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and Function
Topic Outline
Sclerenchyma Ground
Ground tissue
(fiber cells) tissue
connecting
pith to cortex
Pith Epidermis
Key
to labels
cortex
vascular
bundle
pith vascular
cambium
xylem
Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
Monocot Stem
phloem
Anatomy
epidermis
vascular
bundles
ground
tissue
xylem
1) Primary Growth:
• Apical Meristems:
Mitotic cells at “tips” of roots / stems length
1) Increased length
2) Specialized structures (e.g.
fruits)
2) Secondary Growth: girth
• Lateral Meristems:
Mitotic cells “hips” of plant
Young
leaf
Developing
vascular
strand
Axillary bud
meristems
Cortex
Pith Primary
phloem
Primary
Root apical xylem Secondary
meristems Secondary phloem
xylem
Vascular cambium
primary phloem
Vascular cambium
• Is a lateral meristem in the vascular
tissue of plants.
• It is a cylinder of unspecialized
meristematic cells that divide to give
rise to cells that further divide,
differentiate and specialize to form
the secondary vascular tissues.
Note:
• The vascular cambium is a type
of meristem - tissue consisting of
embryonic (incompletely
differentiated) cells from which
other (more differentiated) plant
tissues originate.
pith
cortex
C X C
C
After one year After two years
C of growth of growth
Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings
Production of Secondary Xylem and Phloem
– The accumulation of this tissue over the years
accounts for most of the increase in diameter of a
woody plant.
– Secondary xylem forms to the interior and secondary
phloem to the exterior of the vascular cambium.
C=cambium cell
X=2o xylem
P=2o phloem
D=derivative
Cork cambium
• Another lateral meristem is the cork
cambium, which produces cork, part
of the bark. Growth
• Together, the secondary vascular
ring
tissues (produced by the vascular
cambium) and periderm (formed by Vascular
the cork cambium) makes up
the secondary plant body. ray
Heartwood
• Vascular cambia are found
in dicots and gymnosperms but Secondary
not monocots, which usually lack
secondary growth. xylem Sapwood
Secondary phloem
• Capon, Brian (2005). Botany for
Gardeners (2nd ed.). Portland, OR: Bark
Timber Publishing
Layers of periderm
Cork Cambium
• The cork cambium is a lateral
meristem and is responsible for
secondary growth that replaces
the epidermis in roots and stems.
heartwood
(xylem)
sapwood
(xylem)
vascular
cambium
phloem
annual ring
Sapwood = Young xylem, water
late Heartwood = Old xylem, support
xylem
Seasonal Growth = annual rings
early Secondary phloem = grows outward
xylem older phloem crushed
1 epidermis
3 primary phloem
4 fascicular cambium
5 interfascicular cambium
7 pith
2 cortex
6 primary xylem
Legend:
1 fascicular cambium
3 phloem
4 xylem http://www.vcbio.science.ru.nl/en/virtuallessons/fascicularcambium/
2 interfascicular cambium
The differentiation of the interfascicular cambium
• Ray initials
*In xylem and phloem these are the
parenchyma cells
*small cells perpendicular to the
axis of stem
*from sclerenchyma cells or ray
parenchyma that radially divide the
stem (like slices of pie)
*rays elongate perpendicular to axis
to stem
*cells transport water and dissolved
solutes radially
http://www.cas.muohio.edu/~meicenrd/mudescd/dendrology/SCDRYGTH/VC.gif
• Fusiform initials
a. tapered, prism-shaped cell, periclinal division
b. oriented parallel to stem axis
c. produce secondary xylem and secondary phloem
between rays
d. conducting cells of this tissue transport water and
dissolved solute longitudinally
e. Produces the axial (vertical) transport system
• Generally the xylem-producing cells are
more active than the phloem producing
cells
• In temperate areas, the cambium is active
from the spring to the fall and is inactive in
the winter
• The yearly activity of the cambium
produces the annual rings in the xylem
Periderm (cork)
A. Function of the periderm:
– Increase in diameter of the stem occurs with the activity of the
vascular cambium
– This causes the protective epidermis to crack and split open
– Thus, a need for a meristematic layer at the outer edge of the
phloem for the internal protection of the stem
– Thus a layer of cork cambium forms outside of the phloem.
– The cork cambium forms a layer of waxy cork cells
– The cylinder of cork cambium increase in diameter as the stem
increases in diameter
Formation of the cork
In the young stem (1 year old or less)
a. Cortical cells just under the epidermis become
meristematic
b. Produces a layer 1-2 cells thick of cork cambium
(phellogen)
c. Phellogen produces a layer of cork cells 4-6 cells
thick external (toward the epidermis) to the phellogen
d. Phellogen produces a single layer of cells,
phelloderm, internal (toward the xylem and phloem) to
the phellogen
Structure of Periderm