1

Atlas_2015.indd 1 25-04-2015 14:17

 Borissiak Paleontological Institute
Russian Academy of Sciences

Upper Vendian macrofossils

of Eastern Europe.
Middle Dniester area and Volhynia

A.Yu. Ivantsov, V.P. Gritsenko, V.M. Paliy, V.A. Velkanov, L.I. Konstantinenko ,
A.Sh. Menasova, M.A. Fedonkin, M.A. Zakrevskaya, E.A. Serezhnikova

Moscow
PIN RAS 2015

Atlas_2015.indd 2 25-04-2015 14:17

 . ..

.. , .. , .. , .. , .. ,
.. , .. , .. , ..

2015

Atlas_2015.indd 3 25-04-2015 14:17

 ISBN 978-5-903825-32-5

551.72(447.8)

.
. .. . - .: ,
2015. 144 ., 25 ., 10 . ISBN 978-5-903825-32-5
.

,
() .
, ,
.
.

.
, , .
: - , ,
, , , , , , Metazoa, Metaphyta,
.

Upper Vendian macrofossils of Eastern Europe.

Middle Dniester area and Volhynia. A.Yu. Ivantsov et al. - Moscow.: PIN RS,
2015. 144 p., 25 ill., 10 plates. ISBN 978-5-903825-32-5
In russian and english.
Publication represents report of main data on all species of Vendian and Early
Cambrian macrofossils, that were ever discovered on Middle Dniester area
(Podolia) and Volhynia. All taxonomic descriptions, images of type specimens,
data on their occurrence and storage places at State museums of Kiev and Moscow
were gathered into this edition. Descriptions are predominantly represented by
early edits. Overview of geological study condition and material on the history of
paleontological investigations of Vendian deposits in this region are also included
into report. It is recommended for geologists, paleontologists and biostratigraphers.
Key words: East European Platform, Middle Dniester area, Podolia, Volhynia, Ediacaran,
Vendian, Early Cambrian, Macrobiota, Metazoa, Metaphyta, trace fossils.

. 2015
ISBN 978-5-903825-32-5 : .. . 2015

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 /Content

 6

(.. , .. , .. )7

(Metazoa, )
-
- (.. )20

(.. , .. , .. , .. ,
.. , .. , .. )23

 52

/Pictures and captions 55

Introduction 76

Vendian of Middle Dniester area (V.A. Velkanov, V.P. Gritsenko, L.I. Konstantinenko)77

History of the studies of macrofossils (Metazoa, problematics and trace fossils)

in the Vendian and Lower Cambrian deposits of south-western margin
of the East European Platform (V.M. Paliy)89

Atlas of macrofossils from Upper Vendian and Lower Cambrian of Middle Dniester area
and Volhynia (A.Yu. Ivantsov, V.P. Gritsenko, V.M. Paliy, A.Sh. Menasova,
M.A. Fedonkin, M.A. Zakrevskaya, E.A. Serezhnikova)92

References119

/Plates and captions 123

Atlas_2015.indd 5 25-04-2015 14:17

 1.

-
, ( -
) 100 .
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-
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, (. 1).
-
( 53/111-2013) (
13-05-90435).

Atlas_2015.indd 6 25-04-2015 14:17

 2.
() ,

. -
, . , , -
, , , , ()
, ( , -
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,
(. 3). .
. 150. -
(, 1928; - ., 1968; , , 1980;
, 1981; , 1985; ., 1983, 1990; , 1985, 1988; , 1988,
1988; ., 1988; , 1993).

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(-).
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1952, 1972; Sokolov, 1964 .). - -
-, -
, , - , -
1960- 1990- 2000-.
.., .., .., ..-,
.., .., .., .., .., .. .
, -
.
- .
-
( ., 1983; , 1985;
, 2011).

Atlas_2015.indd 7 25-04-2015 14:17

 -
.
( , .).
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(1928). . -
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1931 ., : (.
: , , 2001).
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(-
., 1968). -
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100150
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) () . -
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.

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70-. ,
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(, , 1980; , 1981; ., 1983, 1990; , 1985; -
, 1987; , 1997) .

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20.
(. 7).
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-
( ., 1983; , 2006; , , 1985, ;
, 1985, 1987).
- ( ) -
- 30 (. 8).
Nemiana simplex Palij,

(.911).

,
. 20
(, 2012).
- - -
, -

10

Atlas_2015.indd 10 25-04-2015 14:17

 .
(, ), , -
( ., 1983). 25 (.12).
, .
. ( 10) -
, - -
, . -
(0,10,8), . (
7) -, , -
(.13).
Serebrina crustacea Istch., Eoholynia
Gnil. (.14). ( 5), , - -
, , e .. B.C.-
- (-
., 1968; , 1976).
(58)
, --
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-, , , -
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( ., 1983). 25.
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15 () (.15). -
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. -
, :
4 , 5,5 . 50. -
(, 1988; , , 1985,
), .
- -
( 1,01,5 )
(.16). -

11

Atlas_2015.indd 11 25-04-2015 14:17

 , : , ,
, .
, , , -
, .
. -
. - -
. , .. .. -
, (, 1985, 1988;
, 1987).
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25. ( 5070%)
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(-).
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, ()
, .
15. 45.
-
, -
(23); -
.
60 (.17).
-
, (. 18), ( ., 1983;
, 1987; , 1988; , 2006).
, -
- (, 1988).
. -
-
Phycodes (=Trichophycus) pedum, -
- (. ,
1992; Landing, 1994).
( ) ,
.
, -, -
.

12

Atlas_2015.indd 12 25-04-2015 14:17

 , -
(. 19).
( );
: Kamenecia stella Gur., Ternavellus
vialovi Gur.; (Planolites sp., Curvolithus sp.); -
() Sokoloviina sp., Parasabellidites sp. (-
., 1983; , 1993). 1517.
-, -
-, -
( 56), ( 0,3). -
: 1), 6,514,0;
2), ( 0,1) 2131; 3)-
, 1824 .
. .
.
,
. : Nemiana simplex Palij, Cyclomedusa minuta Fedonkin,
Elassenia zhuravleva Gur., Kullingia concentrica Glaessn. : Bergaueria
major Palij, Didymaulichnus tirasensis Palij, Treptichnus bifurcus Miller, T. triplex Palij, Planolites sp.
. . , , -
(, 1988; ., 1990).
-
.
, ,
-
( 70% ) . -
, .
.
. , -
( ) .
- 45 .
.
Planolites sp., Didymaulichnus tirasensis Palij, Treptichnus bifurcus
Miller (, 1988). , , -
- , -

. .. ( ., 1982) , -
, , -
, .
.., , , . -
.
.
-, .

- .
450 ( -
).
,
, 10.

13

Atlas_2015.indd 13 25-04-2015 14:17


1
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:
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( 70%) 0,7;
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() .
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- (2050 . .),
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1214. , ,
, , Nemiana simplex
Palij (.10. 11).
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, 7), -
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(-
). -
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(.7).
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( 3 5).
3
. , 1 , .. -
, (
):
: ; 23 (
);

14

Atlas_2015.indd 14 25-04-2015 14:17

 : - ,
; Cyclomedusa plana Glaessner;
2 ;
: ( - 1 )
(); -
; 56.
4
. . , . 0,5
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15

Atlas_2015.indd 15 25-04-2015 14:17

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( 2,5) - 4,6;
10) 5,1;
16

Atlas_2015.indd 16 25-04-2015 14:17

 :
11) (2 ) -, ,
, , -
3,7;
12) - ( 20%)
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-, , 7,6 .
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(Sokoloviina sp.) 6;
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-, - -,
. Phycodes pedum 1,7;

17

Atlas_2015.indd 17 25-04-2015 14:17

 6) , -
-, , Phycodes pedum, Didymaulichnus
sp. -, , , -
0,9.
(0,8 ) -, -
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7) , , -, ,
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, , ,
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(?) ,
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.

-
. . -
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Metazoa, , .
-
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) .
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, -
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. . (?), -
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, ( ).
, -
.
, -
.
-
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.-
, -
. .
. -
. , .-
.

18

Atlas_2015.indd 18 25-04-2015 14:17

 -
(
).
.
-
.,
, , -
. 2010. -
,
.
.
, -
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, ,
.

. -
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. . , -
-
.
-
(, )

,
.

19

Atlas_2015.indd 19 25-04-2015 14:17

 3. (METAZOA,
)

-
-
1916 ,
..,
, , -
( )
, .. (1916)
.
1920- . . -
.- ( )
.. (1928), -
() -
.
.
, ( -
-)
,
-

.
.. -
: , , , -
, -
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, ,
, ()
, 60- . ( ..-
(1939); - ..
(1958); .. (1956)).
: -
. (Walcott, 1898),
. , -
., ., . .
(Sprigg, 1947, 1949; Glaessner, Daily, 1959; Glaessner, Wade, 1966 .) -
, .
, , -
, -
, .
, ,
, .
- , -
1:200000 1:50000 ,
, ,
, -
, -

20

Atlas_2015.indd 20 25-04-2015 14:17

 . 1960-.
, -
-,
( ) , -
(, 1952 .). , -
, ,
, ,
, (Glaessner, 1959),
.
-
. 1960-. ( )
..- - -
, , ,
-
. ,
, -
. ,
1966 . . -
, . . ,
., ., ., ..
-
. -
( 100120)
, -
.

. (1966.) ..-
. . -
, Bronicella podolica (-, 1965)
Beltanelloides .., .. (1968) -
( ) . Sabellidites cambriensis, -
.
.. (1971) Vendotaenia
Tirasotaenia; -
.. .. (, 1983; ., 1988). -
,
() , -
.. (1976, 1988 .).
(. .--
) Cyclomedusa plana (-
., 1968). .. -

. 20
(. )
,
Nemiana, ,
(-, , 1968; -
, , 1968). Cyclomedusa Sprigg
Cyclomedusa serebrina (, 1969), ,
Tirasiana -
(, 1976; ., 1979).

21

Atlas_2015.indd 21 25-04-2015 14:17


()
() -
Harlaniella, .. (-
) . (, 1972), Palaeopascichnus,
.. . . (, 1976; .,
1979). ..
.. Bergaueria Didymaulichnus -
. (, 1974; , 1976; ., 1979),
.
..
(1975) -
Metazoa ( )
(,
1974). -
() Metazoa ,
, -
- . , ..
, -
. , ,
,
.. (1972) Charniodiscus planus, Planomedusites grandis Medusinites
patellaris, -
. ..
(, 1976; ., 1979),
(Paliy et al., 1983).
-
.. (1983, 1985, ), -

(Conomedusites, Dickinsonia, Ediacaria, Tribrachidium), , -
Metazoa.
.. (1983-, 1984, 1985, 1987, 1988), ..
(, 2003, , 2006) .. (2009).

- -
,
-
(, 2011). , -
- -
,
- ( ) (
., 1974 ). -
(Nemiana, Tirasiana Palaeopascichnus)
: - -
- , ,
- , ,
(Paliy, 1987; , 1996, 2010, 2013; Fedonkin et al., 2007; Ivantsov et al., 2014).

,
, -
(-, 1973), -
() (- , 1974),
(, 1990) .
22

Atlas_2015.indd 22 25-04-2015 14:17

 4.

-
() : , -
, , -
.
, , -
- .
, -
, -
, .
, ( ), ,
. -
, ,

.
-
. ,
, -
, , , -
..
( ),
Metazoa, -
.
, , ,
.
,
(?).
:
.
- .


. .. .
,

Beltanella Sprigg, 1947
Beltanella velikania Menasova, 2003
. I, . 1
(, 2003). ,
( ). .
. , .
(.. , ). Beltanella gilesi -
.
(.. , ). ..
-83501-, , 17134 (17142 ).

23

Atlas_2015.indd 23 25-04-2015 14:17

 Bessarabia Gureev, 1988
(, 1988). , , - -
-, -. -
, .
Bessarabia bukatchuki Gureev, 1988
. IV, . 8
(, 1988). -,
. -
.

Bronicella Zaika-Novatsky in Zaika-Novatsky et Palij, 1968

(, 1976). ,
, ,
.
(, 1976). Bronicella Beltanelliformis
Menner (= Beltanelloides Sokolov . ),
, ( 3-4 ) -
.
Bronicella podolica (Zaika-Novatsky, 1965)
. III, . 1
(, 1976).
(2-5 ) ( )
. , , ,
, , .
, -
, .

Charniodiscus Ford, 1958

Charniodiscus planus Sokolov, 1972
(, 1972). , -
72 .
(.. , .. , ). ,
(

). : , -,
; , - , , .
.. (1972, . I, . 4), .

Conomedusites Glaessner et Wade, 1966

(, 1985). , -
, . -
.
. .
Conomedusites cf. lobatus Glaessner et Wade, 1966
. IV, . 1
(, 1985). , -
, , -
- . .
. ,
, .
, . .
24

Atlas_2015.indd 24 25-04-2015 14:17

 -
, ()
, .
Cyclomedusa Sprigg, 1947
Cyclomedusa cf. davidi Sprigg, 1947
. II, . 8
. .
Cyclomedusa minuta Fedonkin in Palij et al., 1979
. IV, . 6
(, 1987). , -
. , ,
, . . -
.
( ., 1979). Cyclomedusa davidi
, C. wadea , C. conifor-
mis .
Cyclomedusa plana Glaessner et Wade, 1966
. I, . 8
(- ., 1968). , -
(?), ,
. -
, . . -
, ,
2/3 . -
. , .
, . ,
, ,
, , , .
Cyclomedusa serebrina Palij, 1969
. I, . 2
( ., 1979). 40-45 ,
( ) 3-5 . -
, -
, .
(, 1969). C. davidi
.
Dickinsonia Sprigg, 1947
Dickinsonia costata Sprigg, 1947
. VI, . 1
(, 1985). - ,
, .
. ,
. -
, ,
<>.
Dickinsonia cf. tenuis Glaessner et Wade, 1966
. VI, . 2
(, 1985). , -
.

25

Atlas_2015.indd 25 25-04-2015 14:17

 -
. , ,
, 25 1 .
.

Ediacaria Sprigg, 1947

(, 1985).
. , ,
. -
.
. , -, ,
.
Ediacaria flindersi Sprigg, 1947
(, 1985). <>. , , -
, -
,
.
-
, .
. -
.
, , .
,
.
. , , -
. <>.
(.. , ). -
.. (1983, . XXIX, . 1; 1985, . I, . 1, . II, . 4)
: , . , ; , -
- , , . ,
. 3994/668, 3994/287 .
Ediacaria sp.
. II, . 7
. .
Elasenia Fedonkin, 1983
(, 1983). , -
, -
,
, .
(, 1983). Medusinites Glaessner et Wade,
- ,
.
Elasenia aseevae Fedonkin, 1983
. II, . 6
(, 1983). , , , -
, ,
. , -
, , ,
: , -
. . , ,
. .
26

Atlas_2015.indd 26 25-04-2015 14:17

 Elasenia zhuravlevae Gureev, 1988
. III, . 3
(, 1988). , -
. -
.
(, 1988). ( . ), -
(E. aseevae Fedonkin) .

Glaessneria Gureev, 1987

(, 1987). . -
1/3 , --
. , -.
(, 1987). Planomedusites Sokolov -
, Kullingia Glaessner -
, .
Glaessneria imperfecta Gureev, 1987
. II, . 9
(, 1987). , , -
.
(, 1987). Glaessneria plana
Glaessner et Wade, 1966) . ), -
, ,
.
Gritcenia Menasova, 2003
(, 2003).
.
(, 2003). Nimbia , Broni-
cella .
(.. , ). Nimbia Fedonkin.
Gritcenia nana Menasova, 2003
. I, . 5
(, 2003). ,
. , , ,
, .
(.. , ). .. . 3/3, -
, . 1768.

Gureevella Menasova, 2003

(, 2003).
, .
(.. , ). Ediacaria Sprigg -
.
Gureevella elliptica Menasova, 2003
. I, . 3
(, 2003). ,
, ,
. , , -
. .
(.. , ). .. . 2/14,
, . 17150.
27

Atlas_2015.indd 27 25-04-2015 14:17

 Hiemalora Fedonkin, 1984 (=Pinegia Fedonkin, 1980)
(, 1980). --
, -
. , .
Hiemalora cf. stellaris Fedonkin, 1980
. VI, . 4
. .

Jampolium Gureev, 1988

(.. , ). , -
. ,
60.
(.. , ). 1988 . ..
,
, . , -
. , ,
, ,
.. .
(.. , ). Cyclomedusa Sprigg, 1947 -
, .
Jampolium wyrzhykowskii Gureev, 1988
. IV, . 2
. .

Irridinitus Fedonkin, 1983

(, 1983). , -
. . .
.
(, 1983).
Nemiana Palij, -
, .
Irridinitus multiradiatus Fedonkin, 1983
. II, . 5
(, 1983). I. multiradiatus
-, . , -
. , ,
. <...>, , -
, .
,
. ,
<>. -
. (-
) (
?), . ,
.
40 .

Kaisalia Fedonkin, 1984

(, 1987). -
. .

28

Atlas_2015.indd 28 25-04-2015 14:17

 Kaisalia levis Gureev, 1987
. II, . 2
(, 1987). .
, , , -
-. -
, .
(, 1987). -
.
Kamenecia Gureev, 1988
(, 1988). . , , -
, . 6 7.
Kamenecia stella Gureev, 1988
. V, . 7
. .
Kelleria Gureev, 1988
Kelleria kelleri Gureev, 1988
. .
(.. , ). Jampolium wyrzhykowskii Gureev,
1988 , . -
: , . , . , -
; , - , , .
() .. (1988, . XI, . 3), .
.
Kimberella Wade, 1972
aff. Kimberella sp.
. V, . 8
. .
(.. , ). ,
( ., 1985), .
Ternavellus vialovi Gureev, 1988.
Kullingia Glaessner in Fyn et Glaessner, 1979
(, 1987). , , -
. , .
.
(, 1987). Glaessneria Gureev Planomedusites Sokolov
.
Kullingia concentrica Glaessner, 1979
. IV, . 5
(, 1987). -
, -
( ).
, . -
. -
.
Lomosovis Fedonkin, 1983
(, 1983). .
-

29

Atlas_2015.indd 29 25-04-2015 14:17

 .
, . -
,
. .
Lomosovis malus Fedonkin, 1983
. VI, . 5
(, 1983). -
, , -
. - .
, , ,
, .
,
. :
, .
, , ,
( ) . -
.
Marnium Gureev, 1988
(, 1988). , -
. ().
(, 1988). Tirasiana Palij, -
,
Marnium Tirasiana .
Marnium cristatum Gureev, 1988
. VI, . 3
. .
Medusinites Glaessner et Wade, 1966
(, 1987). <> , -
.
Medusinites asteroides (Sprigg, 1949)
. III, . 4
(, 1987).
( ) -
, , . ,
.
(), -
, . -
, -
.
(, 1987). Medusinites paliji -
.
Medusinites paliji Gureev, 1987
. III, . 7 ,
(, 1987). Medusinites -
, .
.
Medusinites paliji Beltanelloididae Gureev, 1987.
(.. , ). (, 1988)
, 2127/23, -

30

Atlas_2015.indd 30 25-04-2015 14:17

 . .. (,
1987). , -
, 2127/19. -
(, 1987), , -
. III, . 7 .
Medusinites patellaris Sokolov, 1972
(, 1972). -
(20 ), , 65 .
(.. , .. , ). ,
( -
-
). : , -, ;
, - , , .
.. (1972, . II, . 2), .
Medusinites sokolovi Gureev, 1985
. IV, . 7
(, 1985). . , -
(?) ( ?).
(?). -
.
5:1.
(, 1985). M. patellaris Sokolov,
1972, M. asteroides (Sprigg, 1949), M. patellaris (Fedonkin, 1980) (
. ) -
, , , -
.
(.. , ). .. -
. -, .

Nemiana Palij, 1976

(, 1976).
. -
, () .
(, 1976). Bergaueria Prantl ( )
Nemiana , -
, ; Tirasiana
( ).
Nemiana simplex Palij, 1976
. III, . 2
(, 1976). ( ) ,
, , , .
, , .
.

; .
, -
, -
.
. ,
. , -
, .
31

Atlas_2015.indd 31 25-04-2015 14:17

 Nimbia Fedonkin, 1980
(, 1985). ,
( -
). .
, .
Nimbia dniesteri Fedonkin, 1983
. II, . 10
(, 1983). -
,
. ,
. , ,
. .
(, 1983). Nimbia occlusa Fedonkin,
, -
.
Nimbia occlusa Fedonkin, 1980
(, 1985). . -
3-5 . -
, ,
, .
.
(.. , ). .. (1985, . III, . 1, 6,
7) 3 (, 3994/533, 479-, 644) : -
, . , ; , - , -
, . .
Nimbia paula Gureev, 1985
. IV, . 4
(, 1985). , -
, . -
, . .
(, 1985). N. occlusa Fedonkin, -
.

Onuphionella Kirjanov, 1968

(, 1968). ,
, , -
.
Onuphionella agglutinata Kirjanov, 1968
. IX, . 4 ,
(, 1968). ,
(?) . -
.
. -
. , . -
0,1-0,3 .
, . 1,5 ,
6,5 . 4 .
Palaeospinther Menasova, 2003
(, 2003). , .
(, 2003). Dickinsonia Sprigg .
32

Atlas_2015.indd 32 25-04-2015 14:17

 Palaeospinther nucis Menasova, 2003
. V, . 2
(, 2003). - , -
, .
, (-
).
(.. , ). .. -
3/2, , . 17151.
Palaeospinther conoideus Menasova, 2003
. V, . 1
(, 2003). -
, .
-
. , .
(.. , ). P. nucis (
, ) , .
(.. , ). ..
-83503-, , . 1781.
Paliella Fedonkin, 1980
(, 1985).
, -
. . ( -
) .
(, 1985). Paliella Cyclomedusa,
C. plana
Glaessner et Wade. Paliella Medusinites asteroides Sprigg,
, .
Paliella patelliformis Fedonkin, 1980
. II, . 4
(, 1985).
, -
, , , -
. ,
, -
, . -
, .
, ,
. , Paliella
. (
: ), , -
,
.
Planomedusites Sokolov 1972
(, 1972). ,
, 125 .
Planomedusites grandis Sokolov 1972
. .
(.. , .. , ). ,
(

33

Atlas_2015.indd 33 25-04-2015 14:17


). : , -,
; , - , , .
.. (1972, . II, . 1), .
Platysolenites Pander, 1850
Platysolenites antiquissimus Eichwald, 1860
. IX, . 2
(, 1968). . -
, , , ,
. , , -
. , -
. <>.
Podoliina Gureev, 1988
(, 1988).
- , , -
. , , ,
. , ,
.
Podoliina crassa Gureev, 1988
. VIII, . 1
. .
Podolimirus Fedonkin, 1983
(, 1983). - , -
, -
. 80-85,
. -
.
, .
(, 1983).
Pteridinium Grich , , Pteridiniidae Richter.
,
.
Podolimirus mirus Fedonkin, 1983
. V, . 5
(, 1983). () -
, . , , .
, , . ,
.
, -
, .
-
. 10-12 ,
, . -
, , ,
. -
, -.
Pollukia Gureev, 1987
(, 1987). ( ) -
.
34

Atlas_2015.indd 34 25-04-2015 14:17

 (, 1987). Tirasiana Palij ,
.
Pollukia shulgae Gureev, 1987
. II, . 1
(, 1987). -
, .
() . .
, .
(, 1987). Pollukia serebrina (Cyclomedusa serebrina Palij, 1969
. ) , .

Propaleolina Menasova, 2003

(, 2003). , -
. . -
.
(, 2003). Paleolina
.
Propaleolina vendiensis Menasova, 2003
. IX, . 1 ,
(, 2003). -
,
. , , . -
, , , .
. .
(.. , ). ..
. 6/1, , . 1782.

Protodipleurosoma Sprigg, 1949

(, 1985). , -
(?) , ,
, (?)
( ?),
, , (-
?), .
Protodipleurosoma rugulosum Fedonkin, 1980
(, 1985).
.
, ,
. -
, . -
,
. , -
, .
(, 1985). Protodipleurosoma wardi Sprigg
,
.
(.. , ). , 3994/406,
.. (1983, . XXXI, . 5), : ,
. , ; , - , , -
. .

