Jur. Biologi, FMIPA, Univ. Jember November 2011 Evolusi = evolve proses perubahan Perubahan genetik yg tjd pada populasi dan berlangsung dr generasi ke generasi (dalam jangka waktu yg lama) Dpt menjelaskan kesatuan (unity) dan keanekaragaman makhluk hidup Mikroevolusi : perubahan frekuensi alel pd gene pool dr generasi ke generasi Makroevolusi : pembentukan species baru (spesiasi) EXIT BAGAIMANA PERSEPSIMU TENTANG GAMBAR- GAMBAR INI? DAN APA PENDAPATMU TENTANG GAMBAR INI? EXIT How Do We Know That Evolution Has Occurred?
The evidence for evolution has primarily come from four sources:
1. the fossil record of change in earlier species 2. the chemical and anatomical similarities of related life forms 3. the geographic distribution of related species 4. the recorded genetic changes in living organisms over many generations
Geological strata containing an evolutionary sequence of fossils The Fossil Record Archaeopteryx fossil Archaeopteryx recreation Archaeopteryx tail feathers section of a DNA molecule Amino acid simple protein molecule Chemical Evidence for Evolution Summary of Evolution of Life Formation of the earths early crust and atmosphere Small organic molecules form in the seas Large organic molecules (biopolymers) form in the seas First protocells form in the seas Single-cell prokaryotes form in the seas Single-cell eukaryotes form in the seas Variety of multicellular organisms form, first in the seas and later on land Chemical Evolution (1 billion years) Biological Evolution (3.7 billion years) Anatomical Evidence for Evolution Homologous structures: structures with different appearances and functions that all derived from the same body part in a common ancestor The bones in the forelimb of mammals are homologous structures Different functions, same ancestor structure Homology of the bones of the forelimb of mammals Anatomical Evidence for Evolution Embryology Geographic distribution of some of the major colour pattern forms of Heliconius melpomene. Names of various geographical races/subspecies are given on the figure. Chris Jiggins. the geographic distribution Garis Webber Dan Wallance
Di wilayah ini dijumpai tumbuhan anggota suku Raflesiaceae, anggota famili Dipterocarpaceae misalnya Keruing ( Dipterocarpus sp), Meranti (Shorea sp), Kayu garu (Gonystylus bancanus), dan Kayu kapur (Drybalanops aromatica). Wilayah Indonesia Bagian Barat Shorea leprosula
Shorea resinosa Raflesia arnoldi Raflesia padma Wilayah Indonesia Bagian Timur
Flora yang dapat ditemukan : Sagu, Rotan, matoa, Eboni, Meranti Eucalyptus (mirip dengan di Australia) Hewan-hewan di Indonesia memiliki tipe Oriental (Kawasan Barat Indonesia) dan Australia (Kawasan Timur Indonesia) serta peralihan. ciri-ciri hewan di Kawasan Barat Indonesia :
1.Banyak species mamalia yang berukuran besar, misalnya gajah, banteng, harimau, badak. Mamalia berkantung jumlahnya sedikit, bahkan hampir tidak ada. 2.Terdapat berbagai macam kera, misalnya: bekantan, tarsius, orang utan. 3.Terdapat hewan endemik, seperti: badak bercula satu, binturong (Aretictis binturang), monyet (Presbytis thomari), tarsius (Tarsius bancanus), kukang (Nyeticebus coucang). 4.Burung-burung memiliki warna bulu yang kurang menarik, tetapi dapat berkicau. Burung-burung yang endemik, misalnya: jalak bali (Leucopsar nothschili), elang jawa, murai mengkilat (Myophoneus melurunus), elang putih (Mycrohyerax latifrons).
