit (Özdikmen & Şahin 2006); Rize Central (Kavaklı), Kahramanmaraş–Andırın road (Körsülü
province: Central, Fındıklı, Pazar; Trabzon province: bridge env.), Pazarcık (Sakarkaya–Çağlayancerit road,
Yeşilova (Tozlu et al. 2002); Kırıkkale province: Sakarkaya village, Kısık), Çağlayancerit (Bozlar), Andırın
Sulakyurt (Özdere) (Özdikmen & Demir 2006; (Kahramanmaraş–Andırın road, Çuhadırlı village)
Özdikmen 2008, 2011a, 2011b; Özdikmen & Turgut (Özdikmen & Okutaner 2006); Ankara province: (Lodos
2009c; Özdikmen et al. 2009). Çanakkale province: west 1998); Kazan (Orhaniye village) (Özdikmen & Çağlar
of Küçükkuyu (Maican & Serafim 2009). 2004), (Özdikmen et al. 2005; Özdikmen 2008; Özdikmen
2011a; Özdikmen 2011b; Özdikmen et al. 2009); Antalya
Global distribution. Europe, Near East, North Africa province: Akseki–Manavgat road, Akseki–Güzelsu, İbradı;
(Maican & Serafim 2009). Alanya: Karapınar village, Karapınar–Sarımut, Akseki:
Mahmutlu village, Murtiçi–Güzelsu, Güçlüköy Bademli
beli–Tekebeli, Alanya: 6 km to Dikmetaş plateau; Konya
Procallimus egregius (Mulsant & Rey 1863) province: Beyşehir–Akseki road: Tepearası return Hadim:
Red List category in Europe. DD. Beyreli village (Turgut & Özdikmen 2010).
It is also recorded as P. budensis m. bitlisiensis ab. affinis
Distribution in Turkey. Anatolia (Winkler 1924–1932); and P. budensis productus Plavilstshikov 1940 by research-
Antalya, İçel province: Namrun (Demelt 1963), Tarsus ers (Gül-Zümreoğlu 1975; Adlbauer 1992). Distribution of
(Çamlıyayla) (Adlbauer 1988); Hatay province: Antakya P. budensis m. bitlisiensis ab. affinis in Turkey: İzmir pro-
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(İskenderun) (Villiers 1967); Turkey (Danilevsky & vince: Karaburun–Balıklıova (Gül-Zümreoğlu 1975).
Miroshnikov 1985; Lodos 1998; Sama & Löbl 2010; Distribution of P. budensis productus Plavilstshikov 1940
Özdikmen 2011b). in Turkey: Osmaniye province: Nurdağı (Adlbauer 1992).
Global distribution. Macedonia, Greece (without Crete), Global distribution. Europe, Turkey, Caucasus,
Crimea (Danilevsky 2007), Turkey (Özdikmen 2011b). Transcaucasia, Middle East, Cyprus, Syria, Lebanon,
Israel (Sama et al. 2010).
Erdemli–Güzeloluk road (Özdikmen 2007); Turkey (Lodos province: Alanya (Demelt 1963), Toros Mountains
1998; Sama & Rapuzzi 2000; Jenis 2001); Adıyaman (Elmalı) (Villiers 1967), Alanya (Erdem & Çanakçıoğlu
province: Karadut village (Rejzek & Hoskovec 1999) 1977; Çanakçıoğlu 1983; Çanakçıoğlu & Mol 1998;
Nemrut Mt.; Bingöl province (Tauzin 2000); Muş Svacha & Danilevsky 1986), Bambus Camp (Adlbauer
province: Buğlan pass (Malmusi & Saltini 2005); Aydın, 1988); Kahramanmaraş (Kanat 1998); Adana province:
Mardin (Özdikmen et al. 2005; Özdikmen 2008, 2011b). Balcalı, Karataş; Burdur province: Central; Hatay
It is also recorded as P. dalmatinus apicalis Pic 1905 province: Central, İskenderun (Cırtıman); İçel province:
in Siirt (Central and Baykan) (Fuchs & Breuning 1971; Erdemli, Tarsus, Erdemli (Karahasanlı village);
Özdikmen 2008, 2011b). Osmaniye province: Kadirli (Kabayar) (Tozlu et al.
2002), Bahçe road (Özdikmen et al. 2010; Özdikmen
Global distribution. Eastern Mediterranean from Balkans
2008; Özdikmen 2011b; Özdikmen & Turgut 2009c).
to the Near East, Syria, Jordan, Lebanon, Israel (Sama
et al. 2010). Global distribution. Balkan peninsula from Dalmatia to
the southern Greece, Bulgaria Turkey, Caucasus, Iran,
Syria, Israel, Cyprus (Heyrovský 1948; Sama 1993;
Purpuricenus (Purpuricenus) kaehleri (L. 1758) Althoff & Danilevsky 1997; Sama et al. 2010), Albania,
Red List category in Europe. LC. Yugoslavia, Macedonia (Danilevsky 2007). Europe
(Albania, Serbia, Macedonia, European Turkey),
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Distribution in Turkey. Rize province: Pazar (Sekendiz Georgia, Transcaucasia, Near East, Turkey (Özdikmen &
1976) (Alkan & Eroğlu 2001); Ordu province: Akkuş Turgut 2009c).
(Sama et al. 2011); Artvin, Istanbul, Kırıkkale, Rize,
Trabzon (Özdikmen 2007, 2008).