35

Atlas_2015.indd 35 25-04-2015 14:17

 Protodipleurosoma wardi Sprigg, 1949
. IV, . 9
. .
Pseudorhizostomites Sprigg, 1949
Pseudorhizostomites sp.
. .
(.. , ). , 3994/594,
.. (1983, . XXXI, . 2), : ,
. , ; , - , , -
. .
Sabellidites Yanischevskii, 1926
Sabellidites cambriensis Yanischevskii, 1926
. IX, . 5 ,
(, 1968). , , ,
, . -
, , -
; , .

Sekwia Hofmann, 1981

(, 1987).
.
Sekwia kaptarenkoe Gureev, 1987
. III, . 5
(, 1987). ( ) -
,
. , . -
. .
(, 1987). Sekwia excentrica Hofmann -
.

Serpulites Murchison, 1839

Serpulites (?) petropolitanus Yanischevsky, 1926
. .
(.. , ). , 1731/5,
.. (1968, . V, . 24), : ., . . , . 5,
. 180,1 ; , , ( -
). .

Sokoloviina Kirjanov, 1968

(, 1968). ,
, ,
.
(, 1968).
Paleolina Sokolov. , Sokoloviina , -
, .
Sokoloviina costata Kirjanov, 1968
(, 1968). ,
, , ,
.

36

Atlas_2015.indd 36 25-04-2015 14:17

 , , , , .
0,03-0,1 . , .
.

. 15 , 0,3 1 .
(.. , ). , 1731/2,
.. (1968, . III, . 4), : ., ,
. , . 17, . 202,1-202,2 ; , ,
( ). .
Studenicia Gureev, 1983
Studenicia galeiforma Gureev, 1983
. VIII, . 5
(, 1983 ( , .
)). , -
. .
(, 1983 ( ,
. )). Nemiana , Tirasiana
, .
(.. , ).
(. Monocraterion sp.). .. (1983)
1970/64, , . 2088/9.
Ternavellus Gureev, 1988
(, 1988). (
). .
Ternavellus vialovi Gureev, 1988
. V, . 8
(, 1988). -
( ). .
(, ?). , -
. .
, .
Tirasiana Palij, 1976
(, 1976).
. , -
. -
.
(, 1976). Nemiana Palij Tirasiana
, ,
.
Tirasiana coniformis Palij, 1976
. I, . 7
(, 1976). T. coniformis
.
. -
. , --
, -
. , -
, .
, -, , .
37

Atlas_2015.indd 37 25-04-2015 14:17

 (, 1976). T. disciformis -
.
Tirasiana disciformis Palij, 1976
. I, . 6
(, 1976). (3-4 )
.
, ,
.
. -
- .
, , -
. ( 3-4 ) .
.
(, 1976). T. coniformis Palij T. disciformis
.

Tribrachidium Glaessner, 1959

(, 1985). ,
. (?) -
, , -
, . Y-
.
(). 0,7
. -
, , -
, -
. (?)
.
Tribrachidium heraldicum Glaessner, 1959
. V, . 3
(, 1985). . -
( . ) .
,
. -
(
20 ), .
,
, -
. , -
( ),
, (
, . -
). .
,
.
. -
,
.
, <>. ,
, -
, .

38

Atlas_2015.indd 38 25-04-2015 14:17

 , , ,

, .
, , -
-
, .
, ,
- <>.
(.. , ). .. ,
3994/580-A; , 4158/26, . .. .

Valdainia Fedonkin, 1983

(, 1983). . -
, . -.
. . -
, .
, . .
(, 1983). Pteridinium Grich
, , , -
, , , -
.
Valdainia plumosa Fedonkin, 1983
. V, . 4
(, 1983). -
,
, -
, . ,
. ,
. -
. -
, . ,
,
. ,
, -
. , , -
, .
,
30-40.

Vaveliksia Fedonkin, 1983

(, 1983). , -
, :
, , ,
, , ,
, . , -
, ,
, .
Vaveliksia svetozarovae Gureev, 1988
. V, . 6
(, 1988). (-
). . -

39

Atlas_2015.indd 39 25-04-2015 14:17

 - . ,
, , (-
). .
(, 1988). (V. velikanovi Fedonkin) -
, , .
Vaveliksia velikanovi Fedonkin, 1983
. III, . 6
(, 1983). 3-4, 1,5
. ,
,
.
, . -
, ,
.
Vendella Gureev, 1987
(, 1987). , -
.
(, 1987). Nimbia Fedonkin
.
Vendella haelenicae Gureev, 1987
. II, . 3
(, 1987). , , .
( ) , -
( ) .
, , , .
(, 1987).
.
Vendella larini Gureev, 1987
. IV, . 3
(, 1987). -
, ( ?) .
,
.
(, 1987). V. sokolovi V. haelenicae -
.
Nimbia Fedonkin.

. VIII, . 6
( ., 1979). 6-11
1-2 , 3-4 -
- <>. , ,
, , 15-28
3-5 . 3-5 .
( ., 1979). Nemiana Tirasiana, -
.

40

Atlas_2015.indd 40 25-04-2015 14:17

 ,
, ,

Anulichnus Gureev, 1983

Anulichnus segmentatus Gureev, 1983
(, 1983 ( , .
)). , -
, .
(.. , ). , .. (1983,
. I, . 7), : , .
. ; , , . -
.
Asterichnus Bandel, 1967
Asterichnus vialovi Gureev, 1984
(, 1984). , , -
, , , -
. , ,
. 25.
(, 1984). Asterichnus lawrencensis
Bandel
-
.
(.. , ). , .. (1984,
. 1 ), : , . , . ; , -
, , . ..
: , 2088/3,
.
Aviculaichnus Gritsenko, 2009
(, 2009). - ,
Phycodes pedum , -
.
.
(, 2009).
Phycodes pedum, .
Aviculaichnus gureevi Gritsenko, 2009
. VIII, . 3
(, 2009). - , -
.
. .
4-5 20 .
. . 4-5
10-12 .
(.. , ). ..
. 2480/1, , 2525/93.
Bergaueria Prantl, 1945
(, 1976).
.
- , -
. .

41

Atlas_2015.indd 41 25-04-2015 14:17

 Bergaueria major Palij, 1976
. VII, . 5
(, 1976).
, -
. , -. -
,
, .
, . -

.
, Bergaueria major .
, -
. .
, .
(, 1976). Bergaueria
-
. B. major <>

Chomatichnus Donaldson et Simpson, 1962

Chomatichnus loevcensis Gureev, 1984
(, 1984). , ,
, -
. ( 4 ) ,
; , .
( 4-5 1 ). -
-
( ).
(, 1984). Chomatichnus wegbergensis Donaldson et Simpson (
) , , -
.
(.. , ). , .. (1984,
. 2 ), : , . , . ; ,
, , . ..
: , 2088/4,

Circulichnus Vialov, 1971

Circulichnus montanus Vialov, 1971
(, 1983).
. , , . -
(2 ),
. ,
- , .
(, 1983).
(8-10 ), Circulichnus montanus Vialov (35-40 ).

.
(.. , ). , .. (1983,
. . 9), : , . , . ; , -
, , . ..
: , 1970/57, .

42

Atlas_2015.indd 42 25-04-2015 14:17

 Cochlichnus Hitchcock, 1858
Cochlichnus isp.
. VII, . 3
(, 1976).
. , ,
. , .

.
Didymaulichnus Young, 1972
Didymaulichnus tirasensis Palij, 1974
. VII, . 6
( ., 1979). -
, 8-14 .
, .
0,5-2,5
1,5 , , .
(, 1974). Didymaulichnus miettensis Young -
, , , -
.
Didymaulichnus cf. miettensis Young, 1972
. .
. .
.. (1984) : , . 2088/6.
. , 1984, . 1 .
(, 1984).
, .
, -.
8-10 .
(, 1984). Didymaulichnus miettensis Young,
,
, .
. , - , -
, . , . ,
. .
(.. , ). , .. (1984,
. 1 ), : , . , . ;
, - , , .
.. : , 2088/6, -
.
Harlaniella Sokolov, 1972
(, 1976). ( ), -
, .
.
Harlaniella podolica Sokolov, 1972
. VI, . 6
(, 1976). , -
, 45-80
. ,

43

Atlas_2015.indd 43 25-04-2015 14:17

 . . -
, , , -
,
Palaeopascichnus delicatus. , -
: .
, , .

Monocraterion Torell, 1870

Monocraterion isp.
. VIII, . 5
(, 1984).
. , -
.
(.. , ). ..
(. Studenicia galeiforma Gureev, 1983).

Palaeopascichnus Palij, 1976

(, 1976).
( ). ,
. , ,
.
(, 1976). Helmintoida Schafhutl -
, Helmintoida -
, .
Palaeopascichnus delicatus Palij, 1976
. VI, . 7
(, 1976). -
, , -
. .
, ,
, . -
. 4 10 .

Phycodes Richter, 1850

Phycodes pedum Seilacher, 1955
. VII, . 7
. .

Planispiralichnus Fedonkin, 1985

Planispiralichnus rarus Menasova, 2003
. VII, . 4 ,
(, 2003). , -
( ). . -
. -
. ,
. , ,
.
(.. , ). .. . -8116,
, . 1741.

44

Atlas_2015.indd 44 25-04-2015 14:17

 Pseudohiemaloraichnus Gritsenko, 2009
(, 2009). (
Pascichnia
) , -
, ,
, .. Hiemalora stellaris.
Pseudohiemaloraichnus podolica Gritsenko, 2009
. VIII, . 4
(, 2009). 35 ,
. ( ) (-
) 2 , , .
-, .
Sokolovichnites Gureev, 1983
(, 1983). -
, -
. -,
.
Sokolovichnites angelicae Gureev, 1983
(, 1983). ( ).
, , . -
, , -
( 3-6 ) .
(.. , ). , .. (1983,
, . 1), : , . . ,
100 . , . ; , ,
. .. :
, -81-53, .
Sokolovichnites isp.
(, 1983). , ,
. ,
.
(, 1983). -
( , .
).
(.. , ). , .. (1983,
, . 8), : , . . ,
150 . ; , , -
. .. : ,
-81-17, .
Treptichnus Miller, 1889
Treptichnus bifurcus Miller, 1889
. VII, . 1
. .
(.. , ). .. ,
3994/26-. . .. ,
4158/32.
Treptichnus triplex Palij, 1976
. VII, . 2
(, 1976). ,
20-40 . ,

45

Atlas_2015.indd 45 25-04-2015 14:17

 , . -
, , -
.
.
. .
(, 1976). ,
Treptichnus, .
Veprina cf. undosa Fedonkin, 1980
. VIII, . 2
(, 2009) ( .. Veprina undosa Fedonkin, 1980
,
. ). , -
, .
20 40 , 2-5 . - -
, . () -
, .
(, 2009).
. Veprina undosa Fedonkin
- - .
(.. , ).
, 2480/11 (, 2009, . 34, . 3, . I, . 8). -
(, 2525/103), .

Beltanelloides Sokolov, 1965

( ., 1988). , 5-30 ,
-
. ,
.
( ., 1988). Chuariacea
, .
(.. , ). Beltanelloides
amorphus Menasova, 2003,
. (-
) Metazoa, .
-

Beltanelloides.
Beltanelloides amorphus Menasova, 2003
. I, . 4
(, 2003). ,
. , -
, . , .
,
, . ,
, .
(.. , ). B. sorichevae Sokolov -
, , .

46

Atlas_2015.indd 46 25-04-2015 14:17

 (.. , ). ..
. 2/49, , . 1742.
Beltanelloides podolicus A. Istchenko in Gnilovskaya et al., 1988
. X, . 1
( ., 1988). -
, , - ,
.
, -
. -
10-20 , , 5 10 -
35 .
(). , , -
, . , , -
. -
, 0,5-1 . ,
, . -
, , , .
1-2 .
. , ,
,
(, ) , -
.

Eoholynia Gnilovskaya, 1975

. ( ., 1988). , ,
. 10 . -
, . , ,
3 , , ,
. 50-100 .
Eoholynia capillaria A. Istchenko in Gnilovskaya et al., 1988
. X, . 3
( ., 1988). , -
, 2-3 .
, , .
, , , -
. ,
. 0,02-0,03 <>.
( ., 1988).
, .
Eoholynia fruticulosa A. Istchenko in Gnilovskaya et al., 1988
. X, . 7
( ., 1988). ,
, 2-3 .
-
. , , ,
. , -
, .
20-30 <>.
( ., 1988). (E. mosquensis Gnilovskaya, 1975
. ) () .

47

Atlas_2015.indd 47 25-04-2015 14:17

 Eoholynia longa A. Istchenko in Gnilovskaya et al., 1988
( ., 1988). , 2-3 , -
. , ,
0,01-0,02 .
0,3-0,5 . , , .
, , -
2-3 .
( ., 1988). -
, . , .
, ,
1-2 .
(.. , ). , 2235/14,
.. ( ., 1988, . IV, . 2), : , .
. ; , - , , . -
.

Fusosquamula Aseeva, 1976

(, 1976). , ,
- .
Fusosquamula vlasovi Aseeva, 1976
. X, . 6
(, 1976). ,
. . .
0,5-1 , -. 3-10 .
, , . -.
, ,
.

Kalusina A. Istchenko in Gnilovskaya et al., 1988

( ., 1988). , , -
. , ,
, .
( ., 1988). Eoholynia Gnilovskaya
.
Kalusina compacta A. Istchenko in Gnilovskaya et al., 1988
. X, . 5
( ., 1988).
. , -
. , ,
. , ,
, , -
. 1-2 , , 0,02-0,03 .
5-6 .

Kanilovia A. Istchenko, 1983

(, 1983). , (-
?), , . ,
.
, .
(, 1983). Vendotaenia Gnilovskaya -
, .

48

Atlas_2015.indd 48 25-04-2015 14:17

 Kanilovia insolita A. Istchenko, 1983
. X, . 9
(, 1983). , , -
, , , 3,5 . , -
. 1,2-1,4 ,
2 . . -
. -
. ,
, , . ,
. 0,03-0,036 , -
0,07-0,075 . 0,08-0,09 .
0,1-0,3 ( ). -
, -
. .

Pilitela Aseeva, 1976

(, 1976). , .
, , .
Pilitela composita Aseeva, 1976
. X, . 4
(, 1976). 0,2
, , , . -
0,5-1,5 ,
. -.
.

Redkinia Sokolov, 1977

Redkinia fedonkini Assejeva, 1988
(, 1988). -
(?). ,
. .
.
150-200 20-30 50-60
- -
5 ,
6 25 . ,
7-8 40-45 . , ,
.
(, 1988). <> R. spinosa Sokolov -
<>.
(.. , ). : -
, . 2 , . 320,0 ; , - , , -
. .. (1988, . XXI, . 1-3), -
.

Serebrina A. Istchenko in Gnilovskaya et al., 1988

( ., 1988). -
, ,
.
. - , -
, .

49

Atlas_2015.indd 49 25-04-2015 14:17

 ( ., 1988). Vendotaenia Gnilovskaya
, .
Serebrina crustacea A. Istchenko in Gnilovskaya et al., 1988
. X, . 2
( ., 1988). , -
- .
, .
-
. -
, 1,5-2 , , 2-3 , ,
, . -
, , , -
. ,
, .
-. , ,
.
, , . , -,
. 0,005 0,05-0,06 ,
0,002-0,003 .

Tawuia Hofmann, 1979

( ., 1988). , -
, , 8 , 0,6 , , -
. , .
. .
Tawuia dalensis Hofmann, 1979
. X, . 8
( ., 1988). , -
: 0,5 2,5 ( , ).
0,8 2-3 , . ,
U S. ,
.

Tyrasotaenia Gnilovskaya, 1971

(, 1971). , ,
. .
(, 1971). Tyrasotaenia
Vendotaenia, .
Vendotaenia ,
. Tyrasotaenia -
, , . , -,
, Vendotaenia , , Tyrasotaenia -
. Tyrasotaenia - ,
Vendotaenia. Vendotaenia
. , -
, . Tyrasotaenia
- , , , -
.
Tyrasotaenia podolica Gnilovskaya, 1971
(, 1971). -
, . -

50

Atlas_2015.indd 50 25-04-2015 14:17

 , -
. , -
. , Tyrasotaenia , -
. , ,
. ,
. , -
40 , 0,5 .
, -
. , , -, -
, .
(.. , ). , 2236/27,
.. (1983, . XIX, . 5), : , . . ;
, , , .
.

Vendotaenia Gnilovskaya, 1971

. ( ., 1988). , , -
-. , ; -
, 0,11 3,5 . , -
, . ,
58 , , -
(?). 50-90 .
Vendotaenia antiqua Gnilovskaja, 1971
. IX, . 3
(, 1976). -
, . 1,5-2,0,
70 . , , . -
, . -
, ( -
.. ,
. ).

51

Atlas_2015.indd 51 25-04-2015 14:17


.. - // -
- --
. : , 1976. . 4063.
.. // .., .., ..
. : , 1983. . 96101.
.. // -
. : , 1988. . 8192.
.. // -
. : , 1988. . 93102.
.. // -
. 1996. 5. . 111131.
.. -
// . 2010. 43. . 1835.
.. //
. 2013. 1. . 5280.
.. // . - -
. . 2. . .: , 1985. . 3567.
.. // . . 2011. 1. . 4249.
.., .., .. . : . 1983. 162 .
.., .., .., .., .. . -
III . ,
1990. 129 .
.. -
// . 1956. . XVI. . 1.
.. ( ) // -
. . 1971. 3. . 101107.
.., .., .., .., .. -
. .: , 1988. 142 .
.. (- )
//
: . . . . ,
2009. . 3035.
.. , -
//
. : , 1983. . 3439.
.. -
// . . -. 20. : , 1983. . 7073.
.. // . .
. 43. 1. : , 1983. . 130132.
.. // .
. . . . . . . 1984. 4. . 59.
.. Vendiata Radialia // -
. .: , 1985. . 117125. (. . . . . 632).
.. . , . 1987. 54 .
.. // -
. : . 1988. . 6580.
.., .., .. -
// . , . . 1985. 6. . 1013.
., .., .. -
. .: , 1982. 105 .
- .. // -
(. .). :
. 1965. . 9899.
- ..
// . . II. . : , 1971. . 158163.

52

Atlas_2015.indd 52 25-04-2015 14:17

 - .., .., .. -
( ) // . . 1968. 2. . 133134.
- .., ..
// . . 1968. 5. . 1. . 130135.
- .., ..
// . . 1974. 11. . 1. . 5965.
. 29- , 1992 ., . ,
1992. 70 .
.. // -
. : , 1983. . 181206.
.. //
.-. . .-. 1928. . 2. . 87103.
.., .., .., .. Metazoa -
// . . . . 1974. 12. . 130134.
.. --
// -. : ,
1968. . 527.
.. - --
// -
. : . 1993. . 4752.
.., .. //
. V . . : , 1968. . 9192.
... - . :
. 1981. 56 .
.., .. -
. : . 1980. 57 c.
.. ( ) //
. . -. ., . . . . . 3. 1916. . 2227.
.. // -
1938 . : . 1939. . 107148.
.. . . 2012. 4. .
97104.
.. // . . . .
2003. . 25. . 2526.
.. //
/ . . 2003.
. 139142.
.. Metazoa
// . . .-. . . 2006. 23 .
.. // . . 1969. 6. . 1. . 110113.
.. Metazoa
( ) // c
- . , 1974. 184 .
.. () // ,
. 1974. 6. . 499503.
..
// -
- - . : , 1976. . 6377.
..
. . 2011. 34. . 8588.
.., ., .. -
// --
. .: , 1979. . 4982.
.., .., .., .., .., ..
- - -
. : , 1976. 168 .

53

Atlas_2015.indd 53 25-04-2015 14:17

 .. // . . .
. 1952. 5. . 2131.
.. // XXIV . . . . . . .:
, 1972. . 114125.
.. . .: Scientific Press Ltd. 1997. 154 .
.., .. (.) . -
. . 1. . .: , 1985. 221 .
.., .. (.) . -
. . 2. . .: , 1985. 238 .
.. // . .
. . . 1958. . 21. 44 .
.. // .., .., -
.. . : , 1983. . 128139.
.. Radialia // -
.: , 1984. . 3058.
.. Bilateria Articulata //
. .: , 1985a. . 7992. (. . .
. . 632).
.. Metazoa // . --
. . 1. . .: , 1985. . 70106.
.. Metazoa. .: , 1987. 174 . (.
. . 226).
.. - // .
. . . 1952. 4. . 320323.
.., .. - -
// . -. . . 2001. . 15. . 135143.
Fedonkin M.A. Vendian body fossils and trace fossils // Ed. S. Behgtson. Early Life Earth/ New York: Columbia
University press, 1992a. P. 370388.
Fedonkin M.A. Vendian Faunas and the Early Evolution of Metazoa // Eds J.H.Lipps, P.W.Signor. Origin and the
Early Evolution of Metazoa. New York: Plenum Press, 1992b. P. 87129.
Fedonkin M.A., Gehling J.G., Grey K., Narbonne G., Vickers-Rich P., Clarke A.C. The Rise of Animals: evolu-
tion and diversification of the Kingdom Animalia. Baltimore: Johns Hopkins University Press, 2007. 326 .
Glaessner M.F. Precambrian Coelenterata from Australia, Africa and England. Nature. 1959. V. 183. P. 14721473.
Glaessner M.F., Daily B. The geology and late Precambrian fauna of the Ediacaran fossil reserve. Records of the
South Australian Museum. 1959. V. 13. P.369401.
Glaessner M.F., Wade M. The Late Precambrian fossils from Ediacara, South Australia // Palaeontology. 1966.
V. 9. P. 599628.
Ivantsov A.Yu., Grytsenko V.P., Konstantynenko L.I., Zakrevskaya M.A. Revision of the problematic Vendia mac-
rofossils Beltanelliformis (=Beltanelloides, Nemiana) // Paleontological Journal. 2014. V. 48. 13. P. 126.
Landing E. Precambrian-Cambrian boundary global stratotype ratified and a new perspective of Cambrian
Time // Geology. 1994. V. 22. P. 179182.
Paliy V. M. Ichnology Newsletter. 1987. July 16. P. 20.
Paliy V.M., Posti E., Fedonkin M.A. Soft-bodied Metazoa and animals trace fossils in the Vendian and Early
Cambrian // Upper Precambrian and Cambrian palaeontology of the East-European platform. Warszawa:
Wydawnictwa geologiczne, 1983. P. 5694.
Sokolov B.S. The Vendian and the problem of the boundary between the preCambrian and Paleozoic Group // Re-
port of The Twenty-Second Session of International Geological Congress. Pt X. Archaean and PreCambrian
geology. New Delhi, 1964. P. 288304.
Sokolov B.S. Vendian of Northern Eurasia // Arctic Geology. Proc. Second Intern. Symp. Arctic Geology. Febr.
14. 1971. San Francisco, Cal., Tulsa, Okla., USA. 1973. P. 204218.
Sprigg R.C. Early Cambrian ? jellyfishes from the Flinders Ranges, South Australia // Trans. roy. soc. South
Australia. 1947. 71. P. 221224.
Sprigg R.C. Early Cambrian ? jellyfishes of Ediacara, South Australia, and Mouht John, Kimberley District,
Western Australia // Trans. roy. soc. South Australia. 1949. 73. P. 7279.
Walcott C.D. Fossil medusae. U.S. geol. surv. V. 30. Washington. 1898. 201 p.

54

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PICTURES AND CAPTIONS

. 1. .. ( , 2011 .).
Text-fig. 1. L.I. Konstantinenko (vicinity of Dnestrovskaya Hydroelectric Power Station, October, 2011).

55

Atlas_2015.indd 55 25-04-2015 14:17

 . 2. -
. - ( .,
1990, ). : 1 , 2 ,
3 , 4 , 5 , 6 , 7 -
, 8 , 9 , 10 , 11
, 12 , 13 , 14
.
Text-fig. 2. Scheme of geological structure of Middle Dniester area and geological section across Dniester
river with removed Quarternary and Meso-Cenozoic deposits (by Velikanov et al., 1990, with changing). Legend:
1 crystalline rocks, 2 Grushka Formation, 3 Mogilev Formation, 4 Yaryshev Formation, 5 Nagoryany
Formation, 6 Danilovka Formation, 7 Zharnovka Formation, 8 Krushanovka Formation, 9 Studenitsa For-
mation, 10 Okunets and Khmelnitsky Formation, 11 Ordovician deposits, 12 Silurian deposits, 13 bound-
ary of the pericratonic trough, 14 line of geological section.

. 3. ( ., 1990,
). : 1 ; 2 ; 3 , -
; 4 -, ; 5 -, ; 6 -
; 7 ; 8 ; 9 ; 10 ; 11
; 12 ; 13 Metazoa; 14 ;
15 ; 16 , 17 , 18 .