Kawasan Indonesia Timur Ciri-ciri hewannya adalah:
1.Mamalia berukuran kecil 2.Banyak hewan berkantung 3.Tidak terdapat species kera 4.Jenis-jenis burung memiliki warna yang beragam
Jenis-jenis hewannya seperti kanguru (Dendrolagus ursinus), kuskus (Spiloeus maculatus). burung Cenderawasih (Paradiseae sp). komodo (Varanus komodoensis). Daerah peralihan Jenis hewannya antara lain tarsius (Tarsius bancanus), maleo (Macrocephalon maleo), anoa, dan babi rusa (Babyrousa babyrussa). Genetic Drift Seleksi alam yg mengakibatkan frekuensi gen mengalami perubahan krn ada kesempatan Gene Flow Perubahan frekuensi gen krn adanya migrasi individual Mutation Peristiwa yg tjd scr acak Efek: menguntungkan, merugikan atau tdk berpengaruh. Seleksi alam Tekanan seleksi yg bekerja pd populasi dgn variasi genetik
Life Sciences-HHMI Outreach. Copyright 2006 President and Fellows of Harvard College. ALL IMAGES: http://evolution.berkeley.edu/evolibrary/home.php Map 17.1 POPULATIONS evolution equilibrium forces Phenotypic variation Random Repro- duction Large Popu- lation No net migration Gene flow chance Natural selection adaptive Sexual Recombi- nation Meiotic Crossing over mutation Random Mating/ Offspring success
Genetic drift Direc- tional Disrup- tive Stabil- izing fitness determines If it enhances undergo Never in complete requires No requires With act on include from May be or May be includes Or may get Or may get includes Or may get Variation in Populations Hukum Hardy- Weinberg : frek alel dlm populasi tetap populasi tdk mengalami evolusi Syarat: Perkawinan acak rarely takes place, even in animals Ukuran populasi sgt besar mengurani genetic drift Tdk ada migrasi Tdk tjd mutasi Tdk tjd seleksi alam Mekanisme yg menyebabkan perubahan pd populasi populasi tsb dpt survive dan bereproduksi mewariskan perubahan tsb pd generasi berikutnya Seleksi alam bekerja pd individu yg berubah adlh populasi Terdapat 3 tipe seleksi alam: 1. Directional 2. Disruptive 3. Stabilizing
Directional selection Industrial melanism in Biston betularia Disruptive selection Distinct subpopulations Stabilizing selection Keeps phenotypes stable for long periods of time. Two models exist to describe the rates of speciation. Gradualism: Refers to the model which holds that evolutionary change occurs in a gradual linear fashion. A species will change its morphology over time. Punctuated equilibrium Describes the process in which a species remains relatively unchanged for relatively long periods of time. Following which, a period of stress causes rapid change, followed by another period of relative stability. Allopatric Geographic isolation most common type Reproductive barriers/isolation Adaptive radiation
Sympatric Almost always in plants Without geographic isolation In same population but do not interbreed The notion of evolution was dominated by 2 theories. Lamarckian Theory of Inheretance of acquired characteristics Held that traits acquired throughout life of an organism would be passed on to offspring. "Use it, or lose it principle" Eg. If you lifted weights during your life, your children would be stronger or, if you did'nt exercise your children would be weaker Theory has been replaced by Darwin's theory of natural selection. Natural selection: Based on following premises: There are not enough resources for all organisms to survive. Some organisms are more "fit" to survive. Fitness refers to: The ability of an organism to survive in its environment. The ability of an organism to survive long enough to reproduce. Natural pressures "weed-out" those individuals "less fit" to survive. Thus changing a species over time. New genes/traits arise from random mutations. Most are either Neutral (no effect), or disadvantageous (decrease an organism's chance for survival). The peppered moth is a species of moth which uses camouflage against tree bark to avoid being eaten by birds. It is normally a light grey color. During England's industrial revolution, pollution caused a darkening of the bark of the trees. There was a natural variation in moth coloration with some being darker than others. Those moths of a lighter color, were easier to spot and eat by birds. And were eaten at a faster rate. The survivors tended to be darker, and when they mated, their offspring tended thus to be darker also. Over time, this resulted in the species becoming darker in color.