Rhaphuma gracilipes (Faldermann 1835)
Global distribution. Portugal, Spain, France, Corsica,
Italy, Sicilia, Slovenia, Croatia, Bosnia and Herzegovina, Red List category in Europe. DD.
Yugoslavia, Macedonia, Albania, Greece (without Crete)
Distribution in Turkey. Hatay province: Akbez
Bulgaria, European Turkey, Romania, Hungary, Austria,
(Fairmaire 1884; Özdikmen 2011b).
Belgium, Germany, Czechia, Slovakia, Poland,
Belorussiya, Ukraine, Crimea, Moldova, Russia Global distribution. Poland, Belorussiya, Russia
(southern and central region) (Danilevsky 2007). (northern and central region) (Danilevsky 2007).
Ropalopus insubricus (Germar 1824) 2011). Whereas it is widely seen in Europe, it disappeared
in some regions in Europe and it was first recorded by
Red List category in Europe. NT.
Sama et al. (2012) in Turkey.
Distribution in Turkey. Aegean region (Özdikmen 2008).
Global distribution. Spain, France (without Corsica), Saperda octopunctata (Scopoli 1772)
Italy (without Sardinia and Sicily), Slovenia, Croatia,
Bosnia and Herzegovina, Yugoslavia, Albania, Greece Red List category in Europe. LC.
(without Crete), Bulgaria, Romania, Ukraine (without
Distribution in Turkey. Trabzon (Öymen 1987; Alkan &
Crimea), Russia (central and southern region)
Eroğlu 2001).
(Danilevsky 2007).
Global distribution. Central European species reaching
the Caucasus, Russia and North Africa and Iberian
Ropalopus macropus (Germar 1824) Peninsula (Gutièrrez et al. 2004).
Red List category in Europe. LC.
Global distribution. Slovenia, Croatia, Bosnia and Distribution in Turkey. İzmir province: Efes (Demelt &
Herzegovina, Yugoslavia, Bulgaria, Romania, Hungary, Alkan 1962), Bayındır (Öymen 1987); Antalya province:
Austria, Switzerland, Germany, Czechia, Slovakia, Manavgat (Demelt 1963; Özdikmen 2008, 2011b); Artvin
Poland, Latvia, Belorussiya, Ukraine Crimea, Moldova, province: Saçinka; Trabzon province: Akçaabat (Alkan &
Russia (central and southern region) (Danilevsky 2007). Eroğlu 2001).
Distribution in Turkey. Antalya province: Elmalı et al. 2002), south Germany, Austria (Adlbauer 2001),
(Adlbauer 1992; Özdikmen 2011b). lowlands of Czech Republic, Slovakia, Hungary, Romania,
Bulgaria, Balkans and north-east Italy (Althoff & Danilevsky
Global distribution. Italy (without Sardinia and Sicily), 1997). Some locations of S. flavicornis Küster 1846 in
Croatia, Bosnia and Herzegovina, Yugoslavia, Greece Turkey and Israel were misidentified and belong to S.
(without Crete), Romania, Hungary, Bulgaria, Austria, kraatzi (Brustel et al. 2002; Zamoroka & Panin 2011).
Slovakia, Ukraine, Crimea (Danilevsky 2007).
(Malmusi & Saltini 2005); Afyon (Tozlu et al. 2002); Manisa Muradiye (Tezcan & Rejzek 2002); İçel province: Mut
province: Salihli (Özdikmen et al. 2005); Adana province: (Özdikmen & Şahin 2006); Gaziantep province
Karataş as Stromatium fulvum (Villers 1789) (Adlbauer (Özdikmen et al. 2005); Antalya province: Manavgat
1992), Central, Ceyhan (Tozlu et al. 2002); İçel, Antalya as (Central, Titreyengöl, Bucak) (Özdikmen & Demir 2006;
Stromatium fulvum (Villers 1789) (Lodos 1998), Central, Özdikmen 2008, 2011b).
Lara, Finike, Turunçova, Kumluca, Manavgat, Serik (Tozlu
et al. 2002), Ataş road (Karaduvar) (Özdikmen & Çağlar Global distribution. Circum-Mediterranean species:
2004), Manavgat (Demirciler village, Sülek village) southern Europe, North Africa, Egypt, Cyprus, southern
(Özdikmen & Demir 2006); Hatay province: Central, Turkey, Iraq, Jordan, Israel (Sama et al. 2010).
Erzin, İskenderun (Denizciler); Osmaniye province: Central
(Tozlu et al. 2002); Kahramanmaraş province: Pazarcık
(Bağdınısağır) (Özdikmen & Okutaner 2006), Pazarcık Trichoferus holosericeus (Rossi 1790)
(Bağdınısağır) (Özdikmen et al. 2010); Ankara (Özdikmen Red List category in Europe. LC.
& Şahin 2006; Özdikmen 2008, 2011b; Özdikmen et al.
2009); Mersin province: 30 km NW Gülnar: Köseçobanli, Distribution in Turkey. Aydın as cinereus Villers 1789
Tasdüstü (Sama et al. 2011). (Tuatay et al. 1972), (Özdikmen et al. 2005); İzmir
province: Bornova, Kuşadası as cinereus Villers 1789
Global distribution. Mediterranean: Europe, North (Gül-Zümreoğlu 1972; Özdikmen 2008).