Text-fig. 3. Composite section of the Vendian and Lower Cambrian deposits of Middle Dniester area
(by Velikanov et al., 1990, with changing). Legend: 1 basement rocks; 2 breccias; 3 conglomerates, grit-
stones; 4 coarse-grained sandstones; 5 medium-, and fine-grained sandstones; 6 siltstones; 7 mudstones;
8 basalts; 9 tuffaceous mudstones; 10 phosphoritic concretions; 11 Platysolenitidae; 12 Sabelliditidae;
13 imprints of non-skeletal Metazoa; 14 fossil traces; 15 plant macroremains; 16 microfossils, 17 onco-
lites, 18 red-colored rocks.

56

Atlas_2015.indd 56 25-04-2015 14:17

 57

Atlas_2015.indd 57 25-04-2015 14:17

 . 4. ( -
). : 1 -
, 2 , 3 , 4 ,
5 , 6 -, , 7 . : I -
, II , III , IV , V -
, VI , VII , VIII ,
IX , X , XI .

Text-fig. 4. Structural plan of Bernashevka Rise (Dnestrovskaya Hydro-

electric Power Station area). Legend: 1 crystalline basement, 2 gritstones,
3 varied-grained sandstones, 4 siltstones, 5 mudstones, 6 tuff-mud-
stones, tuffites, 7 quarry. Beds: I Olchedaev, II Lomozov, III Yampol,
IV Lyadova, V Bernashevka, VI Bronnitsa, VII Zinkov, VIII Dzhur-
zhevka, IX Kalyus, X Cenomanian, XI Quarternary.

58

Atlas_2015.indd 58 25-04-2015 14:17

 . 5.
. , . .

Text-fig. 5. Stratotype of Grushka Forma-

tion near Grushka village, right side of the valley
of Murafa river.

. 6.
.

Text-fig. 6. Boulders of the stratotype

of Grushka Formation.

59

Atlas_2015.indd 59 25-04-2015 14:17

 . 7. . , , -
(2011 .).
Text-fig. 7. Left bank of Dniester river, Dnestrovskaya Hydroelectric Power Station quarry, contact of tabu-
late Lomozov Beds mudstones of Mogilev Formation with crystalline basement rocks (photograph of 2011).

. 8. . . ; .

Text-fig. 8. Left bank of Nemija river near Ozarintsy village; sandstones of Yampol Beds.

60

Atlas_2015.indd 60 25-04-2015 14:17

 61

Atlas_2015.indd 61 25-04-2015 14:17

 . 9. . . ; ; -
Nemiana simplex Palij, 1976.
Text-fig. 9. Right bank of Nemija river near Ozarintsy village; sandstones of Yampol Beds;
locality of the holotype of Nemiana simplex Palij, 1976.

. 10. . , , .
Text-fig. 10. Left bank of Dniester river, Dnestrovskaya Hydroelectric Power Station quarry,
sandstones of Yampol Beds.

62

Atlas_2015.indd 62 25-04-2015 14:17

 . 11. Nemiana simplex Palij, , .

Text-fig. 11. Imprints of Nemiana simplex Palij, Dnestrovskaya Hydroelectric Power Station quarry, Yampol
Beds.

63

Atlas_2015.indd 63 25-04-2015 14:17

 . 12. ( 30 ) ,
( )
Text-fig. 12. Lyadova Beds with thick (up to30 cm) bentonite interbed, light line on
the image (Dnestrovskaya Hydroelectric Power Station quarry).

. 13. -
. -.

Text-fig. 13. Bernashevka Beds of Yaryshev Formation with bentonite interbed at the
Borshchov Yar on the periphery of Mogilev-Podolsky town.

64

Atlas_2015.indd 64 25-04-2015 14:17

 . 14. . -, ; Beltanelloides
podolicus A. Istchenko, 1988.

Text-fig. 14. Periphery of Mogilev-Podolsky town, Borshchov Yar; locality of the holotype of Beltanelloides
podolicus A. Istchenko, 1988.

65

Atlas_2015.indd 65 25-04-2015 14:17

 . 15. . .

Text-fig. 15. Kalyus Beds with phosphorites in ravine on the periphery of Tymkov village.

66

Atlas_2015.indd 66 25-04-2015 14:17

 . 16.
. . .

Text-fig. 16. Contact of Kalyus Beds of Nagoryany Formation with Pilipy Beds of Danilovka Formation,
Kanilovka Group. Periphery of Tymkov village.

67

Atlas_2015.indd 67 25-04-2015 14:17

 . 17. . ,
.

Text-fig. 17. Left bank of Dniester river near former Bakota village, landslide of the
upper part of Studenitsa Formation.

. 18. (Harlaniella sp.)

. -
.
Text-fig. 18. Various Vendotaenia algae and bioglyphs (Harlaniella sp.) in Komarovo
Beds of Studenitsa Formation. Outcrop under Bakota cave monastery.

68

Atlas_2015.indd 68 25-04-2015 14:17

 . 19. . -
.
Text-fig. 19. Gradual transition between Studenitsa and Okunets Formations at the outcrop below Kitaygorod
village.

69

Atlas_2015.indd 69 25-04-2015 14:17

 . 20. .
, . (
., 1990, );
. . 25.

Text-fig. 20. Section of Mogilev Formation at

Vysshy Olchedaev village area, Murafa river (by Ve-
likanov et al., 1990, with changing); for legend see
Text-fig. 25.

70

Atlas_2015.indd 70 25-04-2015 14:17

 . 21. , . - ( ., 1990, -
); . . 25.

Text-fig. 21. Section of Borshchov Yar, vicinity of Mogilev-Podolsky town (by Velikanov et al., 1990, with
changing); for legend see Text-fig. 25.

71

Atlas_2015.indd 71 25-04-2015 14:17

 . 22. . , ( ., 1990, ); -
. . 25.
Text-fig. 22. Outcrop near Minkovtsy village, Antonov Yar (by Velikanov et al., 1990, with changing); for
legend see Text-fig. 25.

72

Atlas_2015.indd 72 25-04-2015 14:17

 . 23. . () . (b); -
. . 25.

Text-fig. 23. Outcrops of Studenitsa Formation near Studenitsa (a) and Goraevka (b) villages; for legend see
Text-fig. 25.

73

Atlas_2015.indd 73 25-04-2015 14:17

 . 24. -
. (
., 1990, );
. . 25.
Text-fig. 24. Section of transitional Vendian-Cam-
brian deposits near Kitaygorod village (by Velikanov
et al., 1990, with changing); for legend see Text-fig. 25.

. 25. ( ., 1990, ).
: 1 , 2 , , 3 ,
4 , 5 , 6 , 7 , 8
, 9 , 10 , 11 , 12 ,
13 , 14 , 15 , , 16
, 17 --, 18 , 19 , 2021 -
, 22 , 23 , 24 ,
25 . : 1 , 2 -
; : 3 , 4 , 5 , 6 ,
7 , 8 , 9 , 10 , 11 , 12 ,
13 , 14 , 15 , 16 , 17 , 18 -
, 19 ; : 20 , 21 .
: O , S , K , Q .

Text-fig. 25. Comparison of main sections of Vendian of Podolia (by Velikanov et al., 1990, with changing).
Legend: 1 crystalline rocks, 2 breccias and boulder rock, gritstones, 3 sandstones, 4 clayey sandstones,
5 silt sandstones, 6 siltstones, 7 mudstones, 8 silt mudstones, 9 ash tuff, 10 tuffaceous mudstones,
11 limestones, 12 bentonite interbeds, 13 glauconite, 14 phosphatic mudstones, 15 pebbles of sand-
stone or siltstone 16 phosphoritic concretions, 17 calcite, exhibiting cone-in-cone structure, 18 oolites, 19
red-coloured rocks, 20-21 imprints of macrophytes, 22 imprints of Ediacaran macroorganisms, 23 bio-
glyphs, 24 Sabelliditidae, 25 microfossils. Nemerical symbols: 1 lower subformation of Grushka Formation,
2 upper subformation of Grushka Formation; Beds: 3 Olchedaev, 4 Lomozov, 5 Yampol, 6 Lyadova,
7 Bernashevka, 8 Bronnitsa, 9 Zinkov, 10 Dzhurzhevka, 11 Kalyus, 12 Pilipy, 13 Shebutintsy,
14 Kuleshovka, 15 Staraya Ushitsa, 16 Krivchany, 17 Durnyakovka, 18 Polivanov, 19 Komarovo;
Formations: 20 Okunets, 21 Khmelnitsky. Latin letters mark the contacts with deposits: O Ordovician,
S Silurian, K Cretaceous, Q Quarternary.

74

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 75

Atlas_2015.indd 75 25-04-2015 14:17

 1. INTRODUCTION

Active study of biota from Vendian paleobasins of East Europe is going on almost half of the last
century although first findings of Vendian macrofossils were undertaken in Podolia one hundred years
ago. By now a great volume of information and representative collection of Precambrian fossils was
accumulated in the wide range of state museums and research institute, particularly in Moscow, Kiev
and St-Petersburg. The results of their study could be indicative of strongly pronounced provincialism
of Vendian biota at margins of East European platform. Only five species are common among 215spe-
cies of macrofossils presumably of animal nature and ichnofossils which were discovered in White Sea
region, Southern Ural and Podolia. Such sharp differences between assemblages could be due to tapho-
nomical and paleoenvironmental features of paleobasins, and this requires special detail investigations.
However, probably in higher degree it depends on disconnection of research groups that were studying
fossils. It is obvious that without revision of taxons which were established before, subsequent progress
in research of Vendian macrobiota of East Europe is impossible. And diagnostic of modern conditions
of macrofossils collections that have provided basement for previous paleontological investigations
could be the first step to such revision.
The results of long-lasting study of Vendian macrobiota of East Europe are spread into numerous
publications, part of which were accomplished in little-known regional editions. Over the last decades
the structure and names of organizations, in which paleontological collections are stored, were trans-
formed. Registration numbers of most original specimens were changed and some of type specimens
turn out to be damaged or lost. These changes demand recording.
Present publication opens a series of reports on Vendian macrofossils of South-Eastern and North-
ern margins of East European platform and also Ural. Its main goal is consolidation of information
about all taxons of macrofossils of species rank from Late Vendian and transitional Cambrian deposits
of Middle Dniester area (Podolia) and Volhynia that were ever published in open press.
The authors consider it to be a pleasant responsibility the opportunity to express sincere grati-
tude to our predecessors and colleagues: A.A.Ishchenko, L.V.Korenchuk, V.V.Kirjanov, together with
whom weve been studing Vendian sections for a long time. V.S.Zaika-Novatsky and Yu.A.Gureev left
a good memory behind. We would like to express a deep respect to the memory of academician B.S.So-
kolov a Vendian discoverer, founder and leader of biostratigraphic research of this geological system.
L.I.Konstantinenko, an outstanding expert on Upper Precambrian and Lower Paleozoic sections
of East European platform, our friend and co-author to our deep regret did not live to see the completion
of this work (tex-fig. 1).
The study was supported by State foundation for Basic Research of Ukraine (grant no. 53/111-
2013) and Russian foundation for Basic Research (grant no.13-05-90435).

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 2. VENDIAN OF MIDDLE DNIESTER AREA

The territory of Middle Dniester area (Podolia) includes adjacent regions of Vinnitsa, Khmelnitsky
and Chernovitsky regions of Ukraine and Moldova republic. It is a part of Volhyno-Podolian upland. Its
relief is characterized as gently hilly plain, intensive dissected by river network and ravines. Vendian,
Cambrian, Ordovician, Devonian, Cretaceous, Neogene and Quaternary deposits are well outcropped
at the slopes of ravines, canyon-like Dniester valley and its left tributaries (Derlo, Nemija, Serebria,
Lyadova, Karaets, Kalyus, Ushitsa, Studenitsa and Ternava rivers) (text-fig. 2).
Thick section of Vendian and transitional Cambrian deposits in Middle Dniester area is exposed
with numerous overlapping in a wide range of outcrops and boreholes. Region of Vendian-Cambrian
(?) outcrops is spread from Kitaygorod village on the west to Yampol town on the east, extending on
more than 150 km (text-fig. 3). The section is sufficiently well studied and described in many publica-
tions (Kaptarenko, 1928; Zaika-Novatsky et al., 1968; Korenchuk, Ishchenko, 1980; Korenchuk, 1981;
Velikanov, 1985; Velikanov et al., 1983, 1990; Gureev, 1985, 1988; Aseeva, 1988a, b; Gnilovskaya et al.,
1988; Kirjanov, 1993).
GEOLOGICAL SKETCH
The regular and deep investigations of the oldest rocks of sedimentary cover of Middle Dniester
area were started from 1930syears of XXcentury. Special interest to these rocks could be explained by
excellent outcropping and requirements of USSR in minerals. At that time intensive systematic study-
ing of stratigraphy of sedimentary cover deposits of East-European platform was related to geological
surveying and searching for materials for mineral fertilizers. Long breaks in investigations at different
times were caused by political instability, wars, and economical difficulties. Podolia is located not so
far from the center of Europe. Beneficial geographic position and presence of numerous transportation
routes (railway, auto and air routes) allows to get there from any country. Unique geological structure
attracted geologists of different countries to the western part of Podolia (Middle Dniester area). At dif-
ferent periods scientists from the following countries were working here: R.Kozlovsky and others from
Poland, T.Vascautsanu from Romania, B.S.Sokolov, L.F.Lungersgauzen, G.Dickenstein, O.I.Niki-
forova and many others from USSR, A.Boucot and others from Canada. For a long time the sections
were a field of activity of mostly Ukrainian scientists. Over the last years new international projects and
agreements on studying of deposits and fossil remains from these units have been realized.
Vendian deposits of Podolia became considered as Precambrian in the middle of the 1960s. Previ-
ously they were regarded as Paleozoic (Cambrian-Silurian). Papers by B.S.Sokolov were crucial for
correct definition of the age. B.S. Sokolov was first to describe (1949-1952) Vendian on East-European
platform as a separate complex of sediments (Sokolov, 1952, 1972; Sokolov, 1964 etc.). Huge paleonto-
logical and stratigraphical information and collections on Vendian of Middle Dniester area were col-
lected in the course of geological surveying, prospecting, thematic, and scientific research works, real-
ized here with interruptions since 1960s to the beginning of 1990s and also in the beginning of 2000s.
V.A.Velikanov, P.F.Bratslavsky, V.Ya.Ivanchenko, V.S.Zaika-Novatsky, L.V.Korenchuk, E.A.Ase-
eva, V.M. Paliy, L.I. Konstantinenko, Yu.A. Gureev, A.A. Ishchenko, and others took part in these
works. As a result detailed scheme of section stratification, corresponding to demands of large-scale
mapping, was designed. There are no analogs of this scheme in other regions of East-European platform
on extent of validity at that time. High level of paleontological characterization provided world-wide
reputation and role of hypostratotype of Vendian system to Podolian sequence (Velikanov et al., 1983;
Velikanov, 1985, 2011).
Reliable findings of macrofossil in Vendian deposits of Podolian are known since the beginning of
the last century. However, the first researchers defined them as objects of inorganic genesis (rain drops,
burrows of tadpoles etc.). O.K.Kaptarenko was the first explorer who suggested an organic nature
of these problematic imprints (1928). They were interpreted as imprints of fossil jelly-fishes. There are
data on findings of fossil remains from the same stratigraphic levels in monograph by T.Vascautsanu,
which was printed in 1931. He defined the fossils as inarticulate Brachiopods (Lingula or Obolus) (cited
by Yatsenko, Trophimovich, 2001).

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 A little while later, when there was no more need for searching of materials for mineral fertil-
izers, investigations in Middle Dniester area got new directions. In the publications of that period by
L.F.Lungersgauzen (1939), sedimentary sequence was divided into two Formations Mogilev and Ush-
itsa. L.F.Lungersgauzen considered Mogilev Formation as Cambrian and clayey-shale Ushitsa Forma-
tion as Ordovician. It was later established that most part of soft-bodied imprints were contained within
deposits of Mogilev Formation. He described original sandy bubbles, which are occasionally spread
in thousands on plates of sandstones from Yampol Horizon of Mogilev Formation. He considered
these deposits to have a fluvial-glacial-alluvial genesis. Alluvial facies predominated on his opinion.
Lungersgauzen believed that an influence of running river wave on the low sandy bank had created
such bubbles, which could be compared to analogous structures on the beaches of modern plain rivers.
It must be noted, that concepts of age of sequence, containing imprints, were permanently changing
due to the absence of reliable paleontological data. Only in the beginning of the 60s of XXcentury, in
a process of compilation of State geological map in a scale of 1:200000 and initiate mapping in a scale
of 1:50000, it became possible to correlate deposits of Middle Dniester area with Vendian deposits of
other areas in East-European platform. At the same time Vendian and Lower Cambrian formations were
discovered in numerous boreholes of Volhynia.
Above mentioned type of works specifically provided new level of quality in studying of Ven-
dian deposits. The data, which gave a stimulus to paleontological study of Vendian deposits within
all East-European platform, were obtained during geological surveying and deep geological mapping
with accompanying thematic and scientific research investigations. The finding of Ediacarian fauna
representative by geologist A.P.Koval in 1967 (Zaika-Novatsky et al., 1968) in Bernashevka Beds of
Yaryshev Formation was very important. This finding became the base for subsequent detailed inves-
tigations and opened a new page in paleontological study of the region. It allowed to change the views
on Ediacarian fauna in general.
In geological-tectonical sense the region belongs to southern-western slope of Ukrainian shield.
Crystalline basement occurs on the depth from 100150m in the east part of the region (it is occasion-
ally exposed in river valleys) and up to 1.21.3km in the west part of the region.
Basement consists of suppercrust and ultrametamorphic complexes of Archaean and Palaeopro-
terozoic age, forming a row of linear and dom-like folded structures, which in turn are complicated
by north-western and north-eastern (predominant) faults, along with sublatitudinal and submeridional
ones.
The earliest stage of platform development and destruction of basement appeared in the process
of formation of Volhyn-Polessian depression (northwards from the region of investigation) on the west
slope of Ukrainian shield. This stage was connected with Early Baikalian (Dalslandian) tectogenesis.
The following activation of fault structures and the development of new ones was a result of Late Bai-
kalian (Asintian), Hercynian and Alpine movements.
The most significant evidence of Late Baikalian tectonic activity on the territory of Vendian se-
quence was Podolian fault, which was connected with the effusion of one or two covers of basalt lava.
They belong to one of the youngest trappean complexes of Volhyn-Podolia.
The complex of sedimentary cover consists of two structure stages: Vendian-Lower Paleozoic
and Mesozoic-Cenozoic. These stages differ by various structure layouts and are divided by distinct
planation surface, which was formed during long time interruption in sedimentation from Vendian (on
west part of region Silurian) up to Cenomanian. Deposits, composing lower stage, form the Podolian
monocline of north-western course, gently falling (near 1 or 10-13 m/km) south-westwards. Accord-
ing to bedding conditions and interrelations among stratigraphical units Volhyn (Lower Vendian) and
Mogilev-Podolsky (Upper Vendian), Kanilovka-Baltic (Vendian Lower Cambrian), Berezhtsy (Lower
Cambrian), Ordovician and Silurian structural substages were distinguished in the lower stage. They
are characterized by some differences of structural plans. Their boundaries are expressed by regional
interruptions. Post-Volhyn substages form Dniester pericratonic trough, representing an ancient struc-
ture, spreading along south-western margin of East-European platform. The trough was filled up by
the sediments in Vendian Early Devonian. The pericraton borders with folded Baikalids on the west.

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 Podolian monocline was constructed by above mentioned structural substages and complicated by
morphologically and genetically various structural forms (local elevations are the most common ones).
Among them elevations, closely connected with projections of buried basement relief, which formed
the cores of these structures, predominated. Amplitudes of the elevations on Vendian deposits reach
5070m and on surface of basement they are up to 120150m. All local elevations represent stamp
folds according to forming mechanism. Main phases of activity, which determined modern appearance
of the folds, were associated with Hercynian and, occasionally, Alpine movements. Local depressions,
gently sloping linear folds, flexures, normal faults, overfaults, and non-amplitude disjunctive disloca-
tions got widespread occurrence among other structural elements of Podolian monocline. In addition to
tectonic forms, structures of non-tectonic origin (folds of extrusion and draping, systems of lithogenetic
fracturing and others) are also characteristic for Vendian part of the sequence.
Such structures can be observed near the dam of Dnestrovskaya Hydroelectric Power Station and
Bernashevka village (text-fig.4). Relatively quite hydrodynamic conditions of the basin was supported
by the presence of local uplifts (islands, underwater elevation and shoals) in the beginning of Late
Vendian. The conditions around were favorable for settlement of the sea bottom by representatives of
Vendian benthic communities. Such exact conditions are characteristic for most known locatlities of
Vendian (Ediacaran) fauna (working quarry of Dnestrovskaya Hydroelectric Power Station and also
the former one on the right bank of Dniester river. Engine room of the station was constructed on the
place of this quarry. Posukhov ravine, near Vinozh village, on Lyadova river and etc.).
Mesozoic-Cenozoic structural stage contains three substages: Cretaceous, Neogene and Quater-
nary. Cretaceous structural substage is composed by deposits of Albian and Cenomanian. Its structural
plan in general is similar to that of underlying substage, but incline of bottom surface of Cretaceous is
significantly smaller (33.5m/km).
Regional structural plan of Neogene substage was substantially reconstructed. Miocene (Badenian
and Sarmatian) deposits of this substage has general inclination on south-eastern area under the influ-
ence of Prichernomorskaya (Black Sea) depression. Alpine tectonic activity was most distinctly evident
in activation of Podolian fault with regional overfaults with an amplitude up to 25m. There is also an
extensive trough of north-western spread, which is pressed to Podolian fault. It is playing a factor role
of phosphorite deposits localization.
Quaternary structural substage is polyfacial. The substage is represented by crust of weathering,
alluvial, sloping, and covering deposits. Depending on bedding conditions of pre-Quaternary deposits,
geomorphologic conditions, and facial-genetic specifics of horizons, which take part in the construction
of the substage, these horizons have different limits of occurrence. They make direct contact with any
of the underlying structural units in different regions.

STRATIGRAPHICAL CHARACTERISTIC OF VENDIAN AND EARLY

PALEOZOIC PART OF THE SEQUENCE OF MIDDLE DNIESTER AREA
Vendian and Lower Paleozoic sequence of Podolia has worldwide reputation. Since 70s of the last
century its Vendian part was considered as the most representative section of pre-Cambrian System.
Vendian was considered as its etalon on East-European platform. This section still remains important,
although Ediacarian system of Australia has got a planetary status.
Due to well outcropping, fullness of sequence, simplicity of bedding conditions, and diversity of
animal and plant fossils (Korenchuk, Ischenko, 1980; Korenchuk, 1981; Velikanov et al., 1983, 1990;
Velikanov, 1985; Fedonkin, 1987; Sokolov, 1997), Podolian section acts as a hypostratotype of Vendian
system. It did not loose its significance for interregional and global stratigraphical correlations. Each
stratigraphical unit of Vendian of Middle Dniester area has facial and lithological features, consistent
on large territory, which allow to recognize with confidence and trace these subdivisions in the distinct
sections (text-figs.3, 25). This characteristic of Podolian section allows to prepare extremely detailed
local stratigraphical chart (Velikanov, 1985; Velikanov et al., 1990).
Volyn Group consists of the oldest formations of regional sedimentary cover, distinguished as
Grushka Formation. Its spread is confined to lower parts of erosion relief of the basement. Within this

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 Formation Bakhtyn Beds, represented by mainly red-colored coarse-clastic rocks (gravelites, coarse-
grained arcose sandstones, conglomerates, breccia) were distinguished in the lower part , and Vinkovt-
sy Beds, mainly grey-colored mudstones and siltstones, in the upper part. In the south-eastern part of
the region one basalt cover occurs on the level of Vinkovtsy Beds, while there are two basalt covers in
the west part. Weathering crust is well-developed on these covers. Thickness of the Formation is vary-
ing from 010m up to 5070m (text-figs.5, 6).
Mogilev-Podolsky Group consists of Mogilev, Yaryshev and Nagoryany Formations. Terrig-
enous content of rocks from this Group mainly includes quartz and feldspars, formed after disintegra-
tion of Ukrainian shield rocks.
Mogilev Formation includes following Beds (bottom-upwards): Olchedaev, Lomozov, Yampol,
Lyadova.
Olchedaev Beds are represented by mainly light-grey, coarse- and multi-grained, arcose, often
cross-bedded sandstones and gravelites. Thickness is up to 25m.
Lomozov Beds are represented by thin horizontal interbedding of dark-grey mudstones and thin-
grained sandstones, which thickness is up to 20m. In middle part of the beds thick interbeds and units
of arcose gravelites occur (text-fig.7). Lomozov Beds represent the most ancient level of the section,
where Vendian macrofossils were recognized. Lomozov assemblage is the richest in systematic and
morphological diversity among all known in the reference section (Velikanov et al., 1983; Menasova,
2006; Sokolov, Ivanovsky, 1985a, b; Fedonkin, 1985, 1987).
Yampol Beds are represented by light-grey oligomictic (or quartz), occasionally solid, cross-bed-
ded, fine- and middle-grained sandstones, which thickness is up to 30m (text-fig.8). Numerous imprints
of Nemiana simplex Palij commonly occur in Yampol Beds. Its colony sometimes forms carpet covers
on bedding surfaces of tens and hundreds sq. meters in area (text-figs.9-11).
Over recent years paleoichnological and paleontological characteristic of Yampol Beds was es-
sentially extended due to the findings and description of forms, previously unknown within these beds.
Imprints of more than 20species of macroorganisms and ichnofossils in total were found in the upper
units of Yampol Beds (Martyshin, 2012).
Lyadova Beds are represented by greenish-grey and brown thin- and horizontal-bedded micaceous
mudstones, which are connected with underlying beds by gradual transition. The beds contain bitumi-
nized remains and fragments of films of algal (probably, fungal) origin and assemblage of microphy-
tofossils, typical for Mogilev Formation (Velikanov et al., 1983). Thickness of the beds is up to 25m
(text-fig.12).
Yaryshev Formation combines Bernashevka, Bronnitsa and Zinkov Beds.
Bernashevka Beds are composed by three units in the type sections. Lower unit (up to 10m) is
represented by solid, clayey, middle- and fine-grained, feldspar-quartz sandstones, interbedded with
siltstones. Practically all over the beds, they contain layer of bentonite clay (0.10.8m), representing
a well-marked level. Middle unit (up to 7m) consists of dark-grey and green, thin-bedded mudstones,
sometimes with thin-bedded lenses of sandstones and interbeds of bentonite clays (text-fig. 13). Oc-
casionally remains of Ediacaran organisms and accumulations of Serebrina crustacea Istch. and Eo-
holynia Gnil. algae (text-fig.14) are presented here. Upper unit (up to 5m, sometimes is absent in the
section) is represented by coarse-grained, and sometimes gravelly, arcose sandstones.V.M.Paliy and
V.S.Zaika-Novatsky described assemblage of Precambrian macrofossils from this unit for the first time
in Eurasia (Zaika-Novatsky et al., 1968; Paliy, 1976).
Bronnitsa Beds are represented in lower part (58m) by very solid pelitomorphic rocks: tuffaceous
siliceous mudstones and pelitic, massive or low-bedded tuffites of chocolate-brown color (light-green
in lower part). In the upper part of beds mudstones are chocolate-brown, fine-fractured and slightly
micaceous; they contain several thin interbeds of bentonite clay near top. These levels coincide with
color changing and are accepted as assumed upper boundary of the beds. In several outcrops representa-
tive assemblage of macrofossils was described from lower part of the beds (light-colored) (Velikanov et
al., 1983). Thickness of the beds is up to 25m.

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 Zinkov Beds gradually replace Bronnitsa Beds. The beds are composed of greenish-blueish-grey
and grey-green thin-bedded mudstones and siltstones with interbeds of clayey and carbonate sand-
stones. Occasionally they consist lenses and interbeds of phosphatic mudstones. The richest in the
Vendian sequence assemblage of microphytofossils (acritarchs, sphaeromorphytes, colonial forms and
others) is associated with Zinkov Beds. It has markers significance (Velikanov et al., 1983; Ryabenko
et al., 1976). Thickness of the beds is up to 30m.
Nagoryany Formation is divided into Dzhurzhevka Beds and Kalyus Beds.
Dzhurzhevka Beds are represented in the lower part by mainly light-grey multi-grained feldspar-
quartz sandstones, which are often interbedded with greenish-grey mudstones, siltstones and clayey
fine-grained sandstones. Layer of massive multi-grained sandstone or gravelite (up to 11.5m) is fre-
quently presented at the base of the beds. Upper part of the beds is represented by greenish-grey fine-
middle-grained micaceous sandstones, sometimes glauconitic, interbedded with siltstones and mud-
stones. In the upper part of the section accumulations of the thalli and detritus of fossil multicellular
algae Vendotaenida commonly occur (Aseeva, 1988; Sokolov, Ivanovsky, 1985a, b). Thickness of the
beds is up to 20m.
Kalyus Beds are represented by homogeneous, dark-grey, thin-bedded mudstones. Characteris-
tic feature of the beds is the occurrence of phosphorite concretions which form up to 15 levels in the
middle part of the section (text-fig.15). Kalyus Beds are connected with Dzhurzhevka Beds by gradual
transitions, which are expressed in lower part of the section by presence of interbeds and lenses of
fine-grained carbonate sandstones. Kalyus Beds have two levels enriched by remains of Vendotaenian
algae. Lower level is located in the bottom surface and has thickness near 4m. Upper level is located
near top surface and has thickness up to 5.5m. Thickness of the beds is up to 50m. Nagoryany Forma-
tion includes an assemblage of microphytofossils (Aseeva, 1988; Sokolov, Ivanovsky, 1985a, b), which
distinguishes this Formation from underlying one.
Kanilovka Group is separated from underlying Mogilev-Podolsky Group by weathering crust
(zone of sialitic clay at the top of Kalyus Beds with thickness, up to 1.0-1.5 m) and regional strati-
graphical unconformity (text-fig. 16). Group consists of four rhythms of sedimentation, and each of
them corresponds to one Formation: Danilovka, Zharnovka, Krushanovka, and Studenitsa. In each
Formation more coarse-clastic lower part with dominating sandstones and siltstones, and more clayey,
frequently multicolored, upper part of the section with distinct prevalence of mudstones, could be rec-
ognized. These parts of section are distinguished as Beds in local chart. Remains of fauna are practi-
cally absent in Kanilovka Group. However, in comparison with Mogilev-Podolsky Group degree of
bioturbation of primary sediments is considerably higher. According to the data by Yu.A.Gureev and
M.A.Fedonkin, bioglyphs are extensively represented in different levels of the section (Gureev, 1985,
1988; Fedonkin, 1987).
Danilovka Formation consists of interbedding of fine-grained sandstones, siltstones, and mud-
stones. It is divided into two parts. The lower part of the section is represented by greenish-grey fine-
grained rocks Pilipy Beds. The thickness is near 30m (text-fig.16). In the upper part of the section
more clayey rocks are disposed, mainly represented by thin interbedding of mudstones and siltstones,
multicolored or brown-colored. These are Shebutintsy Beds, which thickness is near 25m. Fragments
of volcanic glass, effusive rocks, phyllites, siliceous and jasper rocks predominate (up to 5070%) in the
content of sandy fractions of Formation. They represent products of destruction of complex of Baikal
folded platform margins (pre-Carpathian).
Zharnovka Formation, as an underlying one, is represented at the base by multi-grained, often
cross-bedded, sandstones (23m), which are replaced upwards by interbedding of greenish-grey sand-
stones, siltstones and mudstones (Kuleshovka Beds). The thickness is up to 25m. Upper part of the sec-
tion of this Formation is represented by red- and brown-colored, commonly slightly micaceous, clayey
(mudstones and siltstones) rocks (Staraya Ushitsa Beds). The thickness is up to 15m.
Krushanovka Formation is divided into two parts. Krivchany Beds occur in the lower part. They
are represented by basal fine- and medium-grained sandstones (34 m). Greenish-grey monotonous
sandstones, siltstones and mudstones (thin interbedding) occur in the upper part. Thickness of Krivch-

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 any beds reaches 30m. Durnyakovka Beds are deposited above. The beds are represented by thin
interbedding of brown-colored siltstones, mudstones (predominant) and sandstones, occasionally with
lenses of limestones and phosphate mudstones. Thickness of Durnyakovka Beds is close to 15m. Over-
all thickness of deposits of Krushanovka Formation is up to 45m.
Studenitsa Formation is represented by Polivanov beds in the lower part. These beds represent in-
terbedding of grey sandstones, siltstones and mudstones with unit of multi-grained sandstones (23m),
commonly isolated at the base. In the upper part Komarovo Beds occur. Komarovo Beds are repre-
sented by relatively homogeneous thin interbedding of mainly grey mudstones and siltstones. Total
thickness of Studenitsa Formation is up to 60m (text-fig.17).
All Formations of Kanilovka Group include macrofossils of Vendotaenian algae, different bio-
glyphs (text-fig.18) and occasionally body imprints (Velikanov et al., 1983; Fedonkin, 1987; Gureev,
1988; Menasova, 2006). Two lower and two upper Formations of Kanilovka Group contain separate as-
semblages of microphytofossils, which are available for intra- and trans-regional correlations (Aseeva,
1988a).
Kanilovka Group upwards the section passes into Baltic Group without interruption. The bottom
surface of Baltic Group corresponds to Vendian-Cambrian boundary, accepted in Ukraine, and practi-
cally coincides with location of the base of bioglyph zone Phycodes (=Trichophycus) pedum, which is
considered as international standard of Precambrian-Cambrian boundary (see, Informational report...,
Kyoto, 1992; Landing, 1994).
Baltic Group contains (bottom-upwards): Okunets, Khmelnitsky and Zbruch Formations.
Okunets Formation is represented by grey and greenish-grey mudstones with rare interbeds and
lenses of siltstones and fine-grained sandstones. Lower boundary of Okunets Formation is not dis-
tinct, while upper boundary is traced by commonly continuous layer of glauconite-quartz basal sand-
stones of Khmelnitsky Formation (text-fig. 19). Phytofossils acritarchs and macroalgae (Tirasotaenia
and Vendotaenia) predominate among fossil remains in Okunets Formation. There are rare remains of
Ediacaran macroorganisms: Kamenecia stella Gur., Ternavellus vialovi Gur. Bioglyphs (Planolites sp.,
Curvolithus sp.) occur more frequently. Sometimes representatives of skeletal fauna (Sabelliditida)
Sokoloviina sp., Parasabellidites sp. are abundant (Velikanov et al., 1983; Kirjanov, 1993). Thickness of
Formation is up 1517m.
Khmelnitsky Formation is represented by dark-grey and greenish-grey thin-bedded mudstones,
with minor interbeds of siltstones and glauconite-quartz sandstones (up to 56m); occasionally con-
glomerates and breccias (up to 0.3 m) occur there. In most full sections Khmelnitsky Formation is
divided into three units: 1)basal unit, represented mainly by sandstones 6.514.0m; 2)middle unit,
represented mainly by mudstones with limestone interbeds (up to 0.1m) 2131m; 3)upper unit, con-
sisting mainly of siltstones with minor content of sandstones 1824. In the outcrops along Ternava
river only lower part of basal unit is represented. In most of the outcrops this unit occurs under the level
of water storage reservoir.
Organic remains in Khmelnitsky Formation are more abundant and diverse then in Okunets For-
mation. Among macrofossils the following species are known: Nemiana simplex Palij, Cyclomedusa
minuta Fedonkin, Elasenia zhuravleva Gur. and Kullingia concentrica Glaessn. The following bio-
glyphs are strongly representative: Bergaueria major Palij, Didymaulichnus tirasensis Palij, Treptich-
nus bifurcus Miller, T. triplex Palij, Planolites sp. and others. Remains of Sabelliditida were found in
the middle unit. Khmelnitsky Formation (as well as Okunets Formation) is characterized by Rovno
assemblage of acritarchs (Aseeva, 1988; Velikanov et al., 1990).
Zbruch Formation differs from all other underlying parts of the section by the fact, that it is not
represented in the natural outcrops. It was discovered only in borehole sections. In Dniester area of
Podolia this Formation is distinctly separated from Khmelnitsky Formation, commonly with erosion.
Zbruch Formation is characterized by interbedding of packs of light-grey fine-grained quartz sand-
stones and multicolored mudstones (up to 70% of the volume) and siltstones. The Formation often
contains interbeds of intrastratified conglomerates and desiccation cracks and ripple marks on bedding
surfaces. Lower boundary of the Formation is diachronic. Northward and westward from studying area

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 it passes upward through the section of Baltic Group. Upper boundary of the Formation is also every-
where sharp and is marked at the bottom of basal deposits (gravely sandstones and conglomerates) of
Berezhtsy Group. Thickness of the Formation is reduced from 45m in the south sections to first meters
in the north due to the moving of its lower boundary. In sections of Dniester area Zbruch Formation is
weekly characterized by paleontological material. Here only bioglyphs were described: Planolites sp.,
Didymaulichnus tirasensis Palij, Treptichnus bifurcus Miller (Gureev, 1988). Assemblage of acritarchs,
contained in Zbruch Formation corresponds to Lontova Horizon of western regions of East-European
platform, which Russian researchers consider as age analog of Tommotian stage of Lower Cambrian
in Siberian platform. V.V.Kirjanov (Drigant et al., 1982) established, that Lontova Horizon had pre-
Tommotian age, while stratigraphical interval, corresponding to Baltic Group, was missed in general
stratigraphical scale of Vendian-Cambrian of Russia. Kirjanov believed that this missed interval cor-
responded to geological Series, probably, even System. This issue should be solved by further research.
Next structural substage is formed by Berezhtsy Group of Lower Cambrian. The Group is repre-
sented by light-grey mainly quartz sandstones. Its structure plan is considerably different from Baltic
one due to re-location of sedimentation areas and discontinuance of contacts with basins of Central
regions of East-European platform.
Total thickness of Cambrian deposits on the west part of region is more than 450m (borehole
Bouchach).
Cambrian deposits are overlaid with large interruption by Ordovician deposits represented by two
Formations Goraevka and Suboch, with total thickness less then 10m.

DESCRIPTION OF KEY OUTCROPS

Outcrop 1
Occurs on north margin of Vysshy Olchedaev village on left slope of Lyadova rivers valley (text-
fig.20).
Paleoproterozoic fractured and cataclastic granites and migmatites are exposed in an old quarry.
Their eroded surface is overlying by Vendian deposits (bottom-upwards):
Grushka Formation is represented by:
1) brown breccia, consisting of fragments (up to 15cm) of migmatites, gneisses, altered effusive
rocks, quartz, and feldspar, which are cemented by sandy-clayey mass (up to 50%) 0.50.7m;
2) multicolored (reddish, greenish) detrital gravelites up to 0.5m;
3) dark-brown, solid breccia, consisting of fragments (34cm) and sandy-clayey cement mass (up to
70%) up to 0.7m;
4) greenish-grey, multi-grained, quartz-feldspar sandstones with beds and lenses (up to 0.3m) with
admixture of fine-grained breakstone fragments of basement 1.21.5m; interbedding of feldspar-
quartz grey, brown-grey, coarse to medium-grained sandstones, with lenses of light-grey fine-
grained grey solid sandstone and solid grey multi-grained sandstones with interbeds of fine-pebbly
conglomerates. An the top sandstones are intensively colored by ochre 1.61.9m.
Olchedaev Beds:
Feldspar-quartz grey coarse-grained sandstones with interbeds and lenses of gravelites and fine
pebble (0.150.5m). The clastic material is weakly rounded and sorted. Cement consists of clayey sili-
ceous mass 3m. The section is covered by screes of Cretaceous (Cenomanian) rocks.
Outcrop 2
Quarry located on the left bank of Dniester river near Dnestrovskaya Hydroelectric Power Station.
Pink-grey Paleoproterozoic granites (2050Ma), constructing local projection of the basements surface,
up to 40m in height, over surrounding areas of pre-Vendian relief, are exposed here at 500600m.
They are overlaid by massive, silicificated, multi-grained, laminated at the upper part of the section,
sandstones of Yampol Beds of Mogilev Formation, forming anticline drape (amplitude is up to 15m).
The thickness of Yampol Beds differs on the dome of uplift (34m) and on the slopes (up to 1214m).
Imprints of colonies of macroorganisms, mostly Nemiana simplex Palij, occur in the upper part of beds,
in tabulated sandstones, sometimes as solid carpets (text-figs.10, 11).
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 Lyadova Beds (arenaceous thin-bedded siltstones and mudstones, up to 7m), overlaying Yampol
Beds, are outcropped on eastern wall of the quarry. Eroded surface of Lyadova Beds is covered by al-
luvium deposit of the terrace above flood-plain (text-fig. 12).
In the previously buried and flooded part of the quarry, as well as on the right bank of Dniester
river, during the construction of hydroelectric station, Lomozov Beds (dark-grey mudstones with in-
terbeds of sandstones), adjoining the slopes of granite projection, were discovered under Yampol sand-
stone. The beds contain assemblage of various macroorganisms and bioglyphs, in which many forms
are common to Ediacaran, White Sea, and Newfoundland fossil assemblages. The Podolian Vendian
sequence is lowermost and the richest stratigraphical level, where macrofossils were found (text-fig.7).
At the present time fragments of the Lomozov Beds section are partly exposed at the north-western
and southern walls of the quarry (from 3 to 5m are seen).
Outcrop 3
Located near Bernashevka village, 1 km north of the church, ravine on the left slope of valley of
Zhvan river. The section of Vendian deposits is outcropped here, which is heightened the sequence of
the Quarry near Dnestrovskaya Hydroelectric Power Station (bottom-upwards):
Lyadova Beds: green thin-bedded micaceous mudstones. Visual thickness was 23m (at the pres-
ent time this part of the section is covered by diluvial and proluvial deposits);
Bernashevka Beds: at the bottom interbedding of grey-green sandstones and mudstones, up-
per green mudstones. Imprints of Cyclomedusa plana Glaessner were found on surface of sandstones.
Incomplete thickness is 2m;
Bronnitsa Beds: coarse-tabulated lilac (at the lower part light-green 1m) tuffaceous mudstones
(tuffites). Uncertain imprints of Ediacaran macroorganisms occur in light-colored mudstones. Thick-
ness is 56m.
Outcrop 4
Southern outskirt of Zhvan village. Ravine, which cut the left slope of the valley of Zhvan river
near 0.5km northwards from outcrop 3. Following beds are outcropped here (bottom-upwards):
Bronnitsa Beds:
1) massive coarse-tubulated lilac tuffaceous mudstones, light-colored at the bottom (1.01.5m), on
bedding surfaces of which sometimes imprints of non-skeleton fauna occur. Visible thickness is 5m;
2) fractured chocolate brown mudstones. Visible thickness is 5m.
Boundary with Zinkov Beds coincides with sharp color change of rocks and appearance of thin
(12cm) interbeds of betonite clay. Thickness of outcropped Bronnitsa Beds is about 10m;
Zinkov Beds:
3) brown-green micaceous fine-fractured mudstones. Visible thickness is about 15m.
Upper, Dzhurzhevka Beds are well outcropped in right slope of the ravine:
4) grey-green fine-grained and clayey sandstones are interbedded with siltstones 4m;
5) dark-grey and brown mudstones and siltstone with interbeds of clayey fine-grained sandstones and
member (1m) of yellowish light-grey siltstones and mudstone in the upper part 67m;
6) solid medium-grained quartzit-like sandstones, often calcareous (0.52.0m).
In the ravine exotectonic fold of extrusion is well outcropped, which consists of Bronnitsa and
Zinkov Beds. The fold has extension, which coincides with spread of the ravine.
Outcrop 5
It is located 3 km on north-east from Kurazhin village near mouth part of Kalyus river.
On the rocky right bank of deep scour of right valley slope, near Vila creek Nagoryany Formation
section is outcropped on the distance of 300m (bottom-upwards):
Dzhurzhevka Beds:
1) interbedding of light-grey, greenish-grey fain-grained sandstones with siltstones (predominant)
2.5m;

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 2) thin-bedded greenish light-grey sandy siltstones, containing lenses of siltstones and mudstones in
the upper part 1.3m;
Kalyus Beds:
3) thin interbedding of dark-grey mudstones with fine-grained sandstones which contains numerous bi-
tuminized accumulations and single thalli of multicellular algae (Vedotaenida) and its detritus 0.6m;
4) dark-grey and grey homogeneous shaley mudstones, with phosphoritic concretions in the upper
part of the sequence, distributed as horizontal rows 1820m.
Marls with inclusions of flints occur upwards (Cenomanian stage).
Outcrop 6
It is located on northern outskirt of Shebutintsy village, right valley slope of Danilov creek. Fol-
lowing beds are outcropped in the scour with abrupt slopes:
Kalyus Beds:
1) dark-grey, greenish-grey, thin-bedded mudstones 10m.
Pilipy Beds:
2) light-green siltstones (predominant) interbedded with aleuritic sandstones and silty mudstones,
dispersed debris of Vendotaenia algae and biogliphs occur in the upper part 4m;
3) light-green silty mudstones (predominate) interbedded with siltstones 2.8m;
4) mudstones (predominate) interbedded with siltstones and sandstones. The rocks are tabulate,
solid, massive, dark-green, yellowish-grey, containing brown members in the middle with interbeds
(0.1m) of bentonite-like clay 5.76.0m;
Shebutintsy Beds:
5) pink-grey and brown cross-bedded fine-grained sandstones in bottom part 1.3m;
6) greenish-grey horizontally-layered sandstones interbedded with green siltstones 1.5m;
7) greenish-grey mudstones with interbeds of brown and lilac mudstones; greenish-grey horizon-
tally-layered siltstone and sandstones (1.5m) occur in the middle part, ichnofossils are rarely ob-
served 4.5m;
8) pink-grey sandstones interbedded with brown mudstones and siltstones 3m;
9) brown slightly micaceous mudstones, sometimes with interbeds (up to 25cm) of pink sandstones
and siltstones 4.7m;
10) thin interbedding of mudstones (predominant) and siltstones; rocks are mostly brown and lilac
in color 3.4m;
11) thin (up to 2cm) interbedding of brown and grey-green mudstones and siltstones 3.0m;
Kuleshovka Beds:
12) at the base (0.5 m) brown-grey, fine-grained, cross-bedded, micaceous sandstones occur; up-
wards it is changing by dark-brown, greenish, micaceous siltstones and silt-sandstones 3.9m.
Upper part of the section is covered by white marls of Cenomanian stage.
Outcrop 7
It is located on the left bank of Dniester river, 1.9km upstream the Danilov creek, 1.0km south-
wards of Berezovka village. Following beds are exposed in the deep ravine with abrupt slopes (bottom-
upwards):
Kalyus Beds:
1) dark-grey thinly laminated mudstones, rarely with interbeds of black limestones and phosphatised
mudstones (25 10.3%). They contain interbeds of calcite with cone-in-cone structure and some
levels of concretionary phosphorites with polymetallic mineralization at the holes of compaction
(desiccation). At the top surface (3m) rock color changes into light-greenish-grey (specific crust of
weathering). The boundary with overlying deposits of Kanilovka Group is lithologically sharp with
signs of a break (rocks are colored by ochre at the contact). Visible thickness is 1012m.
Pilipy Beds:
2) light-greenish-grey, thinly-horizontal-bedded (rarely cross-bedded) silt-sandstones 1.5m;
3) greenish-grey, micaceous thin-bedded mudstones with accumulations of bituminized debris of
Vendotaenia algae 4.4m;