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Sweden, Belorussiya, Ukraine, Crimea, Moldova, Russia Global distribution. The Osmoderma eremita (s.lat.)
(central and southern region) (Danilevsky 2007). complex includes in Europe at least two distinct clades
(Audisio et al. 2007). The first clade includes the West
European O. eremita Scopoli 1763, and the two Italian
Xylotrechus arvicola (Olivier 1795) endemic taxa O. italicum Sparacio 2000 and O. cristinae
Red List category in Europe. LC. Sparacio 1994, from southern peninsular Italy and Sicily.
O. eremita Scopoli 1763 (s.str.) distributed in western
Distribution in Turkey. Hatay province: Iskenderun Europe (northern Spain, France, Belgium, the
(Amanos Mountains) (Öymen 1987); Adana province: Netherlands, Denmark, southern Sweden, southern
Kozan (Feke) (Adlbauer 1992; Özdikmen 2011b). Norway, Italy, Switzerland, western Austria, western
Slovenia, most of western and northern Germany). The
Global distribution. Poland, Spain, France, Corsica, Italy, second clade includes the widespread East European O.
Sardinia, Sicilia, Slovenia, Croatia, Bosnia and Herzegovina, barnabita Motschulsky 1845 (most of Germany, eastern
Yugoslavia, Macedonia, Albania, Greece (without Crete), Austria, Slovenia, Czech Republic, Poland, Hungary,
Bulgaria, European Turkey, Romania, Hungary, Austria, Rumania, Bulgaria, Ukraine, Belarus, Latvia, Lithuania,
Luxembourg, Belgium, Netherlands, Germany, Czechia, Estonia, southern Finland, European Russia, Slovakia,
Slovakia, Poland, Estonia, Latvia, Lithuania, Belorussiya, Croatia, Bosnia and Herzegovina, Albania, Serbia,
Ukraine (without Crimea), Moldova, Russia, European Montenegro, Macedonia) and the southern Balcanic O.
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Kazakhstan (Danilevsky 2007). lassallei Baraud & Tauzin 1991 from Greece and
European Turkey (Audisio et al. 2007).
Xylotrechus rusticus (L. 1758) Remarks. It is very widely distributed in Europe and it is
Red List category in Europe. LC. frequently mentioned in several studies recently since it is
supposed to be an indicator species. Audisio et al. (2008)
Distribution in Turkey. İzmir province: Bergama (Dikili) stated in his study that some different European species and
(Adlbauer 1992; Özdikmen 2008). subspecies take place in O. eremita complex. This complex
is divided into two clades at least, one of them contains a
Global distribution. Poland, Spain, France (without taxon that is widespread in western Europe; the other one
Corsica), Italy (without Sicily), Sardinia, Slovenia, contains a taxon in middle and eastern Europe. These species
Croatia, Bosnia and Herzegovina, Yugoslavia, are affected by several negative environmental factors,
Macedonia, Albania, Greece (without Crete), Bulgaria,
especially destruction of their habitat, and so they have
Romania, Hungary, Austria, Denmark, Germany,
been conserved in many European countries.
Czechia, Slovakia, Poland, Estonia, Latvia, Lithuania,
Belorussiya, Ukraine, Crimea, Moldova, Russia,
European Kazakhstan (Danilevsky 2007).
Osmoderma lassallei (Baraud & Tauzin 1991)
Red List category in Europe. EN.
Family Scarabaeidae, Subfamily Cetoniinae
Distribution in Turkey. European Turkey (Audisio et al.
Gnorimus variabilis (L. 1758)
2007).
Red List category in Europe. NT.
Global distribution. Southern Balcanic (Greece and
Distribution in Turkey. No information was found about European Turkey) (Audisio et al. 2007).
locations in Turkey.
Global distribution. Endemic of the Asiatic part of Distribution in Turkey. Edremit Bay area and Ida
Turkey) (Audisio et al. 2007). Mountain of western Turkey (Ayıderesi, Yedikardeşler)
(Abacıgil et al. 2012).
Protaetia (Cetonischema) aeruginosa aeruginosa Global distribution. Albania, Austria, Belgium, Bosnia
(L. 1767) and Herzegovina, Croatia, Czech Republic, Denmark,
France, Germany, Great Britain, Greece, Hungary, Italy,
Red List category in Europe. NT. Latvia (Barševskis 2005a, 2005b), Norway, Poland, Russia,
Slovakia, Spain, Sweden, Ukraine (Pedroni & Platia 2010).
Distribution in Turkey. Aegean region (İren & Ahmed
1973; Krell et al. 2012). Remarks. This species, which was first recorded by
Abacıgil et al. (2012) from Turkey, is in the Near
Global distribution. Palaearctic (Europe, Mediterranean, Threatened category of the European Red List. According
Balkan, Croatia, Dalmatia, Poland) (Global Species 2011). to a search made in Skoklaster parks in Sweden, this species
has been identified as vulnerable (Jonsell 2004). Similar to
this study in Sweden, in another study in Cowdray Park in
Protaetia (Eupotosia) mirifica (Mulsant 1842) England, this species has been marked with red (Telfer
Red List category in Europe. VU. 2012). Considering these data, this species is among the
species that need conservation.
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Romania, Russia, Slovakia, Spain, Sweden, Switzerland, Global distribution. Europe: Azerbaijan, Armenia,
Ukraine, Iran, Turkey (Pedroni & Platia 2010). Bosnia and Herzegovina, Bulgaria, Georgia, Greece,
Russia (southern territory of European Russia), Ukraine;
Asia: Turkey (Cate 2007).
Ampedus elongatulus (Fabricius 1787)
Red List category in Europe. NT.
Ampedus (Ampedus) pomonae (Stephens 1830)
Distribution in Turkey. İzmir province: Kemalpaşa,
Red List category in Europe. LC.
Kemalpaşa–Kurudere, Ödemiş–Bozdağ (Gülperçin &
Tezcan 2009). Distribution in Turkey. Yozgat province: Çekerek
(Kabalak & Sert 2011).
Global distribution. Western, southern, central and eastern
Europe, northern to Latvia (Barševskis 2005a, 2005b), Global distribution. Azerbaijan, Armenia, Austria,
Estonia (Silfverberg 2010), Siberia (Gülperçin 2006). Belgium, Bosnia and Herzegovina, Bulgaria, Belarus,
Croatia, Czech Republic, Denmark, Estonia, Finland,
France, Germany, Georgia, Hungary, Ireland, Italy, Latvia
Ampedus (Ampedus) nigerrimus (Lacordaire in (Barševskis 2005a, 2005b), Luxembourg, Moldova,
Boisduval & Lacordaire 1835) Netherlands, Norway, Poland, Portugal, Romania, Russia,
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Red List category in Europe. NT. Slovakia, Slovenia, Spain, Sweden, Switzerland, Ukraine,
Serbia and Montenegro, Iran, Kazakhstan, Mongolia,
Distribution in Turkey. It is stated by Mertlik & Platia Turkey, Siberia (Pedroni & Platia 2010).
(2008) and CesaNews (centre for entomological studies
Ankara) that this species lives in Turkey but its location is
not known (Mertlik & Platia 2008; Cate 2007; Gülperçin Ampedus pomorum (Herbst 1784)
& Tezcan 2009).
Red List category in Europe. LC.
Global distribution. Europe: Austria, Belgium, Bosnia
and Herzegovina, Bulgaria, Croatia, Russia (central and Distribution in Turkey. Ankara province: Çubuk as A.
southern territories of European Russia), Czech Republic, pomorum pomorum (Herbst 1784) (Kabalak & Sert 2005).
Denmark, France, Great Britain, Germany, Greece,
Global distribution. Azerbaijan, Albania, Armenia,
Hungary, Italy, Moldova, Netherlands, Poland, Romania,
Austria, Belgium, Bosnia and Herzegovina, Bulgaria,
Slovakia, Slovenia, Spain, Sweden, Switzerland, Ukraine,
Belarus, Croatia, Czech Republic, Denmark, Estonia,
“Caucasia” Asia: Turkey (Cate 2007).
Finland, France, Great Britain, Germany, Georgia, Greece,
Hungary, Italy, Latvia (Barševskis 2005a, 2005b),
Luxembourg, Moldova, Netherlands, Norway, Poland,
Ampedus nigroflavus (Goeze 1777)
Romania, Russia, Slovakia, Slovenia, Spain, Sweden,
Red List category in Europe. LC. Switzerland, Ukraine, Serbia, Montenegro, Kazakhstan,
Mongolia, Siberia, China (Pedroni & Platia 2010).
Distribution in Turkey. Yozgat province: Şefaatli
(Kabalak & Sert 2011).
Global distribution. Albania, Austria, Belgium, Bulgaria, Ampedus praeustus (Fabricius 1792)
Belarus, Czech Republic, Denmark, Estonia, Finland, Red List category in Europe. LC.
France, Germany, Greece, Hungary, Italy, Latvia
(Barševskis 2005a, 2005b), Luxembourg, Moldova, Distribution in Turkey. Sinop province: Boyabat
Netherlands, Norway, Poland, Romania, Russia, (Kabalak & Sert 2011).
Slovakia, Slovenia, Spain, Sweden, Switzerland, Ukraine
(Pedroni & Platia 2010). Global distribution. Azerbaijan, Armenia, Austria,
Belgium, Bosnia and Herzegovina, Bulgaria, Belarus,
Croatia, Czech Republic, Denmark, Estonia, Finland,
Ampedus (Ampedus) ochropterus (Germar 1844) France, Germany, Greece, Hungary, Ireland, Italy, Latvia
Red List category in Europe. LC. (Barševskis 2005a, 2005b), Luxembourg, Moldova,
Netherlands, Norway, Poland, Portugal, Romania,
Distribution in Turkey. Çankırı province: Ilgaz (Kabalak Russia, Slovakia, Slovenia, Spain, Sweden, Switzerland,
& Sert 2011); Artvin province: Atilla valley, Genya Ukraine, Serbia and Montenegro, Kazakhstan, Turkey,
Mountain; Kars province: Sarıkamış (Kesdek et al. 2006). Morocco (Pedroni & Platia 2010).
24 S.S. Avgın et al.
Distribution in Turkey. Bursa province: Uludağ; Antalya Distribution in Turkey. Artvin province: Atilla valley
province: Termessos (Mertlik & Platia 2008) (Kesdek et al. 2006).