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 4) greenish-grey mudstones (predominant) interbedded with grey slightly micaceous siltstones
3.8m;
5) interbedding of mudstones (predominan at the lower part), siltstones, and sandstones. In the upper
part of the member fragments of Vendotaenia thalli occur 4.3m;
Shebutintsy Beds:
6) lilac and brown mudstones interbedded with pink-grey, coarse-grained, polymictic sandstones
(20%) 3m;
7) interbedding of greenish-grey lilac siltstones with sandstones (up to 0.9m) 3.4m;
8) interbedding of light-green and brown mudstones and siltstones (up to 20 %) 7.5m;
9) brown, slightly micaceous mudstones with interlayers of pink-grey, coarse-grained sandstones
(0.10.2m) and light-green siltstones (up to 2.5cm) 4.6m;
10) brown thin-bedded mudstones 5.1m;
Kuleshovka Beds:
11) at the base (2 m) brown-grey, coarse-grained, cross-bedded sandstones, with small pebbles of
green mudstones occur, in the upper portion green and brown siltstones with thin cross-bedding
3.7m;
12) greenish-grey sandstones (up to 20%) interbedded with green, bluish-grey, thin-bedded mud-
stones and siltstones. In middle part light-green, plastic, bentonite-like clay (8cm) as well as debris
and fragments of Vendotaenia algae occur 7.6 m. Upwards siliceous structures of Cenomanian
stage occur.
Outcrop 8
It is located 3 km westwards from Goraevka village, on the left slope of Dniester river valley, near
cave monastery (text-fig.23). Deposits of Studenitsa Formation are outcropped along the rocky scarp
slopes, at the distance of 1.52km (bottom-upwards):
Polivanov Beds:
1) interbedding of grey mudstones, siltstones, and sandstones (last-mentioned are predominant)
7m;
Komarovo Beds:
2) interbedding of dark-grey mudstones (predominant), siltstones and sandstones. At the base (0.3m)
light-grey silty sandstone occurs 20m.
Upwards Upper Ordovician is deposited with gap and evidences of erosion:
Goraevka Formation:
3) light-grey, carbonate-quartz, coarse-grained, solid sandstones with numerous remains of brachio-
pods, trilobites, cephalopods, gastropods, pelecypods, crinoids, bryozoans, rugose and tabulate cor-
als up to 6m;
Suboch Formation:
4) grey massive limestones with rare fauna up to 2m.
Outcrop 9
It is located on the left valley slope of Ternava river, near the bridge on the road, which leads from
Kitaygorod village to Demshin village (text-figs.19, 24). Starting from the base of the slope following
beds are outcropped (bottom-upwards):
Studenitsa Formation:
1) interbedding of mudstones with grey, greenish-grey, slightly micaceous, thin-layered siltstones
(215cm). Bioglyphs, imprints of algae and films of organic matter are represented here 4.5m;
2) greenish-grey and grey mudstones, thinly tabulated at the lower part (2m) with interbeds of silt-
stones (up to 20cm) and light-grey, fine-grained, calcareous sandstone (up to 70cm). Clasts (up to
2cm) of gravel and mudstones occur at the bottom and in the middle part of the member. Layer of
light-grey fine-grained sandstone (0.7m) occurs at the top surface 4.3m.
Upwards following beds are outcropped at the wall of small no longer functioning quarry, at the
distance about 20m above the road:

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 Okunets Formation:
3) grey, bluish or greenish-grey, thin-layered mudstones, solid in the upper part with ohre on bedding
surfaces. Debris, fragments, and imprints of Vendotaenia along with Tirasotaenia remains occur in
the lower part. Sabelliditids (Sokoloviina sp.) occur in the upper part 6m;
Khmelnitsky Formation:
4) glauconite-quartz dark-green, fine-grained, carbonated sandstones with pebbles of siltstones
0.3m;
5) greenish-grey, thin-layered, slightly micaceous siltstones interbedded with light-grey, sometimes
greenish-grey, glauconite-quartz sandstones, which have rounded type of jointing. Phycodes pedum
bioglyphs are observed closer to bottom surface 1.7m;
6) upwards the slope is covered by turf, but nearby the closed areas greenish-grey, thin-bedded mud-
stones, sometimes with bioglyphs of Phycodes pedum and Didymaulichnus sp., can be observed.
Greenish-grey, fine-grained, quartz sandstones with glauconite occur in the middle of the member
0.9m;
Next to turfed area (0.8 m) greenish-grey, glauconite-quartz, fine-grained sandstones with thin
interbeds of mudstones and siltstones of darker color are outcropped. The sandstones contain bioglyphs
Bergaueria major Palij 0.4m;
7) upwards, after the closed area greenish-grey, thin-layered mudstones sometimes with glauconite.
Closer to the bottom, there are interbeds (57cm) of glauconite-quartz, multi-grained sandstones,
which are interbedded with interlayers of mudstones; bioglyphs Planolites sp. and others are devel-
oped on bedding surfaces 0.9m.
Upwards by the slope Cambrian (?) deposits are overlaid by diluvial deposits with fragments and
debris of Ordovician sandstones and limestones along with Silurian limestones. Bedrock exposure of
these rocks occurs up to the upper edge of abrupt slope of the valley.

MAIN LOCALITIES OF MACROFOSSILS AND MODERN CONDITION

OF THE SECTION
Thirty-forty year ago the section of Vendian in the Middle Dniester area was excellently out-
cropped almost without interruptions. Construction of the dam of Dnestrovskaya Hydroelectric Power
Station led to unfortunate results: canyon of Dniester river lost near 60meters of height due to the rising
of water storage reservoirs level. Consequently, some stratotypes of geological section turned out to
be in the flooded zone, while some other stratotypes became unreachable, because of the covering with
slope deposits. On the other hand, a lot of new outcrops appeared along the banks of Dniester water
storage reservoir due to abrasion.
As a result of many years of research main localities of Vendian macrofossils (Metazoa, prob-
lematics, algae and trace fossils) were discovered. The most important and favorable places for search-
ing for Vendian fossil remains are interbedding of clayey silty and sandy interbeds in the Lomozov and
Yampol Beds of Mogilev Formation, Mogilev-Podolsky Group. Sometimes imprints can be found in
interbeds of mudstones (Bronnitsa Beds) of Yaryshev Formation. Imprints and trace fossils are frequent
in Studenitsa Formation of Kanilovka Group (Komarovo Beds). Significantly, findings in Podolia are
confined to slope facies of paleoelevations, which were discovered as a result of geological prospecting
and searching investigations as early as in 1960s 1970syears.
Such localities occur in natural outcrops and artificial sections on banks of Ternava river near
Kitaygorod village (Vendian and Cambrian?), near Bakota cave monastery (Studenitsa Formation). The
richest locality was discovered in the quarry near Dnestrovskaya Hydroelectric Power Station, where
two levels of findings were recognized (in Lomozov and Yampol Beds). Recultivation was started at
the quarry, which can destroy an access to the most perspective locality of Vendian macrofossils in
Ukraine. After discontinuance of water pumping, flooding will reach the level of Yampol Beds. The
recultivation of quarry slopes can completely cover the section of Lyadova Beds and lower part of Ber-
nashevka Beds. According to our suggestion, a part of the slopes and quarry walls are supposed to be
left outcropped in order to have an opportunity for their investigations in the future.

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 Remarkable locality occurs in outskirts of Mogilev-Podolsky town in Borshchov Ravine. The sec-
tion along the creek starts with upper Lomozov Beds and ended by lower part of Bronnitsa Beds. Yam-
pol and Lyadova Beds are badly outcropped there. Imprints and trace fossils were found in Bernashevka
and Bronnitsa Beds of Yaryshev Formation. Other localities expose the sandy facies of Lomozov Beds
near Popelukha village, and Bronnitsa Beds of Yaryshev Formation near Yastrebna village.
Less productive localities are not characterized by diverse findings and often occur in inaccessible
regions (dirt roads become totally impassable in rain seasons). Part of already known outcrops fall into
flooded zone of water storage of Dnestrovskaya Hydroelectric Power Station.
At the present time scientists from different fields of science and geology propose to include the
middle part of Dniester valley, where reference sections of Vendian, Lower Cambrian, Silurian and
Lower Devonian of East-European platform are outcropped, to European list of geological heritage.
In 2010 International symposium on protection of Geological heritage was organized and successfully
hold. Decision of the symposium also supported this idea. Podolian section can be estimated as one
of the most important stratigraphical sequences of global value for definition of the boundaries and
correlation of deposits. Studied region is a polygon for international geological excursions, seminars,
symposiums and a base for holding work focused on realization of basic scientific projects; present
work could serve as such an example. In Podolia it is possible to get factual material for paleontological,
stratigraphical, facial, and other kinds of geological researches.
Middle part of Dniester valley is partially included in the list of geological monuments Geological
monuments of Ukraine. Great numbers of geological objects, represented in this region, of different
types, for example, stratigraphical, paleontological, mineralogical, and others, define geological unique-
ness of Podolia. They could become a base of geological park of international scale as a result of their
revision, organization of protection and proper management.
Urgent preparation of presentation for foundation of geological or natural park (reserve or pre-
serve) on territory of western part of Vinnitsa region in Dniester valley is needed. The main task is to
obtain conservation of the quarry near Dnestrovskaya Hydroelectric Power Station with minimal recul-
tivation, which create danger of whole artificial elimination of the unique Vendian section.

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 3. HISTORY OF THE STUDIES OF MACROFOSSILS (METAZOA,
PROBLEMATICS AND TRACE FOSSILS) IN THE VENDIAN
AND LOWER CAMBRIAN DEPOSITS OF SOUTH-WESTERN
MARGIN OF THE EAST EUROPEAN PLATFORM

The beginning of the reported investigations should be considered in 1916, when in the article of
the famous Ukrainian geologist A.V. Krasovsky, published in Notes of the Imperial Society of Natural
Science, Anthropology and Ethnography, the mysterious structures on the surface of Silurian sand-
stones in the Podolian Dniester area were first mentioned. A.V.Krasovskii (1916) himself supposed that
it represented the fossil imprints of the raindrops.
At the end of 1920s O.K.Kaptarenko (1928) started to study those problematic imprints from sand-
stones of Ozarintsy near Mogilev-Podolsky town on Dniester river (now Vinnitsa region of Ukraine).
In this research he convincingly criticized the hypothesis of their inorganic (meteorological) origin, and
suggested an idea that they might belong to ancient organisms.
The Silurian age of the above-noted deposits should not confuse the readers. According to the
ideas of that time the siliciclastic sedimentary rocks of the Middle Dniester area (and also volcanic-
sedimentary rocks in the lower part of the section) were considered as a lower unit of the thick Paleozoic
Group, exposing in the valleys of Dniester river and its tributaries eastward of a line Ternopol Ivano-
Frankovsk. It was also considered as composed mostly by the well paleontologically characterized Si-
lurian and Devonian sediments.
It is unlikely that the article by O.K.Kaptarenko mentioned above attracted a lot of attention from
the professionals at that time. Many years had to go by before, on the one hand, the true age of depos-
its under consideration was established, on the other hand seemed unbreakable understanding of the
lifelessness of the Precambrian was shaken and, finally, when the various problematics were no longer
considered as some geological curiosities and become the object of thorough in-depth study. Neverthe-
less, O.K.Kaptarenkos thorough research was favorably differed from the others, rather superficial as-
sumptions about the nature of these imprints that continued to appear in the literature after the release of
her paper, until the early 60s of the last century (traces of air bubbles L.F.Lungersgauzen (1939); traces
of drops falling at one point from the overhanging rock M.F.Staschuk (1958); traces of burrowing of
the primary chordate animals or crustaceans A.N.Voznesensky (1956)). In particular, Olga Kaptarenko
expressed an opinion that they belong to jellyfish on the basis of the burial characteristics, morphology
of the imprints and mainly using the fundamental work of Charles Walcott on fossil jellyfish (Walcott,
1898). When we consider that the first explorers of the classical Ediacaran fauna R.Sprigg, M.Glaessner,
B.Daily and M.Wade (Sprigg, 1947, 1949; Glaessner, Daily, 1959; Glaessner, Wade, 1966 etc.) had at-
tributed much of the described forms to Medusoids, we should pay tribute to her insight. But again we
should notice that according to the ideas of that time and under that conditions the research in this area
were not of the big current interest and were stimulated practically by nothing, but the common curiosity.
Fundamentally the situation began to change after the World War II, and, as often happens, the
practical needs played a significant role. These factors were the exploration of the Lviv- Volhyn coal
basin, as well as the geological mapping of the territory of the USSR on the scale of 1:200000 and
1:50000 in western Ukraine, including Podolsky slope of the Ukrainian shield adjacent to the Pre-
Carpathian and Pre-Dobrudzhinsky trough. They led realization of the extensive and comprehensive
investigations including geological mapping and prospecting-exploration drilling that allowed to clarify
the stratigraphic division of the sequences and the age of the rocks composing it. By the early mid
1960s the presence of the Cambrian deposits besides of the previously known Devonian and Silurian-
Ordovician ones was established for the section of Podolian Dniester area. The Precambrian (Riphean
and Vendian) age of the underlying siliclastic strata that overlie on the crystalline basement was sub-
stantiated (Shulga, 1952 and others). Around the same time, in light of the numerous data, the theory
about the Late Precambrian representing the period of the unique organic world development, which
included highly enough organized forms of fauna, finally started to dominate (Glaessner, 1959). And its
sedimentary deposits became the promising object of paleontological exploration and research.

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 The Geology Department of the University of Kiev became one of the centers of such research
in the Ukraine. Here in the mid 1960s under the guidance of V.S.Zaika-Novatsky, associate profes-
sor (later professor), the expedition in Dniester area was active within the scientific research sector.
Among others it has a purpose of target search of any possible evidences and signs of the existence
of ancient organisms in the Precambrian deposits. As usual, a large part of the expedition team was
represented by the students of Geological department. Valery Semenovich Zaika-Novatsky attract-
ed many of them to scientific work from the very beginning of their studies at the University. Thus,
in the season of 1966 the main labor power in field trips were a graduate student L. Konstantinen-
ko, B. Sukach who finished the second grade, V.Aleshin, V.Burmistrov, V.Gritsenko, A.Matvienko
and V.Paliy who all finished the first grade.
The study area of the expedition was the middle steams of Dniester river and its numerous left-
bank tributaries Rusava in the east and the Golden Lipa in the west. Deep (up to 100m) erosional
trench created excellent conditions for studying the whole section from Quaternary deposits to the
crystalline basement, although the main focus of research remained the Lower Paleozoic and Upper
Precambrian deposits. And in the latter deposits significant place was occupied by shallow marine fa-
cies the most promising in terms of detection of fossil organic remains. By this time (1966) V.Zaika-
Novatsky had already reported the finding in the Bronnitsa Beds of Vendian at the Yastrebna settlement
on Lyadova river of specific small rounded imprints of sheath type, which he called Bronicella podolica
(Zaika-Novatsky, 1965) that is the most close to Beltanelloides of B.S.Sokolov. V.V.Kirjanov (1968)
described Sabellidites cambriensis from Khmelnitsky Formation (Lower Cambrian) on Ternava river,
previously known from the Lower Cambrian sediments of the Baltic Group of the Baltic. M.B.Gnilovs-
kaya (1971) described the algae Vendotaenia and Tirasotaenia from Sokoletsky and Komarovo Beds of
Kanilovka Formation of Vendian Dniester area. Later L.V.Korenchuk and A.A.Ishchenko got involved
in the study of these and other mainly plant remains (Ishchenko, 1983; Gnilovskaya et al., 1988). The ac-
ritarchs represented a separate area of research microscopic capsule-shaped structures presumably of
plant (algal) origin. E.A.Aseeva (1976, 1988b etc.) made a significant contribution to these studies. Then
the real sensation was the finding of Cyclomedusa plana imprint basic form of Ediacaran fauna in
Bernashevka Beds of Yaryshev Formation (Serebria settlement on the western outskirts of the Mogilev-
Podolsky city) (Zaika-Novatsky et al., 1968). V.M.Paliy was recommended to focus on the searching
and studying of the problematic imprints from Yampol sandstones of Vendian Mogilev formation that
were mentioned at the beginning of this article. As a result of thorough preparation of the material from
more than 20 localities he was able to refute all the hypothesis of its non-organic origin (see above). He
took into account a number of individual characteristics and was able to prove the belonging of these
imprints named Nemiana, primitive sedentary soft-bodied organisms that inhabited the relatively shal-
low waters of the Vendian marine basin (Zaika-Novatsky, Paliy, 1968; Paliy, Konstantinenko, 1968).
Later, the same author described a new species of the genus Cyclomedusa Sprigg Cyclomedusa sere-
brina (Paliy, 1969), and opened a completely new, previously unknown forms of the primitive fossil
organisms Tirasiana in Bernashevka Beds of Yaryshev Formation of Vendian on the Moldavian bank
of Dniester river (Paliy, 1976; Paliy et al., 1979).
At that time it also began to be possible to extend the understanding of the Vendian organic world
by studying trace fossils (ichnofossils) from the most primitive locomotion (crawling) tracks on the
surface of the muddy layer to rather complex sculptured ones, such as Harlaniella, that was found by
B.S.Sokolov in the sediments of Komarovo (Studenitsa) Formation at the entry of Ushitsa river area
(Sokolov, 1972), and Palaeopascichnus, discovered and described by V.M.Paliy from the same horizon
at Molodovo-Chernovitsky region (Paliy, 1976; Paliy et al., 1979). L.I.Konstantinenko, the leading ex-
pert on the geological section of Dniester area, handed over for description the specimens of trace fos-
sils Bergaueria and Didymaulichnus from the Khmelnitsky Formation at the entry of Ternava river area
to V.M.Paliy (Paly, 1974, 1976; Paliy et al., 1979). The localities opened by this researcher became the
source of the new paleoichnological findings. Ph.D thesis of V.M.Paliy (1975) Fossil remains of Meta-
zoa and trace fossils in the ancient siliciclastic strata (Vendian Lower Cambrian) in Podolian Dniester
area, defended at the Institute of Geological Sciences of the Academy of Sciences USSR was concern
with the analysis and synthesis of all materials mentioned above. It was the first thesis work in this area

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 in the world (Paliy, 1974). In addition to monographic descriptions of all known at that time from these
deposits body fossils (imprint) of suggested Metazoa and trace fossils, considerable attention in the
thesis was given to determination of their stratigraphic, paleogeographic and evolutionary-biological
significance. It should be noted that even at that time V.M.Paliy warned against jumping to a conclusion
and superficiality that sometimes happen to even respected researchers, while they are getting to know
some of the Precambrian problematics. For example, he showed that the structures from Bronnitsa Beds
of Vendian of Dniester area, mentioned by honored B.S.Sokolov (1972) as medusoids Charniodiscus
planus, Planomedusites grandis and Medusinites patellaris, in reality represented weathering struc-
tures on the lithological heterogeneities on the surface of samples. The conceptual issues of the thesis of
V.M.Paliy were published in two collective monographs (Paliy, 1976; Paliy et al., 1979), the second of
which was subsequently republished in English in Warsaw (Paliy et al., 1983).
The fossil search in the Vendian deposits of Dniester area was successfully continued by M.A.Fe-
donkin (1983, 1985a, b). He was able to identify some common Ediacaran forms (Conomedusites, Dick-
insonia, Ediacaria, Tribrachidium), as well as a number of new, previously unknown species of Meta-
zoa in Lomozov Beds of Mogilev Formation in the construction site of Dniester Hydroelectric power
station. Later Y.A.Gureev (1983, 1985, 1987, 1988) was studying the paleontology of Vendian deposits
of Dniester area, and in recent years A.Sh.Menasova (Menasova, 2003a, b, 2006) and V.P.Gritsenko
(2009).
All of the above-mentioned findings had epoch-making significance to recover the early stages of
the organic life evolution on the Earth based on the material of southwestern margin of the East Europe-
an platform, global stratigraphic correlations and conformation of the outcrop of Upper Proterozoic and
Lower Paleozoic sediments of Dniester area as a standard (Velkanov, 2011). We can also note that the
paleontological findings in Vendian of the Middle Dniester area to some extent served as a benchmark
for further studies by Soviet geologists and paleontologists, during which the Vendian biota assemblage
in the north of the East European Platform (White Sea coast) was soon discovered and described (Keller
et al., 1974 and subsequent publications). Some typical Podolian forms of macroorganisms and trace
fossils (Nemiana, Tirasiana and Palaeopascichnus) were identified in Upper Precambrian sequences
in sufficiently distant regions: in Ust-Pinega Formation in the north of the East European platform, in
Asha Group of the Southern Urals, in Khatyspyt Formation of northern Yakutia, in Dabis Formation of
Namibia, in Rawnsley Quartzite of South Australia (Becker, 2010; Paliy, 1987; Fedonkin et al., 2007).
Dniestrian sections of Vendian and Lower Cambrian and localities of organic remains became im-
portant objects of geological excursions for participants of numerous scientific conferences, symposia
and sessions, among which was the III All-Union Seminar on trace fossil (Kamenets-Podolsky, 1973),
Session on the stratigraphy of the Upper Precambrian (Riphean) of the Russian Platform (Kamenets-
Podolsky Kishinev, 1974), the Third International Symposium on the Cambrian system (Novosibirsk,
1990) etc.

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 4. ATLAS OF MACROFOSSILS FROM UPPER VENDIAN
AND LOWER CAMBRIAN OF MIDDLE DNIESTER AREA
AND VOLHYNIA
Summary report contains selected information on all known species of Vendian and Early Cam-
brian macrofossils of Middle Dniester area (Podolia) and Volhynia: description, images of most impor-
tant specimens, data on location of these specimens, current place of its storage and valid collection
number. Nomenclative types of the species and specimens, which do not belong to type category, but
their photographs were used for illustration of the species descriptions, are considered as important
here. Many species belong to monospecific genera and it can be tricky to distinguish among generic
and specific characters. Therefore, the diagnosis of some genera and generic comparisons, which can
be found in publications on fossils of this region, are also represented in this text. The descriptions are
provided in limited extent, only diagnosis (for genera), description, comparisons and remarks
sections are provided. Besides, we predominantly used early and mainly original authors versions,
which more truly reflect conception of a researcher, who established a new taxon or determined an af-
finity of found fossils to already known taxa.
Material is arranged in three sections according to initial visions of nature of fossils. In each sec-
tion taxons are provided in alphabetical order without determination of over-generic categories. This
was motivated by uncertainty of systematic position due to the incompleteness of material, question-
able interpretations, unsettled opinions etc. Information on macrofossils of Ediacaran type preservation
(positive imprints on the bottom of the layer without any traces of organic matter) is provided in the
first section. They were interpreted by authors of original descriptions as body remains of Metazoa.
Information on organic and agglutinated inhabited tubes is also included. The second section contains
data on ichnofossils and problematic bioglyphs, described as living, feeding and locomotion traces of
multicellular animals. In the third section diagnoses and descriptions of suggested plant remains, pre-
served as organic films (phytoleims ?) are provided.
Used abbreviations:
KTSNU Geological Museum of the Taras Shevchenko National University of Kiev
NMNH Geological Museum of the National Museum of Natural History at the National Academy of Sciences
of Ukraine
IGS Institute of Geological Sciences, National Academy of Sciences of Ukraine
PIN A.A. Borissiak Paleontological Institute, Russian Academy of Sciences

MACROFOSSILS DESCRIBED AS BODY REMAINS OF METAZOA

Genus Beltanella Sprigg, 1947

Beltanella velikania Menasova, 2003
Pl. I, fig. 1
D e s c r i p t i o n (Menasova, 2003a). Flat, rounded body, which does not go beyond the substrate
plane (negative epirelief). The small depression occurs in the center. The imprint is separated from the
substrate by the groove. It is possible that the edges of an umbrella were folded over during the lifetime
of an animal.
C o m p a r i s o n s (A.Sh. Menasova, herein). It differs from Beltanella gilesi in the presence of
central depression and uneven line of umbrellas edge.
R e m a r k s (A.Yu. Ivantsov, herein). In paper by A.Sh. Menasova holotype has number -83501-,
the new one KTSNY, specimen no. 17134 (17142 counterpart).

Genus Bessarabia Gureev, 1988

D i a g n o s i s (Gureev, 1988). Flatten, elongated, biradially or bilaterally symmetric, elongate-
ovoidal body. Peripheral and central zones separated by a groove are clearly distinguished.

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 Bessarabia bukatchuki Gureev, 1988
Pl. IV, fig. 8
D e s c r i p t i o n (Gureev, 1988). Central zone is irregularly undulating and rather compressed in
the middle. Peripheral zone has approximately equal width and is divided by transverse grooves into
segments.

Genus Bronicella Zaika-Novatsky in Zaika-Novatsky et Palij, 1968

D i a g n o s i s (Paliy, 1976). Small round imprints with convex, umbrella-like or flatten shape;
with consolidated glazed smooth or sculptured surface with very thin wrinkles and folds.
C o m p a r i s o n s (Paliy, 1976). Bronicella is the most similar to the imprints of Beltanelliformis
Menner (= Beltanelloides Sokolov editors note), but differs from them in more convex umbrella-like
shape, smaller (3-4 times) sizes and absence of numerous concentric folds on the periphery.
Bronicella podolica (Zaika-Novatsky, 1965)
Pl. III, fig. 1
D e s c r i p t i o n (Paliy, 1976). Typical specimens of this species represent small (2-5 mm in di-
ameter) more or less convex round molds (positive hyporelief) with corresponding negative epirelief.
The surface of the imprints is generally smooth glazed and seems like covered with a thin film. This is
caused by a consolidated structure of the rock. A transverse section reveals a real shape of the imprints
surface, which is umbrella-like convex down shape with the edges of an umbrella turned up to the center
of an imprint.