Global distribution. Azerbaijan, Armenia, Austria, Global distribution. Azerbaijan, Armenia, Austria,
Bosnia and Herzegovina, Bulgaria, Croatia, Czech Bosnia and Herzegovina, Bulgaria, Croatia, Czech
Republic, Denmark, France, Great Britain, Germany, Republic, France, Germany, Georgia, Greece, Hungary,
Georgia, Greece, Hungary, Italy, Moldavia, Poland, Italy, Moldova, Poland, Romania, Russia, Slovakia,
Portugal, Romania, Slovakia, Slovenia, Spain, Russia: Slovenia, Switzerland, Ukraine, Serbia and Montenegro,
southern European territory. Sweden, Ukraine, Serbia Iran, Turkey (Pedroni & Platia 2010).
and Montenegro, Algeria, Morocco, Iran (Cate 2007;
Mertlik & Platia 2008).
Brachygonus gratiosus Platia & Schimmel 1991
Red List category in Europe. DD.
Ampedus sanguineus (L. 1758)
Distribution in Turkey. Mersin, Erdemli (Platia &
Red List category in Europe. LC.
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Schimmel 1991).
Distribution in Turkey. Kars province: Sarıkamış (Tozlu
Global distribution. Europe: Greece; Asia: Turkey (Cate
2001; Tozlu et al. 2010).
2007).
Global distribution. Azerbaijan, Albania, Austria,
Belgium, Bosnia and Herzegovina, Bulgaria, Belarus,
Croatia, Czech Republic, Denmark, Estonia, Finland, Brachygonus megerlei (Lacordaire 1835)
France, Germany, Georgia, Greece, Hungary, Italy, Red List category in Europe. NT.
Latvia (Barševskis 2005a, 2005b), Liechtenstein,
Luxembourg, Moldova, Netherlands, Norway, Portugal, Distribution in Turkey. It is stated by Mertlik (2008a)
Romania, Russia Leone, Slovakia, Slovenia, Spain, and Pedroni & Platia (2010) that this species lives in
Sweden, Switzerland, Ukraine, Iran, Kazakhstan, Turkey but its location is not known.
Mongolia, Turkey, eastern region, Siberia, Morocco,
Global distribution. Austria, Belgium, Bulgaria, Croatia,
Algeria (Pedroni & Platia 2010).
Czech Republic, France, Germany, Greece, Hungary, Italy,
Poland, Romania, Russia, Slovakia, Spain, Switzerland,
Ukraine, Turkey (Pedroni & Platia 2010).
Ampedus sanguinolentus (Schrank 1776)
Red List category in Europe. LC.
Calais parreysii (Steven 1830)
Distribution in Turkey. Aksaray province: Ortaköy;
Red List category in Europe. NT.
Eskişehir province: Mahmudiye; Karaman province:
Central County; Kayseri province: Tomarza, Özvatan;
Distribution in Turkey. İzmir province: Balçova,
Konya: Ilgın, Sarayönü; Niğde province: Çamardı; Sivas
Balçova–Teleferik, Bornova–Çiçekli, Kemalpaşa–Çambel,
province: Central County, Yıldızeli; Yozgat province:
Kemalpaşa–Kurudere, first record for İzmir fauna
Şefaatli, Çandır (Kabalak & Sert 2011); Ankara
(Gülperçin & Tezcan 2009).
province: Elmadağ (Kabalak & Sert 2005).
Global distribution. Southern Russia, Iran, Turkey, Syria,
Global distribution. Azerbaijan, Albania, Armenia, Ukraine, Greece and Turkey (Gülperçin 2006).
Austria, Belgium, Bosnia and Herzegovina, Bulgaria,
Belarus, Croatia, Czech Republic, Denmark, Estonia,
Finland, Great Britain, Germany, Georgia, Greece, Hungary, Cardiophorus gramineus (Scopoli 1763)
Italy, Latvia (Barševskis 2005a, 2005b), Liechtenstein,
Luxembourg, Macedonia, Moldova, Netherlands, Norway, Red List category in Europe. NT.
Poland, Portugal, Romania, Russia Leone, Slovakia,
Distribution in Turkey. It is stated by Mertlik & Platia
Slovenia, Spain, Sweden, Switzerland, Ukraine, Serbia and
(2008) and Pedroni & Platia (2010) that this species lives
Montenegro, Iran, Kazakhstan, Mongolia, Turkey, Siberia,
in Turkey but its location is not known.
China, Morocco (Pedroni & Platia 2010).
Annales de la Société entomologique de France (N.S.) 25
Global distribution. Azerbaijan, Armenia, Austria, Denticollis rubens Piller & Mitterpacher 1783
Belgium, Bosnia and Herzegovina, Bulgaria, Croatia, Red List category in Europe. LC.
Czech Republic, France, Germany, Georgia, Greece,
Hungary, Italy, Latvia (Barševskis 2005a), Luxembourg, Distribution in Turkey. Artvin province: Cankurtaran
Moldova, Netherlands, Poland, Portugal, Romania, pass (Hopa) (Mertlik & Platia 2008).
Russia, Slovakia, Slovenia, Spain, Sweden, Switzerland,
Ukraine, Serbia–Montenegro, Israel, Turkey (Pedroni & Global distribution. Austria, Belgium, Bosnia and
Platia 2010). Herzegovina, Bulgaria, Belarus, Croatia, Russia: central
European territory, Czech Republic, Denmark, Estonia,
France, Germany, Georgia, Hungary, Italy, Latvia
Cardiophorus ruficollis (L. 1758) (Barševskis 2005a), Liechtenstein, Luxembourg, Moldavia,
Norway, Poland, Romania, Serbia and Montenegro,
Red List category in Europe. LC. Slovakia, Slovenia, Russia: southern European territory,
Sweden, Switzerland, Ukraine (Cate 2007), Spain (Molino-
Distribution in Turkey. İzmir province: Ödemiş–Bozdağ
Olmedo 2005; Mertlik & Platia 2008).
(Gülperçin & Tezcan 2009)
Haterumelater fulvago (Marseul 1868) Sweden, Switzerland, Ukraine, Iran, Japan, Syria (Cate
2007) Turkey (Platia 1994; Mertlik & Platia 2008).
Red List category in Europe. DD.
Distribution in Turkey. Mersin province: Kızılen Isidus moreli Mulsant & Rye 1874
(Jansson & Coşkun 2008; Winkler 1924–1932;
Schenkling 1925, Guglielmi & Platia 1985; Preiss & Red List category in Europe. NT.
Platia 2003).
Distribution in Turkey. It is stated by Mertlik (2008a),
Global distribution. Europe: Bulgaria, Greece, Italy; Cate (2007), Gülperçin & Tezcan (2009), Preiss & Platia
Asia: Cyprus, Israel, Lebanon, Turkey (Cate 2007). (2003) and Winkler (1924–1932) that this species lives in
Turkey but its location is not known.
Haterumelater languidus (Buysson 1891) Global distribution. Europe: France, Italy, Macedonia,
Spain, Russia (southern territories of European Russia);
Red List category in Europe. DD. North Africa: Algeria, Egypt, Morocco; Asia: Cyprus,
Turkey (Cate 2007).
Distribution in Turkey. It is stated by Mertlik & Platia
(2008), Winkler (1924–1932), Schenkling (1925) and
Preiss & Platia (2003) that this species lives in Turkey Lacon gillerforsi Platia & Schimmel 1994
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Global distribution. Austria, Bulgaria, Croatia, Russia Distribution in Turkey. Mersin province: Gülnar (Sama
(northern, central and southern territories of European et al. 2010).
Russia), Czech Republic, Denmark, Finland, France,
Germany, Georgia, Greece, Italy, Latvia (Barševskis Global distribution. Azerbaijan, Albania, Armenia,
2005a), Poland, Romania, Slovakia, Slovenia, Sweden, Austria, Bosnia and Herzegovina, Bulgaria, Belarus,
Switzerland, Ukraine Asia: Iran, Russia (Western Siberia Czech Republic, France, Great Britain, Germany,
of Russia) (Cate 2007). Georgia, Greece, Hungary, Italy, Macedonia, Moldova,
Romania, Russia Leone, Slovakia, Spain, Switzerland,
Ukraine, Serbia and Montenegro, Algeria, Turkey
Lacon punctatus (Herbst 1779) (Pedroni & Platia 2010).
Red List category in Europe. LC.
Global distribution. Azerbaijan, Albania, Armenia, Global distribution. North America, Sweden, Azerbaijan,
Bosnia and Herzegovina, Bulgaria, Croatia, Czech Bulgaria, Czeck Republic, China, Georgia, Caucasus,
Republic, France, Germany, Georgia, Greece, Hungary, Canada, Latvia (Barševskis 2005a), Mexico, Pakistan,
Italy, Malta, Portugal, Romania, Russia, Slovakia, Siberian, Slovakia and Turkey (Winkler 1924–1932;
Slovenia, Spain, Ukraine, Serbia and Montenegro, Schenkling 1925, 1927; Mardzhanyan 1976; Tarnawski
Cyprus, Jordan, Syria, Turkey, North Africa (Morocco) 1984; Guglielmi & Platia 1985; Penev & Tarnawski
(Pedroni & Platia 2010). 1987; Laibner 1988, 2000; Gülperçin 2006).
Poland, Romania, Slovakia, Spain, Switzerland, Ukraine, Stenagostus zuercheri (Reitter 1905)
Serbia and Montenegro, Turkey (Pedroni & Platia 2010).
Red List category in Europe. DD.
Procraerus carinifrons (Desbrochers des Loges 1875) Distribution in Turkey. Adana (Kesdek et al. 2006);
Mersin (Platia & Gudenzi 1998).
Red List category in Europe. DD.
Global distribution. Europe: Greece; Asia: Turkey, Syria
Distribution in Turkey. Antalya province: Gazipaşa; (Cate 2007).
Artvin province: Yusufeli, Altıparmak Mountain,
Erzurum province: Oltu, Pasinler; Konya province: south
of city, Gürağaç (Kesdek et al. 2006). Tetrigus cyprius Baudi 1871
Global distribution. Azerbaijan, Armenia, Bulgaria, Red List category in Europe. EN.
Georgia, Russia (central, southern territories of European
Russia), Ukraine, Turkey (Cate 2007). Distribution in Turkey. Isparta province: Davraz Dağı,
Yukarigökdere; İçel province: Gülnar env. (Mertlik &
Dusánek 2006).
Procraerus tibialis (Lacordaire 1835)
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Red List category in Europe. LC. Global distribution. Greece, Cyprus, Turkey, Lebanon,
Syria (Cate 2007), Israel (Platia 2010).
Global distribution. Edirne (Mertlik & Platia 2008).