Genus Charniodiscus Ford, 1958

Charniodiscus planus Sokolov, 1972
D e s c r i p t i o n (Sokolov, 1972). Discoidal imprint with axial dome. About 72 mm in diameter.
R e m a r k s (V.M. Paliy, A.Yu. Ivantsov, herein). An assumption that present form represents
structure of inorganic origin (a result of selecting weathering over lithological heterogeneities on the
surface of rock) is making. Location of the holotype: Podolia, vicinity of Mogilev-Podolsky, Borshchov
Yar; Vendian, Mogilev-Podolsky Group, Yaryshev Formation, Bronnitsa Beds. Holotype is pictured by
B.S. Sokolov (1972, Pl. I, fig. 4), place of its storage is unknown.

Genus Conomedusites Glaessner et Wade, 1966

D i a g n o s i s (Fedonkin, 1985b). The theca, forming a low cone with concentric wrinkles, is di-
vided by four deep radial grooves. Thus, the peripheral boundary is accordingly tetralobate. Each lobe
can be further divided by small grooves. Along the peripheral boundary, sometimes there is a crown of
rather thick tentacles.
Conomedusites cf. lobatus Glaessner et Wade, 1966
Pl. IV, fig. 1
D e s c r i p t i o n (Fedonkin, 1985b). Large specimen, preserved in a positive hyporelief, is charac-
terized by a relatively smooth, central, conical portion of a circular-square outline. Along the margin of
theca, a distinct ridge is observed. Theca radius in plan outline is approximately one-half that of a body.
Four grooves, cutting the board outer ring, do not extend onto the theca. Margins of the body point to
its tetralobate structure, minor undulations of the margin are also observed. Tentacles are not preserved.
Absence of concentric wrinkles and deep radial grooves in the thecal area suggest that not the outer
(lower) thecal surface, but rather its inner surface or the body surface of the organism is impressed in
central part.

Genus Cyclomedusa Sprigg, 1947

Cyclomedusa cf. davidi Sprigg, 1947
Pl. II, fig. 8
D e s c r i p t i o n . Not provided in the publications on fossils of this region.

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 Cyclomedusa minuta Fedonkin in Paliy et al., 1979
Pl. IV, fig. 6
D e s c r i p t i o n (Gureev, 1987). Small molds of high-conical bodies possessing concentric zonal-
ity. The internal zone is relatively wide, flatten. External zones are narrow and cylinder-shaped. Central
tubercle is relief or submerged. Thin transverse furrows can be observed on the zone following the
internal in one specimen.
C o m p a r i s o n s (Paliy et al., 1979). It differs from Cyclomedusa davidi in more pronounced
zonality; from C. wadea in the presence of relatively wide internal zone; from C. coniformis in smaller
number of zones.
Cyclomedusa plana Glaessner et Wade, 1966
Pl. I, fig. 8
D e s c r i p t i o n (Zaika-Novatsky et al., 1968). Round slightly convex imprint of subumbrella (?),
which surface is divided into three concentric zones external, middle and internal. Internal zone con-
sists of three concentric folds with approximately equal width, which are enclosed in each other. Round
tubercle occurs in the center. Middle zone is the most smooth. It is separated from internal zone by
distinct groove, which is traced for 2/3 of internal zones circuit. Middle zone is separated from external
zone by chains of irregularities. External zone is the most uneven with rare radial grooves. External
edge is rare and slightly ragged. It can be assumed that internal zone represents flatten manubrium; the
middle zone represents the surface of gastral cavity; the external zone free edge of an umbrella. And,
consequently, the imprint corresponds to the surface of subumbrella.
Cyclomedusa serebrina Palij, 1969
Pl. I, fig. 2
D e s c r i p t i o n (Paliy et al., 1979). Round and oval imprints, 40-45 mm in diameter, with depres-
sion (negative epirelief), 3-5 mm in diameter, in the center of a smooth area. Peripheral part of imprint
represents edging formed by thin concentric folds, which are often echelon fold-like overlie each other.
C o m p a r i s o n s (Paliy, 1969). It differs from genotype C. davidi in absence of radial ribbing on
the peripheral part and circular grooves.

Genus Dickinsonia Sprigg, 1947

Dickinsonia costata Sprigg, 1947
Pl. VI, fig. 1
D e s c r i p t i o n (Fedonkin, 1985b). Flat, ovoidal, bilaterally symmetric body, with a distinct seg-
mentation, its length approaching the width. On some impressions, a longitudinal mid-groove is seen.
The body is divided into narrow and long segments, slightly widening towards the outer ends. The
closer to the anterior or posterior ends of the body the segments lie, the greater their curvature towards
the corresponding end and the smaller the angle between them and the mid-line <>.
Dickinsonia cf. tenuis Glaessner et Wade, 1966
Pl. VI, fig. 2
D e s c r i p t i o n (Fedonkin, 1985b). Fragment of a relatively small, flat body whereby the length
is two-fold greater then the width. A distinct mid-ridge is slightly lowered relative to the lateral seg-
mented portions of the body, gradually thinning towards the posterior end. Fine segments (25 per 1 cm
of body length) diverge from deep grooves on each side on the mid-ridge. The description of the speci-
mens is based on an impression on the lower surface of a sandstone bed.

Genus Ediacaria Sprigg, 1947

D i a g n o s i s (Fedonkin, 1985b). Exumbrella surface demonstrates central disc and outer ring. A
sharp annual groove, separating the disc from outer ring, can correspond to the margin of gastral cav-
ity. Radial grooves are mainly confined to the outer ring. On the subumbrella surface there is a round,
central mouth without appendages. One specimen apparently bears numerous long and fine peripheral
tentacles.

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 Ediacaria flindersi Sprigg, 1947
D e s c r i p t i o n (Fedonkin, 1985b). <>. Usually this is flat smooth casts, but rather relief forms
are also frequent. Besides central disk and outer ring, the latter is bearing a prominent small tubercle,
which can be surrounded by several concentric wrinkles as a result of postmortal deformations of the
body. For the same reason concentric wrinkles can be also arranged around central disc, but the disc is
more often separated from the outer ring by distinct low step or groove. Discs radius makes from two
thirds to three fourths of the whole radius. Radius of the central tubercle makes one fourth or one fifth of
the central discs radius. Frequent, very thin and straight grooves, radiating to periphery, are sometimes
visible on the outer ring. They are significantly better preserved on the oral surface of the body, where
they can be observed across almost the whole outer ring. Diameter of the mouth opening is similar to
diameter of the central tubercle or a little less. Thin short tentacles, forming a marginal crown, are more
frequently observed precisely on the oral side. <>.
R e m a r k s (A.Yu. Ivantsov, herein). Photographs of several specimens of the species were pub-
lished by M.A. Fedonkin (1983, Pl. XXIX, fig. 1; 1985b, Pl. I, fig. 1, Pl. II, fig. 4) with the following ref-
erence: Middle Dniester area, right bank of Dniester river, Dnestrovskaya Hydroelectric Power Station;
Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds. These specimens with PIN no.
3994/668 and 3994/287 were not found in PIN collections.
Ediacaria sp.
Pl. II, fig. 7
D e s c r i p t i o n . Not provided.

Genus Elasenia Fedonkin, 1983

Diagnosis (Fedonkin, 1983). Small discoidal organism preserved in a positive hyporelief in a form
of hemispherical mold, which internal zone is occupied by relatively large central disk. A ridge of small
round tubercles adjoining the central disk occurs on the external circular zone.
C o m p a r i s o n s (Fedonkin, 1983). Genus rather resembles Medusinites Glaessner et Wade,
however, an external circular zone of the latter does not possess any tubercles and has only rare radial
furrows.
Elasenia aseevae Fedonkin, 1983
Pl. II, fig. 6
D e s c r i p t i o n (Fedonkin, 1983). Central disk probably, corresponding to gastral cavity, is rela-
tively flat. Its diameter amounts to about one-third of that of the holotype body. Seven rounded tubercles
of the external circular zone, adjoining the central disk, probably correspond to gonads. However, their
distribution around the circumference is not quite regular: some of them are pressed to each other, the
others are spaced at small intervals. On some tubercles, an apical depression is seen. Margins of the
body, judging by their deformation, were fine and delicate. An oral portion is unknown.
Elasenia zhuravlevae Gureev, 1988
Pl. III, fig. 3
D e s c r i p t i o n (Gureev, 1988). The mold in a positive hyporelief has an appearance of a disc
with a superimposed hemisphere. The elongated tubercles with flatten surface occur along the perim-
eter of the hemisphere.
C o m p a r i s o n s (Gureev, 1988). In contrast to the described one (species editors note) the
type species (E. aseevae Fedonkin) has round tubercles which are located on the external flatten zone.

Genus Glaessneria Gureev, 1987

D i a g n o s i s (Gureev, 1987). Large organisms. Rather narrow concentric zones occupy up to 1/3
of body diameter in the central part of the sole, forming a graded-conical appearance. An external zone
is flat, smooth or radially rugate.
C o m p a r i s o n s (Gureev, 1987). It differs from Planomedusites Sokolov in concentric zonal-
ity of the central body part; from Kullingia Glaessner in the absence of a distinct concentric pattern,
which can indicate more dense exumbrella or mesogloea.

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 Glaessneria imperfecta Gureev, 1987
Pl. II, fig. 9
D e s c r i p t i o n (Gureev, 1987). Flat smooth body bears two poorly pronounced concentric zones
and indistinct central tubercle in the central part.
C o m p a r i s o n s (Gureev, 1987). The described species is similar to the type one (Glaessneria
plana (Glaessner et Wade, 1966) editors note), but differs in smaller size of the concentrically znal
central part, smaller number of the concentric zones and indistinct central tubercle.

Genus Gritcenia Menasova, 2003

D i a g n o s i s (Menasova, 2003 b). Small round-shaped mold with central depression and outer
ring.
C o m p a r i s o n s (Menasova, 2003 b). It differs from genus Nimbia in size; from genus Broni-
cella in the presence of central depression.
Remarks (A.Sh. Menasova, herein). Represents the junior synonym of Nimbia Fedonkin.
Gritcenia nana Menasova, 2003
Pl. I, fig. 5
D e s c r i p t i o n (Menasova, 2003b). Small round-shaped mold barely elevated over the substrate.
The central zone is round-shaped with brownish outer ring, probably, due to the iron staining. The outer
ring has slightly higher relief in relation to the central part.
R e m a r k s (A.Yu. Ivantsov, herein). In paper by A.Sh. Menasova number . 3/3 is provided, the
new one KTSNY, specimen no. 1768.

Genus Gureevella Menasova, 2003

D i a g n o s i s (Menasova, 2003a). Almost flat molds with uneven oval-shaped surface; a central
tubercle is round-shaped and displaced in relation to the center of an organism.
C o m p a r i s o n s (A.Sh. Menasova, herein). It differs from the genus Ediacaria Sprigg in flat
central zone.
Gureevella elliptica Menasova, 2003
Pl. I, fig. 3
D e s c r i p t i o n (Menasova, 2003a). Slightly convex or flat elipsoidal body with displaced central
tubercle, possessing flat apex, is separated from outer zone by ring-shaped groove. The outer zone is
unevenly tuberculous, probably, due to the deformations of the soft body during burial. The radial ele-
ments are absent.
R e m a r k s (A.Yu. Ivantsov, herein). In paper by A.Sh. Menasova number . 2/14 is provided,
the new one KTSNY, specimen no. 17150.

Genus Hiemalora Fedonkin, 1984 (=Pinegia Fedonkin, 1980)

D i a g n o s i s (Fedonkin, 1980b). Bowl-shaped body with a smooth or concentrically rugate ab-
oral part and numerous tentacles, radiating from the outer margin of body. Preserved in a negative
epirelief, less frequently, in a positive hyporelief.
Hiemalora cf. stellaris Fedonkin, 1980
Pl. VI, fig. 4
D e s c r i p t i o n . Not provided in the publications on fossils of this region.

Genus Jampolium Gureev, 1988

D i a g n o s i s (A.Yu. Ivantsov, herein). Rounded positive imprints on the bottom of the layer,
consisting of low disc with large tubercle at the center. Disc is crossing over six deep grooves, radiating
from central tubercle at an angle of 60.
R e m a r k s (A.Yu. Ivantsov, herein). In his paper of 1988 Yu.A. Gureev reported on all data on
location of holotype and provided its photograph of good quality, but without reporting on neither genus

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 diagnosis nor species description. Holotype remains in place; and no reasonable revisions of material
were made until the present publication. That is why, while publishing a genus diagnosis for the first
time, I, nevertheless, consider it possible to keep a name and Yu.A. Gureevs authorship, provided earlier.
C o m p a r i s o n s (A.Yu. Ivantsov, herein). It differs from genus Cyclomedusa Sprigg, 1947 in the
presence of well pronounced radial grooves, located symmetrically.
Jampolium wyrzhykowskii Gureev, 1988
Pl. IV, fig. 2
D e s c r i p t i o n . Not provided.

Genus Irridinitus Fedonkin, 1983

D i a g n o s i s (Fedonkin, 1983). Small, rounded casts of the basal part of the polyp, preserved in
a positive hyporelief. Relief is low. Central part is commonly caved in. Numerous thin radial ridges or
wrinkles radiate from the center.
C o m p a r i s o n s (Fedonkin, 1983). Shape of the basal part of the polyp described resembles that
of Nemiana Palij, however, the essential difference is the presence of numerous radial structures detach-
ing from the center to the periphery in representatives of the new genus.
Irridinitus multiradiatus Fedonkin, 1983
Pl. II, fig. 5
D e s c r i p t i o n (Fedonkin, 1983). Radial elements on casts of I. multiradiatus appear differently,
depending on the degree of preservation. Most common are fine wrinkles, irregularly detaching from
the center to the periphery. There are deeper wrinkles, cutting the margin of a body, and very fine ones.
Specimens with radial wrinkles, <> are possibly casts, which display characteristic features of the
polyp basal surface. Therefore, it is difficult to say to what extent the above wrinkles reflect the number
and arrangement order of inner radial structures. Casts, upon which structure of the inner part of these
polyps is impressed <>, appear different. These casts have a relatively low relief and are preserved
on the top of a bed. From a small central depression, which corresponds to an oral aperture, numerous
unbranched radial ridges detach towards the periphery (canals of a gastrovascular system?) and reach a
narrow marginal ridge. Most likely, the marginal ridge corresponds to an annular canal of a gastrovas-
cular system. The number of radial canals can reach 40 and more.

Genus Kaisalia Fedonkin, 1984

D i a g n o s i s (Gureev, 1987). Large organisms with concentrically three-zoned body structure.
A central tubercle is absent.
Kaisalia levis Gureev, 1987
Pl. II, fig. 2
D e s c r i p t i o n (Gureev, 1987). Round imprint in positive hyporelief. The internal and middle
zones are convex and distinct; the external one is flatten, slightly distinguishable from the surface of
mark-bearing layer. On the external zone weak irregularly concentric folds can be observed, which are
traced only fragmentary.
C o m p a r i s o n s (Gureev, 1987). It differs from type species in an absence of clearly pronounced
concentric pattern on the surface of middle and external zones.

Genus Kamenecia Gureev, 1988

D i a g n o s i s (, 1988). Star-shaped organism. The rays are short and wide; the body is flat-
ten and tuberculous. Number of rays is 6 or 7.
Kamenecia stella Gureev, 1988
Pl. V, fig. 7
D e s c r i p t i o n . Not provided.

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 Genus Kelleria Gureev, 1988
Kelleria kelleri Gureev, 1988
D e s c r i p t i o n . Not provided.
R e m a r k s (A.Yu. Ivantsov, herein). As in the case of Jampolium wyrzhykowskii Gureev, 1988
author did not provide any diagnosis of the genus and description of the species. Location of the single
specimen of the species: Vinnitsa region, right bank of Lyadova river, Vinozh village, Popeluhovsky
Yar; Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds. This specimen (holotype)
is pictured by Yu.A. Gureev (1988, Pl. XI, fig. 3), its place of storage is unknown. Loss of the holotype
complicates an understanding of the taxon.

Genus Kimberella Wade, 1972

aff. Kimberella sp.
Pl. V, fig. 8
D e s c r i p t i o n . Not provided.
R e m a r k s (A.Yu. Ivantsov, herein). In the paper, where this identification is provided (Gureev
et al., 1987), the description of the form is absent. Fossil was re-described afterwards as Ternavellus
vialovi Gureev, 1988.

Genus Kullingia Glaessner in Fyn et Glaessner, 1979

D i a g n o s i s (Gureev, 1987). A body is flatten, thin, covered with concentric pattern. The central
part is conical, concentrically zonal or simple. The central tubercle is clearly pronounced.
C o m p a r i s o n s (Gureev, 1987). It differs from Glaessneria Gureev and Planomedusites Soko-
lov in distinct thin concentric pattern throughout the surface of flat exumbrella.
Kullingia concentrica Glaessner, 1979
Pl. IV, fig. 5
D e s c r i p t i o n (Gureev, 1987). Flat mold with conical or hemispherical central tubercle and
numerous concentric thin ridges, evenly covering all the body (concentric pattern). The ridges are sepa-
rated mainly by very thin, sometimes relatively wide grooves. The ridges can be rather thicker in the
center of the mold and make a sufficiently relief projection. Sometimes a thickening of the ridges occurs
on some distance from the center.

Genus Lomosovis Fedonkin, 1983

D i a g n o s i s (Fedonkin, 1983). Large, dendritic, colonial organisms. The basal part is a high,
relatively broad obconical stem with a small discoid basal attachment. Relatively fine, tubular processes
detach from the upper part of the stem and end in a crown of fine, long setae. Such processes can also
diverge from the lateral sides of the basal stem; they often display a dichotomous branching. The sur-
face of the organism is flat.
Lomosovis malus Fedonkin, 1983
Pl. VI, fig. 5
D e s c r i p t i o n (Fedonkin, 1983). The upper part of the basal stem bears two semicircular pro-
cesses, either continuing as long, tubular growths or as shorter brush-like terminations. Below, the stem
has two or three processes of different length. These processes are commonly rather thick proximally;
they then became narrow, and in places of branching, became slightly thicker. The surface of the pro-
cesses and basal stem are usually smooth, though the processes are covered by fine and long mainly lon-
gitudinal wrinkles. The basal stem in one case bears distinct transverse wrinkles: fine almost straight
double grooves, cutting the stem from one edge to another. Judging by the thin occasional wrinkles,
curvatures, traces of twisting and contortion, the processes, as well as the basal stem seem to be rather
soft and flexible. The ends of the processes and offshoots are commonly brush-like

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 Genus Marnium Gureev, 1988
D i a g n o s i s (Gureev, 1988). The body is bizonal; the central tubercle occurs in the center of the
inner zone. Numerous appendages (tentacles) occur along the edge of the body.
R e m a r k s (Gureev, 1988). It is similar to Tirasiana Palij by body shape, however, Tirasiana fos-
sils are known in abundance and of good preservation. Thats why the presence of tentacles in Marnium
cannot be considered as a result of Tirasiana finding of especially good preservation.
Marnium cristatum Gureev, 1988
Pl. VI, fig. 3
D e s c r i p t i o n . Not provided.

Genus Medusinites Glaessner et Wade, 1966

D i a g n o s i s (Gureev, 1987). <> It is characterized by submerged central tubercle, surrounded
by circular hollow.
Medusinites asteroides (Sprigg, 1949)
Pl. III, fig. 4
D e s c r i p t i o n (Gureev, 1987). Small imprints on the lower bedding plane (positive hyporelief)
are in the form of round flatten bodies with central tubercle of the same height as the external zone or
somewhat higher. The tubercle is hemispherical, bounded by the circular hollow. In case of ferruginized
surface of the specimen the top of central tubercle frequently has more fresh appearance (cleavage),
which indicates the lifetime penetration of the central tubercle into the sediment; i.e. allows interpreting
it as an attaching organ. Sometimes several adjoining individuals with united lateral boundary occur,
which indicates the reproduction by longitudinal dividing.
C o m p a r i s o n s (Gureev, 1987). It differs from Medusinites paliji in more stable sizes of central
tubercle.
Medusinites paliji Gureev, 1987
Pl. III, fig. 7 a, b
D e s c r i p t i o n (Gureev, 1987). Medusinites with unstable correlation between body sizes and
size of central tubercle has irregularly round shape. Thick concentric pattern sometimes occurs on the
surface of the outer zone of large specimens. Small Medusinites paliji specimens resemble Beltanel-
loididae Gureev, 1987.
R e m a r k s (A.Yu. Ivantsov, herein). In later publication (Gureev, 1988) the other specimen with
the number NMNH, no. 2127/23 was named and shown as holotype that cannot be accepted. Yu.A.
Gureev considered each imprint as a single individual of Vendian organisms (Gureev, 1987). That is
why designation of holotype, given by him to the assemblage of several imprints, presented on the
specimen No. 2127/19, should be considered as a direct mistake. According to the authors description
(Gureev, 1987), large imprint, shown on Pl. III, fig. 7 b at the center, is chosen as lectotype of the species.
Medusinites patellaris Sokolov, 1972
D e s c r i p t i o n (Sokolov, 1972). Medusoidal saucer-like organism with wide central disc (20
mm), from which radial grooves are slightly shown, diameter is about 65 mm.
R e m a r k s (V.M. Paliy, A.Yu. Ivantsov, herein). An assumption that present form represents
structure of inorganic origin (a result of selecting weathering over lithological heterogeneities on the
surface of rock) was made. Location of the holotype: Podolia, vicinity of Mogilev-Podolsky, Borshchov
Yar; Vendian, Mogilev-Podolsky Group, Yaryshev Formation, Bronnitsa Beds. Specimen is pictured by
B.S. Sokolov (1972, Pl. II, fig. 2), place of its storage is unknown.
Medusinites sokolovi Gureev, 1985
Pl. IV, fig. 7
D e s c r i p t i o n (Gureev, 1985). Medusoids of medium sizes. Smooth concave area of round
shape in a plan view (subumbrella?) with tubercle at the center (oral stem?). The apex of tubercle is
convex to slightly concave (mouth?). A cast of subumbrella is frequently preserved as an unsculptured
wide circular ridge. Ratio of medusoid umbrellas diameter to diameter of tubercle is no less than 5:1.

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 C o m p a r i s o n s (Gureev, 1985). It differs from other species of present genus M. patellaris
Sokolov, 1972, M. asteroides (Sprigg, 1949), M. patellaris (Fedonkin, 1980) (in paper by Fedonkin tax-
on with such specific name is absent editors note) in bigger sizes of umbrella in relation to tubercle,
furthermore it differs from the last two in the absence of any signs of radial morphology.
R e m a r k s (A.Yu. Ivantsov, herein). In papers by Yu.A. Gureev the specimen is shown to the
fuller extent. It appears to have been damaged during storage.

Genus Nemiana Palij, 1976

D i a g n o s i s (Paliy, 1976). Imprints of sedentary animals in a form of positive hyporelief and
negative epirelief. Casts represent convex discs with smooth surface, arranged in groups or (rarely)
isolated.
C o m p a r i s o n s (Paliy, 1976). Nemiana imprints differ from genus Bergaueria Prantl (fossil
traces) in the absence of the cylindrical part, crater-like central depression, surrounded by the crown
of tiny tubercles; from genus Tirasiana in more simple structure (an absence of distinct concentric
sculptures).
Nemiana simplex Palij, 1976
Pl. III, fig. 2
D e s c r i p t i o n (Paliy, 1976). Casts (positive hyporelief) of medium sizes, generally round, con-
vex, with smooth surface. Occurring irregularities have the appearance of irregular dents, grooves, con-
centric wrinkles and folds. Delicate pattern as a range of concentric circles occurs very rarely. Casts are
frequently arranged in groups in number from several specimens to several dozens or even hundreds;
as a result of close contact they can obtain a polygonal shape. On the specimens of better preservation
it is evident that surface edges of the cast are slightly turned up towards the center; in consequence of
what, the cast became short vase-shaped with semispherical bottom and incurved edges. The casts are
separated from the underlying bed by thin layer of clayey rock. The casts with offshoots occur rarely,
which resemble daughter forms of gemmating organisms. Negative epirelief represents depressions,
corresponding to casts, but less distinct.