Stenagostus rhombeus (Olivier 1790) Global distribution. Algeria, Austria, Bosnia and
Herzegovina, Croatia, Czech Republic, France, Germany,
Red List category in Europe. LC. Hungary, Italy, Poland, Romania, Russia: central European
Distribution in Turkey. Samsun (Kabalak & Sert 2011). territory, Slovakia, Sweden (Mouna 2007); Caucasus
(Burakowski 1991; Lucht & Merkl 1993); Turkey and
Global distribution. Austria, Belgium, Bosnia and Ukraine (Mertlik et al. 2007); Georgia (Mertlik et al. 2009).
Herzegovina, Bulgaria, Belarus, Croatia, Czech Republic,
Denmark, France, Germany, Georgia, Greece, Hungary,
Italy, Liechtenstein, Moldova, Netherlands, Poland, Portugal, Dromaeolus maronita (Bonvouloir 1871)
Romania, Russia, Slovakia, Slovenia, Spain, Sweden,
Red List category in Europe. DD.
Switzerland, Ukraine, Turkey (Pedroni & Platia 2010).
Annales de la Société entomologique de France (N.S.) 29
Distribution in Turkey. It is stated by Mertlik & Platia Global distribution. Croatia, Bosnia, Slovenia, Romania,
(2008), Mertlik (2007) that this species lives in Turkey but Bulgaria, southern Russia, Turkey and countries in the
its location is not known. Caucasus region (Mouna 1995, 2007; Lucht 1996) and
Iran (Van Meer 1999; Irurzun 2008).
Global distribution. Russia: central European territory,
Caucasus (Muona 2007); Turkey: Taurus (Reitter 1921),
Greece (Mertlik 2007). Melasis buprestoides (L. 1761)
Red List category in Europe. LC.
Epiphanis cornutus (Eschscholtz 1829)
Distribution in Turkey. It is stated by Mertlik (2008a,
Red List category in Europe. NT. 2008b) that this species lives in Turkey but its location is
not known.
Distribution in Turkey. Rize province: Ovitdaği geçidi,
between Dereköy and Sivrikaya villages, Hotel Çamlık Global distribution. Algiers, England, Belgium, Bosnia
env. (Mertlik 2007). and Herzegovina, Belarus, Bulgaria, Czech Republic,
Global distribution. Algeria, Bulgaria, France, China, Denmark, Finland, France, Croatia, Ireland, Italy,
Russia, Japan, Nearctic region (Mouna 2007), Turkey Luxembourg, Macedonia, Hungary, Germany, Norway,
(Mertlik 2007). Poland, Romania, Russia (central European and southern
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Distribution in Turkey. Mersin province: Derbent European Red List of Saproxylic Beetles.
(Jansson & Coşkun 2008).
Distribution in Turkey. Asiatic part of Turkey.
Global distribution. Greece, Araxestal, Syria, Turkey
(Mitter & Steyr 1987). Global distribution. Endemic to the Asiatic part of
Turkey. Lucanus laticornis Deyrolle has been known for
a long time but is not mentioned in most recent
Lucanus cervus cervus (L. 1758) publications or placed as a synonym or subspecies of
Lucanus cervus L. Since there are significant differences
Red List category in Europe. NT. between Lucanus laticornis Deyrolle and Lucanus cervus
Distribution in Turkey. Kahramanmaraş province: L. in external features as well in ecology and distribution,
Andırın (Kanat & Akbulut 2005); Hatay province: Lucanus laticornis Deyrolle should have the rank of a
Yayladağı, Hassa Antakya, Dörtyol; Adana province: species (Schenk 2012).
Kozan, Feke, Saimbeyli; Osmaniye province: Yarpuz
(Atay & Oğur 2011); Isparta province: Sütçüler (Karaca
et al. 2006). It is also recorded as Lucanus cervus Family Mycetophagidae
akbesianus Planet 1896 in Kahramanmaraş–Göksun; Mycetophagus (Parilendus) quadriguttatus
Hatay–Yayladağı; Adana–Feke; as Lucanus cervus (Müller 1821)
judaicus Planet 1902 Kahramanmaraş–Göksun (Atay &
Oğur 2011) and as Lucanus cervus turcicus (Sturm Red List category in Europe. LC.
1843) in Turkey (Harvey et al. 2011). Distribution in Turkey. Mersin province: Derbent and
Kızılen (Jansson & Coşkun 2008)
Global distribution. Europe, Central Asia, Crimea,
Caucasus and North Africa (Maican & Munteanu 2008). Global distribution. Europe, having been reported
throughout the continent except for Corsica, Crete,
Remarks. This species is easily noticed with its
Cyprus, Estonia, Ireland, Norway, Portugal, and Sicily
mandibula and it is rare in Europe. In the study of
(Nikitsky 2010), and is also found across North Africa,
Hawes (2008) in England, this species was considered to
in the eastern Palaearctic, Asia, Australia (Nikitsky 2010;
need conservation. Fremlin (2009) observed behaviours of
Majka 2010).
this species in urban parks and gardens in Colchester, UK.
He stated that human activities affect their behaviour and
threaten their population. Studies near Kahramanmaraş in
Mycetophagus (Mycetophagus) quadripustulatus
Turkey give the same information about its population.
(L. 1761)
Atay & Oğur (2011) carried out field work in 2009–2010
and identified a 30% decline in its population. Whereas Red List category in Europe. LC.
they claim that this is caused by negative environmental
conditions, there has not yet been a serious study on this Distribution in Turkey. Mersin province: Kızılen
issue. (Jansson & Coşkun 2008).