Genus Nimbia Fedonkin, 1980

D i a g n o s i s (Fedonkin, 1985b). Flat, discoidal organisms preserved as annular or ovate struc-
tures at the base of siltstone interlayers (positive hyporelief). The peripheral part of the disc is preserved
in the form of a relief, marginal ridge. An extensive area inside the ring is generally smooth, rarely with
a small tubercle in the center.
Nimbia dniesteri Fedonkin, 1983
Pl. II, fig. 10
D e s c r i p t i o n (Fedonkin, 1983). Small discoid organisms with a flat and smooth central part
and a clearly protruding marginal ridge with a trapeziform cross-section. The oral aperture is small; it
is preserved as a shallow, rounded depression in the centre of the disc. Rare, fine and short tentacles,
not always preserved, diverge from the marginal ridge. Longitudinal division of the organisms is fairly
common.
C o m p a r i s o n s (Fedonkin, 1983). Unlike Nimbia occlusa Fedonkin, which has a smooth or
segmented marginal ridge with a rounded section, this species exhibits a low-relief ridge of a relatively
greater width with a trapeziform cross-section.
Nimbia occlusa Fedonkin, 1980
I m a g e o f s p e c i m e n s . Fedonkin, 1985 b, Pl. III, fig. 1, 6, 7.
D e s c r i p t i o n (Fedonkin, 1985b). The ring is circular to ovate. The radius of the central smooth
zone is three to five times bigger than the width of the outer ridge. In a low relief, the ridge is gently
sloping and smooth; in a higher relief, in some cases the ridge has regular, but weak constrictions, less
common are transverse furrows. In the center of a smooth zone, a small, gently sloping tubercle with an
acute apex is sometimes preserved.

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 R e m a r k s (A.Yu. Ivantsov, herein). In paper by M.A. Fedonkin (1985b, Pl. III, fig. 1, 6, 7) 3
specimens of the species (PIN, specimen no. 3994/533, 479-, 644) were pictured with the follow-
ing reference: Middle Dniester area, right bank of Dniester river, Dnestrovskaya Hydroelectric Power
Station; Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds. Specimens were not
found in PIN collections.
Nimbia paula Gureev, 1985
Pl. IV, fig. 4
D e s c r i p t i o n (Gureev, 1985). Small medusoid with an imprint of subumbrella, preserved as a
circular ridge, surrounding a smooth central zone. The surface of the ridge is smooth; constrictions and
furrows are absent. The central zone is slightly uplifted.
C o m p a r i s o n s (Gureev, 1985). It is similar to N. occlusa Fedonkin in morphology, differing
in considerably smaller sizes.
Genus Onuphionella Kirjanov, 1968
D i a g n o s i s (Kirjanov, 1968). Large straight light tubes, consisting of mica scales, tightly ad-
joining each other; plane oriented transversely to the tube axis.
Onuphionella agglutinata Kirjanov, 1968
Pl. IX, fig. 4 a, b
D e s c r i p t i o n (Kirjanov, 1968). Large straight light tubes, consisting of scales, tightly adjoin-
ing each other (agglutinate?). The tubes of the majority of specimens are deformed into the thick straight
bands. The tubes of two specimens are not deformed and are filled with the rock. One of the ends of
tubes fragment on one specimen is covered with the same mica material. The tubes are formed of vari-
ous micas, mainly muscovite. The size of mica scales is 0.1-0.3 mm. All mica scales in undeformed
specimens are plane oriented transversely to the tube axis. The length of the tubes fragments in collec-
tion is up to 1.5 cm; the length of imprints with preserved tubes fragments is up to 6.5 cm. The tubes
diameter is about 4 mm.
Genus Palaeospinther Menasova, 2003
D i a g n o s i s (Menasova, 2003a). Circular, fine segmented impressions of convex shape.
C o m p a r i s o n s (Menasova, 2003a). It differs from genus Dickinsonia Sprigg in three-dimen-
sional form.
Palaeospinther nucis Menasova, 2003
Pl. V, fig. 2
D e s c r i p t i o n (Menasova, 2003). Convex bilaterally segmented body with a length close to
a width; at first sight it resembles walnuts nutshell. The medial furrow is shown on the imprint, from
which the segments originate in a form of fine hatching (negative epirelief).
R e m a r k s (A.Yu. Ivantsov, herein). In paper by A.Sh. Menasova number 3/2 is provided for ho-
lotype, the new one KTSNY, specimen no. 17151.
Palaeospinther conoideus Menasova, 2003
Pl. V, fig. 1
D e s c r i p t i o n (Menasova, 2003). Flatten conical impressions with a flat apex and fine radial
hatching, greatly resembling burrows of modern Polychaeta. Something, resembling five-radial symmetry, can
be observed on the images of some imprints. The apex is flat, round-shaped.
C o m p a r i s o n s (A.Sh. Menasova, herein). It differs from species P. nucis in conical (round
instead of ovate in a plan view) shape and absence of longitudinal groove.
R e m a r k s (A.Yu. Ivantsov, herein). In paper by A.Sh. Menasova number -83503- is provided
for holotype, the new one KTSNY, specimen no. 1781.
Genus Paliella Fedonkin, 1980
D i a g n o s i s (Fedonkin, 1985b). Flat and circular body composed of the central tubercle with a
flatten apex and a broad, smooth, outer zone, separated by a narrow annular groove. Tentacles are absent.
Radial elements (commonly irregular, isolated and paired grooves) observed only in the largest specimens.
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 C o m p a r i s o n s (Fedonkin, 1985b). Paliella is similar in its structure to Cyclomedusa, differ-
ing in the absence of concentric grooves over the entire umbrella or in the central part, as in C. plana
Glaessner et Wade. Small forms of Paliella resemble Medusinites asteroides Sprigg, differing in the
absence of distinct radial grooves, intersecting the outer zone of the latter.
Paliella pateliformis Fedonkin, 1980
Pl. II, fig. 4
D e s c r i p t i o n (Fedonkin, 1985b). The largest specimens are noted for a gently undulating, mar-
ginal part of the umbrella or its lateral deformations, as well as deep and short radial grooves, diverging
from the margin of the umbrella, or fine, isolated and paired grooves, extending from the margin of the
umbrella to the central annular groove. The inner portion, surrounded by an annular groove, is devoid
of radial furrows, being relatively flat in large specimens and protruding as a flat-tipped tubercle in
smaller ones. Small juvenile specimens provide more inflated casts than larger ones. Specimens occur
in groups, or in pairs, where individuals are closely pressed to each other and rimmed by a common
narrow zone. Such an occurrence indicates that the reproduction of Paliella proceeded as a simple,
longitudinal division. However, there are clear indications of multiple division (two types: linear and
clustered) and a stolonal type of asexual reproduction, when a long stem-like stolon extends from the
central part, sometimes with a daughter individual on the end.
Genus Planomedusites Sokolov, 1972
D i a g n o s i s (Sokolov, 1972). Discoidal medusoid organism with smooth surface, bearing pe-
ripheral ridge; diameter is about 125 mm.
Planomedusites grandis Sokolov, 1972
D e s c r i p t i o n . Not provided in the publications on fossils of this region.
R e m a r k s (V.M. Paliy, A.Yu. Ivantsov, herein). An assumption that present form represents
structure of inorganic origin (a result of selecting weathering over lithological heterogeneities on the
surface of rock) was made. Location of the holotype: Podolia, vicinity of Mogilev-Podolsky, Borshchov
Yar; Vendian, Mogilev-Podolsky Group, Yaryshev Formation, Bronnitsa Beds. Holotype is pictured by
B.S. Sokolov (1972, Pl. II, fig. 1), place of its storage is unknown.
Genus Platysolenites Pander, 1850
Platysolenites antiquissimus Eichwald, 1860
Pl. IX, fig. 2
D e s c r i p t i o n (Kirjanov, 1968). White segmented thick-walled tubes. The segmentation is
sharp, distinct and even in some specimens, and indistinct, but distinguishable well enough in oth-
ers. The longest tubes fragments in collection are generally getting narrow in one end. In individual
specimens the segmentation is uneven; totally disappearing at one end of the fragment, and the tube
became smooth. <>.
Genus Podoliina Gureev, 1988
D i a g n o s i s (Gureev, 1988). Imprints on the bedding planes in a form of low cross-wrinkled
ridges, composed of the same material as the mark-bearing layer. The rugosity is thick, inaccurate
and irregular. It forms the ribs, which sometimes go beyond the imprints edges. The ridges are long,
strongly curved; superposition and twisting are not observed.
Podoliina crassa Gureev, 1988
Pl. VIII, fig. 1
D e s c r i p t i o n . Not provided.
Genus Podolimirus Fedonkin, 1983
D i a g n o s i s (Fedonkin, 1983). Large, bilaterally symmetrical organism, composed of paired,
arched segments, wide in the proximal part and narrowing distally. Segments, diverging at 80-85 from
the symmetry plane, curve abruptly, becoming subparallel to the above plane in their distal part. Size
of segments decreases successively in the same direction, i.e. towards the growth zone. The ends of the
last segments in this zone become aligned, forming a sort of comb.
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 C o m p a r i s o n s (Fedonkin, 1983). The genus described is similar to Pteridinium Grich in its
segmentation and, probably, both genera can be placed into the family Pteridiniidae Richter. The new
genus differs in much greater dimensions and proportions of the segments, and a structurally different
growth zone.
Podolimirus mirus Fedonkin, 1983
Pl. V, fig. 5
D e s c r i p t i o n (Fedonkin, 1983). On the holotype (impression), nine segments preserved in the
right part; and six in the left one. Smooth, flattened segments with narrow marginal ridges. Their ends
are, probably, irregularly bifurcated. Arrangement of segments is not strictly opposite, with consider-
able displacement. Judging by an irregular decrease in the width of certain segments from the proximal
to distal ends, segments were relatively mobile and could be partly overlapped, at least in their distal
parts. However, one should bear in mind possible disturbances of continuity of the body after the dearth
of an organism and in the course of burial. Distal parts of the last 10-12 segments are subparallel, and
their ends are aligned, forming a pectinate edge. The first segment on the right is much wider than suc-
cessive ones, particularly in its proximal part, where a relatively small trapeziform area with a distinct
relief is located. Unlike the second and subsequent segments, the front part at the beginning of the first
segment is not rounded, but rounded-angular.

Genus Pollukia Gureev, 1987

D i a g n o s i s (Gureev, 1987). Tirasiana with uneven (wrinkled and tuberculous) outer zone.
C o m p a r i s o n s (Gureev, 1987). It differs from Tirasiana Palij in the habitus of the outer zone
and the absence of inner zones relief.
Pollukia shulgae Gureev, 1987
Pl. II, fig. 1
D e s c r i p t i o n (Gureev, 1987). Small cast on the bottom bedding plane of siltstone; the surface
of the cast is covered with pyrite. The outer zone is slightly elevated in relation to the inner one and has
uneven (tuberculous) surface. The margin is slightly uneven. A central tubercle is small and distinct.
C o m p a r i s o n s (Gureev, 1987). It differs from Pollukia serebrina (Cyclomedusa serebrina
Palij, 1969 editors note) in tuberculous outer zone and the way its rising relatively to inner one.

Genus Propaleolina Menasova, 2003

D i a g n o s i s (Menasova, 2003b). Elongated, faintly curved tubes represented inner casts. The
surface is brownish. The tubes look like smooth grooves in a negative epirelief.
C o m p a r i s o n s (Menasova, 2003b). It differs from genus Paleolina in the absence of external
skeleton and sculpture traces.
Propaleolina vendiensis Menasova, 2003
Pl. IX, fig. 1 a, b
D e s c r i p t i o n (Menasova, 2003b). The specimens surface is covered with unsystematically
oriented faintly twisting tube-like structures, sometimes immersing into the substrate. It probably rep-
resents inner core, filled with substrates rock. The tubes surface is brownish, possibly represented
all that was preserved from the organisms skeleton. The gently sloping grooves occur in the negative
epirelief. The traces of sculpture are not observed.
R e m a r k s (A.Yu. Ivantsov, herein). In paper by A.Sh. Menasova number . 6/1 is provided for
holotype, the new one KTSNY, specimen no. 1782.

Genus Protodipleurosoma Sprigg, 1949

D i a g n o s i s (Fedonkin, 1985b). Rounded impression of exumbrella, broad outer ring (velum?)
covered by narrow concentric grooves; mid-field large, circular, with an ovoidal, irregularly compressed
central area (stomach?), divided into lobes, and irregular radial grooves (radial canals?), with non-
dichotomous branching; primary grooves deep, reaching the submarginal groove (radial canal?) giving
rise to shorter furrows: branching near the base.

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 Protodipleurosoma rugulosum Fedonkin, 1980
D e s c r i p t i o n (Fedonkin, 1985b). The median groove, commonly not strictly in the middle
of an ovate impression, divides the central disc into two unequal, elongated lobes. Minor furrows and
wrinkles, radiating from the median groove, thin towards the periphery of the central disc, not affecting
the annular zone. The latter is well-preserved in large specimens, strongly reduced (if at all) in smaller
ones. On the holotype, semicircular processes are seen around the body periphery, which may corre-
spond to reproductive organs or budding daughter individuals. As a rule, one lobe of the central disc has
more radial wrinkles than the outer.
C o m p a r i s o n s (Fedonkin, 1985b). The new species differs from Protodipleurosoma wardi Sprigg
in the greater number of wrinkles, relatively larger size and lateral processes in the largest specimens.
R e m a r k s (A.Yu. Ivantsov, herein). Specimen PIN, no. 3994/406, pictured by M.A. Fedonkin
(1983, Pl. XXXI, fig. 5), has the following reference: Middle Dniester area, right bank of Dniester river,
Dnestrovskaya Hydroelectric Power Station; Vendian, Mogilev-Podolsky Group, Mogilev Formation,
Lomozov Beds. Specimen was not found in PIN collection.
Protodipleurosoma wardi Sprigg, 1949
Pl. IV, fig. 9
D e s c r i p t i o n . Not provided in the publications on fossils of this region.
Genus Pseudorhizostomites Sprigg, 1949
Pseudorhizostomites sp.
D e s c r i p t i o n . Not provided in the publications on fossils of this region.
R e m a r k s (A.Yu. Ivantsov, herein). Specimen PIN, no. 3994/594, pictured by M.A. Fedonkin
(1983, Pl. XXXI, fig. 2), has the following reference: Middle Dniester area, right bank of Dniester river,
Dnestrovskaya Hydroelectric Power Station; Vendian, Mogilev-Podolsky Group, Mogilev Formation,
Lomozov Beds. Specimen was not found in PIN collection.

Genus Sabellidites Yanischevskii, 1926

Sabellidites cambriensis Yanischevskii, 1926
Pl. IX, fig. 5 a, b
D e s c r i p t i o n (Kirjanov, 1968). Black dense non-segmented tubes, flatten in a form of nar-
row bands or non-deformed, filled with the rock. Rugosity is faint, but distinct; its frequency changes
towards one end of the fragment are observed in single specimens; sometimes rugosity is not observed,
however, thin hatching is clearly visible.
Genus Sekwia Hofmann, 1981
D i a g n o s i s (Gureev, 1987). Medusinitiids with tubercle, elevated relatively to outer zone.
Sekwia kaptarenkoe Gureev, 1987
Pl. III, fig. 5
D e s c r i p t i o n (Gureev, 1987). Small convex (positive hyporelief) imprints of animals with
subcylindrical body shape, which can be indicated by the observed incline of some casts. The central tu-
bercle is large and hemispherical. The outer zone is flatten. The groups of cast are frequently observed.
C o m p a r i s o n s (Gureev, 1987). It differs from Sekwia excentrica Hofmann in central position
of tubercle.
Genus Serpulites Murchison, 1839
Serpulites (?) petropolitanus Yanischevsky, 1926
D e s c r i p t i o n . Not provided in the publications on fossils of this region.
R e m a r k s (A.Yu. Ivantsov, herein). Specimen NMNH, no. 1731/5, pictured by V.V. Kirjanov
(1968, Pl. V, fig. 24), has the following reference: Volhynia region, B. Obzyr village, borehole no. 5,
depth of 180.1 m; Lontova Horizon, Baltic Group, Stokhod Beds (age analogue of Zbruch Formation of
Dniester area). Specimen is broken.

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 Genus Sokoloviina Kirjanov, 1968
D i a g n o s i s (Kirjanov, 1968). Black non-segmented tubes of small and medium size with sharp
even rugosity, prominent around the tubes stem in a form of pointed or obtuse circular offshoots.
C o m p a r i s o n s (Kirjanov, 1968). Tubes of described genus are rather similar with tubes of ge-
nus Paleolina Sokolov. Unlike the latter, the tubes of genus Sokoloviina are dense, non-transparent; the
colour of the chitin matter is black.
Sokoloviina costata Kirjanov, 1968
D e s c r i p t i o n (Kirjanov, 1968). Black, non-segmented, non-transparent tubes of small and me-
dium size with sharp rugosity, prominent around the tubes stem in a form of thin pointed or obtuse
circular offshoots. The rugosity is thick in small specimens, and thin, twisting and branching in large
ones. The thickness of wrinkles at the base is 0.03-0.1 mm. The inner surface of the tubes is smooth
with thin hatching. The strokes correspond to grooves on the outer side of the tubes. The chitin matter
of the wrinkles remains in rock under the split of tubes stem from the rock surface; and the wrinkles
branching is distinctly shown. The length of the tubes fragments in collection is up to 15 mm, the width
ranges from 0.3 mm to 1 mm.
R e m a r k s (A.Yu. Ivantsov, herein). Specimen NMNH, no. 1731/2, pictured by V.V. Kirjanov
(1968, Pl. III, fig. 4), has the following reference: Volhynia region, Manevich area, Berezhnitsa village,
borehole no. 17, depth of 202.1-202.2 m; Lontova Horizon, Baltic Group, Stokhod Beds (age analogue
of Zbruch Formation of Dniester area). The specimen fell into small fragments.

Genus Studenicia Gureev, 1983

Studenicia galeiforma Gureev, 1983
Pl. VIII, fig. 5
D e s c r i p t i o n (Gureev, 1983 a (the formal description is not provided, this description is de-
rived from general text editors note)). Small, round in a plan view, body imprints of sedentary ani-
mals on the lower bedding plane. The upper part of imprint possesses the central tubercle.
C o m p a r i s o n s (Gureev, 1983 a (the formal description is not provided, this description is
derived from general text editors note)). It differs from Nemiana in the presence of central tubercle,
from Tirasiana in the absence of circular sculpture and more simple structure.
R e m a r k s (A.Yu. Ivantsov, herein). Interpretation of the fossil by the author has been changing
(see description of Monocraterion sp.). In paper by Yu.A. Gureev (1983a) number 1970/64 is provided
for holotype, the new one NMNH, specimen no. 2088/9.

Genus Ternavellus Gureev, 1988

D i a g n o s i s (Gureev, 1988). Convex cast, triangular in a plan view (isosceles triangle). Three
offshoots occur at the angles and center of the base.
Ternavellus vialovi Gureev, 1988
Pl. V, fig. 8
D e s c r i p t i o n (Gureev, 1988). Flat body with almost upright marginal surfaces (positive hypo-
relief). Contact with the mark-bearing layer is sharp. A row of tubercles of uncertain nature (probably,
mud-eater traces?) occur on the surface of the cast. Outgrowths are short and wide. Middle outgrowth
seems to stretch out the base of a triangle. Distinct bulges are observed on the lateral outgrowths; the
bulge on the middle outgrowth is inexpressive.

Genus Tirasiana Palij, 1976

D i a g n o s i s (Paliy, 1976). Impressions of sessile organisms in the form of a positive hyporelief
and negative epirelief. Casts are convex to conic; the surface has a concentric ornamentation: shelves
and ridges. Arranged in groups of several specimens or isolated.
C o m p a r i s o n s (Paliy, 1976). The imprints of Tirasiana differ from genus Nemiana in the
presence of concentric sculpture of the surface. In addition Tirasiana does not form big accumulations.

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 Tirasiana coniformis Palij, 1976
Pl. I, fig. 7
D e s c r i p t i o n (Paliy, 1976). Typical cast of T. coniformis is interpreted as consisting of three
superimposed discs with consistently decreasing sizes. A round tubercle occurs on the flat surface of the
last disc. The most convex specimen has the least prominent sculpture of the surface. Otherwise, more
distinct concentrically stepped relief occurs on the slightly flattened casts, experienced great compres-
sion in a vertical direction. It can be suggested that concentrically stepped cast reflects the features of
body structure of an animal that became evident under deformation. The general body shape was ap-
parently goblet-shaped, close to conical.
C o m p a r i s o n s (Paliy, 1976). It differs from the species T. disciformis in more convex cone-
shaped form and more complex sculpture in a form of three superimposed discs.
Tirasiana disciformis Palij, 1976
Pl. I, fig. 6
D e s c r i p t i o n (Paliy, 1976). Moderately convex casts (3-4 mm) in the form of medium-sized
discs on the lower bedding planes. The main, slightly flattened, disc is superimposed by the second,
smaller and relatively more convex disc with a small, round tubercle in the centre. The margin of the
main disc surface, extending into the rock, is curved abruptly inward. The casts are separated from the
underlying bed by thin interlayer of the bluish-green mudstone. Unlike the rest of the surface, which is
easily separated from the rock, the central tubercle is often cemented in the underlying layer. The casts
more often are arranged in small groups (3-4 specimens) or isolated. The casts of adherent specimens
are very typical.
C o m p a r i s o n s (Paliy, 1976). The casts of T. disciformis differ from the species T. coniformis
Palij in flatter shape and simple structure: appearance of two superimposed discs.

Genus Tribrachidium Glaessner, 1959

D i a g n o s i s (Fedonkin, 1985b). Discoid organism, slightly convex, with a relatively steep pe-
ripheral boundary. One side (oral?) has three elevated arms, which radiate at the same angle, curving
clockwise, and which join the periphery of the disc, where they become thinner. Occasionally, a small
Y-shaped furrow is seen in the centre between the arms. From each arm on its concave side, a small
elevated area (bulla) diverges. At about 0.7 of the length, distal parts of the arms bear short, strong
tentacles on the outer side and on the acute side of each arm. Between arms sometimes there are fine,
long, straight or slightly curved setiform structures, extending from the convex part of each arm across
an acute portion of a neighboring arm. The opposite side (aboral?) of the disc displays only several
concentric grooves.
Tribrachidium heraldicum Glaessner, 1959
Pl. V, fig. 3
D e s c r i p t i o n (Fedonkin, 1985b). Small discoid fossils. In a negative epirelief (hyporelief edi-
tors note) three arms radiate from the central part. They curve smoothly counterclockwise, and then
parallel peripheral termination occurs, tapering towards the ends. Distal portions of the arms, for about
two-thirds of their length, bear numerous short tentacles (about 20 on each arm), forming a fringe along
the outer margin of the body. Each of the three areas, between branches, ramifies, starting from a convex
part of the arm near the centre and almost touching with its end the concave portion of the neighboring
arm at a point separated by about one-fourth of its length from the distal end. Cast of upper, oral surface
(positive epirelief), not very common in a fossil state, displays the following structural features (the natu-
ral body cast is implied, the imprint of the upper side of the body was considered in the previous para-
graph editors note). The upper side is scutiform and slightly convex. In the middle portion, there are
three arched depressions, oriented towards the centre of the disc with their narrow ends. The outer sides
of the arched depressions are relatively broader and deeper due to small, rounded, terminal widenings.
The disc surface is speckled with fine radial grooves and ridges entering the arched depressions. Such a
structural pattern of the oral side enables the interpretation of a Tribrachidium as a more primitive organ-
ism than it was previously assumed <>. Arched depressions with a terminal widening may belong to

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 oral apertures, though, possibly, only terminal widening served for introducing food. Fine radial grooves,
entering the arched depressions, may be interpreted as microrelief, favouring transportation of nutrient
particles from the entire umbrella area towards the oral apertures by means of ciliated epithelium. A
further analysis of Tribrachidium morphology from the same viewpoint shows that arms, bearing short
tentacles in their distal parts, may and even should be regarded as a distribution system, structurally
similar to canals of the gastrovascular system. If this interpretation proves reasonable, Tribrachidium
should be assigned to coelenterates rather than to some other group of Metazoa <>.
R e m a r k s (A.Yu. Ivantsov, herein). In paper by M.A. Fedonkin number PIN, no. 3994/580-A is
provided, the new one PIN, specimen no. 4158/26, the specimen is in the exposition of Yu.A. Orlov
Museum.