Annales de la Société entomologique de France (N.S.) 31
Global distribution. Albania, Austria, Belgium, Bosnia oaks from different regions in Turkey. For instance,
and Herzegovina, Britain, Corsica, Croatia, Czech Ectamenogonus montandoni (Buysson 1888), Ischnodes
Republic, Danish mainland, Estonia, European Turkey, sanguinicollis (Panzer 1793) and Megapenthes lugens
Finland, French, Germany, Greek mainland, Hungary, (Redtenbacher 1842) are found in the Kasnak Forests in
Italian mainland, Latvia, Lithuania, Macedonia, Turkey.
Moldova, Poland, Romania, Russia, Sicily, Slovakia, According to this study, Osmoderma lassallei (Baraud
Slovenia, Spanish, Sweden, Switzerland, Ukraine, & Tauzin 1991) is recorded in European Turkey but we
Yugoslavia, Near East, east Palaearctic (Silfverberg estimate that this species may also occupy in different
2010; Fauna Europaea 2012). locations in Turkey. Moreover, Callergates gaillardoti
(Chevrolat 1854) is recorded from more than two loca-
tions in Turkey (Figure 2). Although the population den-
sity is not known for these species, it could be inferred
Discussion that these are rare in Turkey.
The countries in Europe and EU countries have signed in Locations where Xylomedes cornifrons (Baudi 1874)
several agreements that aim conservation of biodiversity. can be found are not known in Turkey but Proetia mirifica
The 1979 Bern Convention on the Conservation of (Mulsant 1842) and Ischnodes sanguinicollis (Panzer
European Wildlife and Natural Habitats and the 1992 1793) are recorded from more than one location in
Convention on Biological Diversity are most important. Turkey (Figure 3). In particular, Stenopterus atricornis
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The Bern Convention mentions saproxylic beetles in five Pic 1891 is found in several regions with different climatic
lists (Nieto & Alexander 2010). The first Symposium and features in Turkey.
Workshop on Saproxylic Beetles in Hollow Trees was Although Megapenthes lugens (Redtenbacher 1842)
arranged in Sweden in 1999. It was followed by other and Epiphanis cornutus (Eschscholtz 1829) are recorded
symposia in UK (2002), Latvia (2004), France (2006), in only one location in Turkey, some saproxylic species
Germany (2008), Slovenia (2010) and Spain (2012). In such as Cerambyx cerdo L. 1758, Cerambyx dux
these works, different conservation methods were devel- (Faldermann 1837) and Cerambyx miles Bonelli 1812
oped in addition to identifying key habitats. Ranius et al. are widespread in Turkey (Figures 4, 5).
(2009) prepared some boxes mimicking tree hollows and Approximately 35% of the beetle species assessed by
located 48 of them in Brokind, Bjӓrka Sӓby and Grebo the European Red List are present in Turkey. Some species
regions in Sweden. He tried to make life-like tree hollows have limited distribution in Turkey. Although a lot of the
by putting some materials in them. The goal of his work literature has been checked, there is not enough knowl-
was to develop new methods for providing artificial habi- edge about the population density of these species from
tats to saproxylic beetle species associated with tree hol- localities in Turkey. Data show that threatened categories
lows. It worked mostly but some species, such as Allecula of saproxylic beetles can show similarities with the
morio (Fabricius 1787), Ampedus cardinalis (Schiödte European Red List for some species, although it is uncer-
1865), Chrytohagus quercinus Kraatz 1852, Procraerus tain. The European Red List of Saproxylic Beetles pro-
tibialis (Lacordaire 1835) and Osmoderma eremita vides information about the general threat status, but the
(Scopoli 1763), did not settle on these artificial habitats. regional threat status of these beetles is very different.
The studies show that the importance of saproxylic beetle Some studies show that the threat categories of these
species for the biodiversity of Europe is understood. beetles can change because of geographic range and popu-
Turkey is located at the intersection of Europe, the lation density. For example, Elater ferrugineus (L. 1758),
Middle East, Central Asia and Africa. Turkey is covered which is listed as Near Threatened in the European Red
by parts of biodiversity hotspots: Caucasus, Irano- List, is classified as Critically Endangered (CR) in
Anatolian, and Mediterranean. Furthermore, Turkey's geo- Norway. Similarly, Melasis buprestoides (L. 1761) and
graphical features have resulted in high terrestrial, fresh- Procraerus tibialis (Lacordaire 1835) are classified differ-
water, and marine biodiversity (Şekercioğlu et al. 2011). ently in Norway to the European Red List (Sverdrup-
When the biodiversity of Turkey is taken into account, the Thygeson et al. 2012). In Italy, 59% of the beetle species
number of saproxylic beetle species should be higher than in the European Red List are present and the percentage of
the number of indicated species in this study. During this threat categories are indicated differently for Italy’s
study a few important works about these species in Turkey saproxylic beetles (IUCN 2012). As a result, the threat
were identified. In one, Atalay & Oğur (2011) investigated categories of saproxylic beetles in the European Red List
the distribution of stag beetle populations in the Taurus may be unsuitable for the status of these species in Turkey.
and Amanos mountain. They detected the effect of chan- Research should be done on saproxylic beetle species in
ging environment conditions on this population. Avcı et al Turkey to determine their actual Red List threatened cate-
(2010) recorded saproxylic beetles living on Turkish old gories and criteria as published by IUCN (2012).
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