Genus Valdainia Fedonkin, 1983

D i a g n o s i s (Fedonkin, 1983). Bipinnate, segmented organism. Short, broad body. Rounded-
zigzag mid-line. Broad, smooth segments are gently curved towards the growth zone; distinctly alter-
nating segments. Proximal ends of the segments are rounded; distal ends taper smoothly and obliquely.
Segments are divided by a pair of grooves, with a long narrow ridge in-between. The end of the body is
bifurcated in the growth zone.
C o m p a r i s o n s (Fedonkin, 1983). The genus described is similar to Pteridinium Grich in
the general structural outline, bit differs in proportions, shape of segments and the presence of a ridge
between neighboring segments diverging to the sides; the finest segments in the growth zone form the
bifurcated end of the body.
Valdainia plumosa Fedonkin, 1983
Pl. V, fig. 4
D e s c r i p t i o n (Fedonkin, 1983). Narrowing of segments from the proximal to distal part begins
at some distance from the mid-line, mainly due to a steeper curvature of the segment margin, which
faces the side, opposite the growth zone. The distal ends of the largest segments are thin and taper
obliquely. According to holotype, the segments are separated by a relatively narrow zone, consisting of
two grooves with a narrow ridge in-between. This intersegmental ridge begins near the mid-line and
extends towards the distal ends of the adjacent (one side) segments. The width of the intersegmental
ridges is variable: the broader the ridge, the narrower the segment. Besides, from the side of the growth
zone of the organism, segments are separated by a much shallower groove from the intersegmental
ridges than from the opposite side. These specific features suggest that intersegmental ridges are parts
of the segments, or rather rigidly conjugated structures, located on the side of the growth zone of the
organism. These parts or structures were apparently impressed in the sediment in those parts of the
body, where the segments are to a certain extent deformed. The largest segments extend at an almost
right angle from the body axis, but approaching the growth zone, the angle between the segments and
the body axis decreases; in the growth zone it equals 30-40

Genus Vaveliksia Fedonkin, 1983

D i a g n o s i s (Fedonkin, 1983). Problematic organisms, preserved as elongated casts and impres-
sions with two distinct parts: a relatively small, regular disc, which may imply a discoid attachment
of the organism, and an ellipsoidal part, adjoining the disc by the long axis end and, apparently, cor-
responding to a soft ampullar sac. On the other end of ellipsoidal part, opposite to the disc, no smooth
closure of the ellipse is observed; short linear elements follow the long axis, forming a pectinate margin.
Vaveliksia svetozarovae Gureev, 1988
Pl. V, fig. 6
D e s c r i p t i o n (Gureev, 1988). Convex, triangular in a plan view impression (positive hypore-
lief). The triangle is closed to isosceles. An outgrowth with terminal widening of irregularly rounded
shape, extending from one of the apexes. The side that is opposite to the terminal widening is uneven;
the existence of processes (tentacles) can be assumed. A small round tubercle of indistinct genesis is
located on the cast.

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 C o m p a r i s o n s (Gureev, 1988). In contrast to type species (V. velikanovi Fedonkin) the de-
scribed form is shorten; its width is close to its length.
Vaveliksia velikanovi Fedonkin, 1983
Pl. III, fig. 6
D e s c r i p t i o n (Fedonkin, 1983). Length of ampullar sac is 3-4 times and width is 1.5 times
larger that of the diameter of the discoid attachment area. The sac is joined to the disc by a short stalk,
not always observed in fossils, if overlain by fixing of the disc during crushing of the organism in the
course of burial. An oral end, opposite the disc, bears a crown of short tentacles, surrounding the oral
aperture. The surface of the casts of the ampullar sac is smooth or tubercular, possibly depending on
whether the inner or outer surface of the sac is impressed in the sediment.

Genus Vendella Gureev, 1987

D i a g n o s i s (Gureev, 1987). Vendellids with wide outer zone and small, hemispherical or coni-
cal central depression.
C o m p a r i s o n s (Gureev, 1987). It differs from Nimbia Fedonkin in small sizes of the central
depression.
Vendella haelenicae Gureev, 1987
Pl. II, fig. 3
D e s c r i p t i o n (Gureev, 1987). Flattened, relief casts of regular rounded shape. A small depres-
sion occurs in the center of the cast (positive hyporelief), in a negative epirelief it is corresponding to
sharp-pointed (under good preservation) tubercle. The central depression is not traced on some speci-
mens, which can be due to poor preservation.
C o m p a r i s o n s (Gureev, 1987). It differs from the other species of this genus in very small
sizes of the central depression relatively to the casts diameter.
Vendella larini Gureev, 1987
Pl. IV, fig. 3
D e s c r i p t i o n (Gureev, 1987). Positive hyporelief is represented by circular ridges, bordering
round and oval (result of deformation?) central depressions. The ridge surface of holotype is compli-
cated by evenly distributed, poorly preserved, narrow transverse furrows.
C o m p a r i s o n s (Gureev, 1987). It differs from Vendella sokolovi and Vendella haelenicae in
smaller general sizes and larger sizes of central depression relatively to diameter of the body. The latter
character makes this species close to representatives of the genus Nimbia Fedonkin.

Discoidal casts with adjoining worm-shaped body

Pl. VIII, fig. 6
D e s c r i p t i o n (Paliy et al., 1979). Casts in a form of small discs, 6-11 mm in diameter and 1-2
mm in thickness, with tubercle at the center, 3-4 mm in diameter. Casts occur on the lower surface of
the greenish-grey siltstone bed <>. Three-dimensional, worm-like, a little flattened, body, 15-28 mm
in length and 3-5 mm in thickness, is adjacent to discs on the side, opposite to tubercle. Constrictions
mark its division on 3-5 segments.
C o m p a r i s o n s (Paliy et al., 1979). In contrast to casts of Nemiana and Tirasiana the described
fossil is completely detachable from the rock.

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 ICHNOFOSSILS AND PROBLEMATIC BIOGLYPHS,
DESCRIBED AS HABITATION, FEEDING AND LOCOMOTION TRACES
OF METAZOA
Genus Anulichnus Gureev, 1983
Anulichnus segmentatus Gureev, 1983
D e s c r i p t i o n (Gureev, 1983a (the formal description is not provided, this description is derived
from general text editors note)). Large, circular, superimposed grazing traces, divided by transverse
constrictions on separate segments.
R e m a r k s (A.Yu. Ivantsov, herein). Holotype, pictured by Yu.A. Gureev (1983a, Pl. I, fig. 7), has
the following reference: Middle Dniester area, right bank of Dniester river, opposite the mouth of the
Ternava river; Rovno Horizon, Baltic Group, Khmelnitsky formation. Place of storage of the holotype
is unknown.
Genus Asterichnus Bandel, 1967
Asterichnus vialovi Gureev, 1984
D e s c r i p t i o n (Gureev, 1984). Star-shaped bioglyphs, circular in the plan view, possessing me-
dian field with straight, non-branching, non-intersecting, mainly vane-like rays, radiating from it. Me-
dian field has irregular, close to oval or circular shape. The number of rays is up to 25.
C o m p a r i s o n s (Gureev, 1984). It differs from the species Asterichnus lawrencensis Bandel in
significantly smaller sizes and more regular distribution of the rays.
R e m a r k s (A.Yu. Ivantsov, herein). Holotype, pictured by Yu.A. Gureev (1984, fig. 1 a), has the
following reference: Podolia, right bank of Zhvan river, opposite Posukhov village; Vendian, Mogilev-
Podolsky Group, Mogilev Formation, upper part of Lomozov Beds. In paper by Yu.A. Gureev the fol-
lowing place of storage and number of the specimen is given: IGS, specimen no. 2088/3. However, the
specimen was not found in IGS collections.
Genus Aviculaichnus Gritsenko, 2009
D i a g n o s i s (Gritsenko, 2009). Thin traces with oblique cross-hatching, resembling bioglyphs
Phycodes pedum, and forming trajectories, resembling flying bird - the tick. Sometimes these ticks
unite into solid double curve and their wings become longer.
C o m p a r i s o n s (Gritsenko, 2009). The presence of oblique cross-hatching put the described
traces close to bioglyphs Phycodes pedum, but the latter has straight short tracks.
Aviculaichnus gureevi Gritsenko, 2009
Pl. VIII, fig. 3
D e s c r i p t i o n (Gritsenko, 2009). Thin traces with oblique cross-hatching, forming double arc-
shaped tracks. The width of the traces is maximal at the middle part of convex arc. The hatches are shorter
at the arc joint and their ends. The number of hatches in the arc is from 4-5 to 20 and more. Generally the
traces, according to their trajectories, resemble diagrammatic representation of flying bird the tick.
The length of the hatches reaches 1.5 mm. The length of the arcs on the tracks is from 4-5 to 10-12 mm.
R e m a r k s (A.Yu. Ivantsov, herein). In paper by V.P. Gritsenko number . 2480/1 is provided for
the holotype, the new one NMNH, specimen no. 2525/93.
Genus Bergaueria Prantl, 1945
D i a g n o s i s (Paliy, 1976). Houses of benthic organisms in a form of cylindrical burrows with
spherically rounded and slightly flatter bottom. Positive hyporelief is in a form of short cylindrical
molds, ending with hemispherical or blunt rounding. Cylindrical part is often non-preserved.
Bergaueria major Palij, 1976
Pl. VII, fig. 5
D e s c r i p t i o n (Paliy, 1976). The fullest preserved specimens represent cylindrical molds with a
rounded lower part often in the shape of a regular hemisphere. The end blunt, rarely conically rounded.

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 Many molds are represented by only lower rounded part, because the cylindrical part of the burrow
was destroyed by underwater erosion, often preceding its filling with the sediment. They are either soli-
tary or arranged in small groups without touching one another. The matter of molds represents small
quartz sandstone with significant compound of glauconite grains and rare inclusions of small nodules.
Clayey composition of underlying bed gives evidence for the fact that the organisms, producing Bergau-
eria major, inhabited slimy bottom. In some cases the burrows, left by them, became habitat of small
ground-feeders, which is evident by their traces. On one of the specimens repeated occupation of the
burrow that was left by its previous inhabitant an animal of smaller sizes, was observed.
C o m p a r i s o n s (Paliy, 1976). Present species is different from previously described Bergau-
eria in the absence of depression at the center of rounded lower part along with the features of radial
structure. B. major also exceeds almost all fossil traces of this genus by its sizes <>
Genus Chomatichnus Donaldson et Simpson, 1962
Chomatichnus loevcensis Gureev, 1984
D e s c r i p t i o n (Gureev, 1984). Endoglyphs in a form of simple, non-branching, vertical or
slightly inclined tubes; transverse annular sculpture sometimes occurs on their surface. The traces are
located in the thin (up to 4 mm) siltstone bed, and do not penetrate into the underlying siltstone bed;
when the traces reach the underlying bed, they bend along the bedding plane. The distribution of the
traces is quite tight (4-5 traces per 1 cm of the bed surface on the average). On the upper bedding plane
they appear as cones or truncated cones with central crater (positive epirelief).
C o m p a r i s o n s (Gureev, 1984). It differs from Chomatichnus wegbergensis Donaldson et
Simpson (Carboniferous, England) in considerably smaller sizes, bend of the vertical tube and presence
of the annular sculpture.
R e m a r k s (A.Yu. Ivantsov, herein). Holotype, pictured by Yu.A. Gureev (1984, fig. 2 ), has the
following reference: Podolia, left bank of the Zharnovka Creek, Loevtsy village; Vendian, Kanilovka
Group, Zharnovka Formation, Kuleshovka Beds. In paper by Yu.A. Gureev the following place of stor-
age and number of the specimen is given: IGS, specimen no. 2088/4. However, the specimen was not
found in IGS collections.
Genus Circulichnus Vialov, 1971
Circulichnus montanus Vialov, 1971
D e s c r i p t i o n (Gureev, 1983c). In a positive hyporelief the traces have an appearance of closed
figure of the circular shape. The ridge is low, narrow with well consistent width. On the specimen of best
preservation a bulge (2 mm) is observed in one area, which origin can be explained by the returning of
the trace-creator to the beginning of the trace with an overlap. The ridge surface is simple; the contact
with the mark-bearing layer is gradual, indistinct.
C o m p a r i s o n s (Gureev, 1983c). The traces from Vendian of Dniester area have smaller diameter
(8-10 mm) than Circulichnus montanus Vialov from Upper Triassic of the Pamirs (35-40 mm). However,
equal width of the ridge and simplicity of its surface allows attribution of Vendian traces to this genus.
R e m a r k s (A.Yu. Ivantsov, herein). Holotype, pictured by Yu.A. Gureev (1983c, figure without
number on the page 9), has the following reference: Podolia, right bank of Ushitsa river, Sokolets vil-
lage; Vendian, Kanilovka Group, Danilovka Formation, Shebutintsy Beds. In paper by Yu.A. Gureev
the following place of storage and number of the specimen is given: IGS, specimen no. 1970/57. How-
ever, the specimen was not found in IGS collections.
Genus Cochlichnus Hitchcock, 1858
Cochlichnus isp.
Pl. VII, fig. 3
D e s c r i p t i o n (Paliy, 1976). Traces in a form of sinuous non-branching furrows on the surface
of dense clay shale. Furrows of constant width, with distinct edges, arcuate in cross-section. Trace lon-
gitudinal axis is straight, more rarely curving. Trace pattern in places is asymmetrical due to the change
in amplitude or sine wave length of certain trail sections.

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 Genus Didymaulichnus Young, 1972
Didymaulichnus tirasensis Palij, 1974
Pl. VII, fig. 6
D e s c r i p t i o n (Paliy et al., 1979). Positive hyporelief shows repeatedly curving intersecting
and overlapping ridges 8 to 14 mm wide. In cross-section they are trough-like, some almost cylindri-
cal. Stretching along the smooth surface of ridges are furrows 0.5 to 2.5 mm wide, up to 1.5 mm deep,
wedge-like in cross-section, with rounded edges.
C o m p a r i s o n s (Paliy, 1974). It differs from species Didymaulichnus miettensis Young in an
absence of lateral slopes on ridges, slightly smaller sizes, along with the fact that trace fragments are
very rarely intersecting on one plane.
Didymaulichnus cf. miettensis Young, 1972
D e s c r i p t i o n (Gureev, 1984). Slightly curved exoglyphs in positive hyporelief have an appear-
ance of a ridge, divided by longitudinal groove into two approximately equal parts. Lateral walls of the
ridges are inclined, gradually passing into mark-bearing layer. Width of the ridge is 8-10 mm.
C o m p a r i s o n s (Gureev, 1984). In contrast to Canadian Didymaulichnus miettensis Young,
Podolian forms are single; however, a taxonomic importance of this feature is indistinct, which required
using open nomenclature.
R e m a r k s (A.Yu. Ivantsov, herein). Specimen, pictured by Yu.A. Gureev (1984, fig. 1 ), has the
following reference: Podolia, left bank of Lyadova river, north margin of Iraklievka village; Vendian,
Mogilev-Podolsky Group, Mogilev Formation, upper part of Lomozov Beds. In paper by Yu.A. Gureev
the following place of storage and number of the specimen is given: IGS, specimen no. 2088/6. How-
ever, the specimen was not found in IGS collections.

Genus Harlaniella Sokolov, 1972

D i a g n o s i s (Paliy, 1976). Braid-like molds (positive hyporelief), covered with oblique hatch-
ing that was created by densely and regularly located furrows. Negative epirelief has an appearance of
grooves with oblique ribbing.
Harlaniella podolica Sokolov, 1972
Pl. VI, fig. 6
D e s c r i p t i o n (Paliy, 1976). Positive hyporelief represents ridges, covered with dense oblique
hatching in a form of thin furrows, inclined at 45-80 to longitudinal axis of the trace. The ridges fre-
quently form smooth curves, sometimes getting slightly narrower or wider to one or another side. Their
ends are indistinct. The ridges outlines are often blurred and merged into the rock surface, which turned
out to be covered with specific furrowed relief, resembling fossil traces of Palaeopascichnus delicatus.
The fact that the orientation of hatches on all studied molds is uniform (directed upwards from left to
right), represents an interesting feature. Negative epirelief has an appearance of ribbed grooves, as a
rule of worse preservation.
Genus Monocraterion Torell, 1870
Monocraterion sp.
Pl. VIII, fig. 5
D e s c r i p t i o n (Gureev, 1984). Vertical endoglyphs with funnel-shaped mouth and filled chan-
nel, passing into the rock. The mouth and beginning of the channel is well-preserved, along with their
low-relief impression.
R e m a r k s (A.Yu. Ivantsov, herein). Initially Yu.A. Gureev considered the fossil as a body remain
(see description of Studenicia galeiforma Gureev, 1983).
Genus Palaeopascichnus Palij, 1976
D i a g n o s i s (Paliy, 1976). Systems of traces have an appearance of closely spaced parallel small
grooves (negative epirelief). The grooves break at the edges of system; their endings are indistinct or
rounded. Positive hyporelief is in a form of narrow, closely spaced, parallel ridges.
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 C o m p a r i s o n s (Paliy, 1976). It differs from the genus Helmintoida Schafhutl in the presence
of breaks at the edges of system, while in case of Helmintoida, the whole system represents continuous
trace, repeatedly bending in opposite directions.
Palaeopascichnus delicatus Palij, 1976
Pl. VI, fig. 7
D e s c r i p t i o n (Paliy, 1976). Negative epirelief represents series of parallel, in most cases slight-
ly arcuate, tightly packed small furrows. In positive hyporelief, there are respective ridges of similar
structure. Surface shape of ridges is arcuate in cross-section, their endings are indistinct, gradually
passing into rock surface or blunt. In some cases transverse segmentation of ridges by constrictions is
observed. The number of furrows of one series is from 4 to 10 and more.

Genus Phycodes Richter, 1850

Phycodes pedum Seilacher, 1955
Pl. VII, fig. 7
D e s c r i p t i o n . Not provided in the publications on fossils of this region.

Genus Planispiralichnus Fedonkin, 1985

Planispiralichnus rarus Menasova, 2003
Pl. VII, fig. 4 a, b
D e s c r i p t i o n (Menasova, 2003b). Trace in a form of thin smooth ridge, untightly curled as a
right-handed helice (as in electric stove). The terminations end abruptly. The height of the ridge over
the substrate is inconsiderable. The width in general is uniform throughout the trace length. There are
two spiral traces on the specimen, one of which is represented by the fragment of significantly bigger
diameter. Besides spiral traces, there are direct traces (feeding and locomotion traces), which seem to
belong to the same organism.
R e m a r k s (A.Yu. Ivantsov, herein). In paper by A.Sh. Menasova number . -8116 is provided,
the new one KTSNY, specimen no. 1741.

Genus Pseudohiemaloraichnus Gritsenko, 2009

D i a g n o s i s (Gritsenko, 2009). Grazing traces (Pascichnia) with oblique hatching and irregular
trajectory, sometimes creating coils, resembling in appearance impressions of jelly-fish, described by
M.A. Fedonkin under the name Hiemalora stellaris.
Pseudohiemaloraichnus podolica Gritsenko, 2009
Pl. VIII, fig. 4
D e s c r i p t i o n (Gritsenko, 2009). There is an oval projection (3x5 mm) at the central part of a
trace, from which thin traces of eating the substrate radially spread. The trace, 2 mm in width, moves
off (or comes to) the coil; the trace has composite path, resembling sinusoid. The trace intersects the
furrows-hatches, directed obliquely towards the animals movement.

Genus Sokolovichnites Gureev, 1983

D i a g n o s i s (Gureev, 1983b). In the positive hyporelief bioexoglyph has an appearance of curved
and sinuous ridge, ending with well-marked terminal widening. The free end of the ridge gradually
turns into the mark-bearing layer, although there is a possibility of the presence of two terminal widen-
ings on the ends of the sinuous trace.
Sokolovichnites angelicae Gureev, 1983
D e s c r i p t i o n (Gureev, 1983b). Small bioexoglyphs (positive hyporelief). The ridge is curved
and sinuous, solid, sometimes dashed. Terminal widening is sometimes cone-shaped, more frequently
flatten, with round or irregular shape in plan view, always significantly (3-6 times) bigger than the ridge.
R e m a r k s (A.Yu. Ivantsov, herein). Specimen, pictured by Yu.A. Gureev (1983b, Plate without
number, fig. 1), has the following reference: Podolia, right bank of Ternava river, near Kitaygorod vil-

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 lage, 100 m upwards the mouth of Okun Creek, Suhoy log outcrop; Rovno Horizon, Baltic Group,
lower part of Khmelnitsky Formation. In paper by Yu.A. Gureev the following place of storage and
number of the specimen is given: IGS, specimen no. -81-53. However, the specimen was not found in
IGS collections.
Sokolovichnites isp.
D e s c r i p t i o n (Gureev, 1983b). The ridge is sinuous and solid; terminal widening with round
shape in a plan view. The ridge ends at the edge of the specimen, where the presence of the second ter-
minal widening might be supposed.
C o m p a r i s o n s (Gureev, 1983b). It differs from the type species in considerably bigger siz-
es (the width of the ridge is approximately twice bigger, and the terminal widening is three times
bigger editors note).
R e m a r k s (A.Yu. Ivantsov, herein). Specimen, pictured by Yu.A. Gureev (1983b, Plate without
number, fig. 8), has the following reference: Podolia, left bank of Ternava river, near Kitaygorod village,
150 m downwards the mouth of Okun Creek; Rovno Horizon, Baltic Group, lower part of Khmelnitsky
Formation. In paper by Yu.A. Gureev the following place of storage and number of the specimen is
given: IGS, specimen no. -81-17. However, the specimen was not found in IGS collections.

Genus Treptichnus Miller, 1889

Treptichnus bifurcus Miller, 1889
Pl. VII, fig. 1
D e s c r i p t i o n . Not provided in the publications on fossils of this region.
R e m a r k s (A.Yu. Ivantsov, herein). In papers by M.A. Fedonkin number PIN, no. 3994/26-
is provided. After putting into exposition of Yu.A. Orlov Museum number was changed to PIN,
no. 4158/32.
Treptichnus triplex Palij, 1976
Pl. VII, fig. 2
D e s c r i p t i o n (Paliy, 1976). Positive hyporelief is in a form of ridges arranged at an angle of
20 to 40 to the trace longitudinal axis. Ridges of irregular width, their external ends are rounded or ta-
pered with internal ones overlapped by preceding fragments. In vertical plane ridges are slightly curved,
right-angled in a cross-section; the convex part directed downwards. Ridge surface is subdivided into
three segments of equal width by two longitudinal grooves. And the surface of each secondary ridge
is arcuate in cross-section. Arrangement of the fragments in a system is from spike- to zigzag-shaped.
C o m p a r i s o n s (Paliy, 1976). Present species differs from the other known in literature forms,
which can be attributed to genus Treptichnus in triplicate structure of fragments.
Veprina cf. undosa Fedonkin, 1980
Pl. VIII, fig. 2
D e s c r i p t i o n (Gritsenko, 2009) (in paper by V.P. Gritsenko Veprina undosa Fedonkin, 1980 is
interpreted as trace, which is opposite to authors vision of a fossil as body impression editors note).
Tracks form sinuous curves, composed of arcuate segments, which seemed to be adhered to each other
and created the chain of traces. The tracks length is from 20 to 40 mm, the width is 2-5 mm. Depres-
sions between arcuate ridge-sectors are considerably narrower, than width of the segments themselves.
In a rough comparison a trace (track) resembles a lenses column, which was put on one side.
C o m p a r i s o n s (Gritsenko, 2009). Present tracks differ from nominotypical species in simpler
path and an absence of the tracks closure. Holotype of Veprina undosa Fedonkin from Ust-Pinega
Formation has denser irregularly rounded path.
R e m a r k s (V.P. Gritsenko, herein). In the first publication specimen NMNH, no. 2480/11 (Grit-
senko, 2009, 34 p., fig. 3, Pl. I, fig. 8) was pictured. The image of the different specimen from the same
locality (NMNH, specimen no. 2525/103) is given in the present publication.

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 PALEOFLORISTIC REMAINS AND PROBLEMATICS
DESCRIBED AS ALGAE

Genus Beltanelloides Sokolov, 1965

D i a g n o s i s (Gnilovskaya et al., 1988). Large spherical envelopes, 5-30 mm in diameter, fossil-
ized as flatten phytolaims or forming impressions on the rock. Phytolaims have rounded shape, distinct
edges, concentric crumpling folds on periphery and smooth central part.
C o m p a r i s o n s (Gnilovskaya et al., 1988). It differs from other representatives of the family
Chuariacea in large sizes and smooth non-cellular envelope.
R e m a r k s (A.Yu. Ivantsov, herein). Description of Beltanelloides amorphus Menasova, 2003,
preserved as a positive impression on the bed sole without any trace of organic matter on it, is also in-
cluded in this section. This type of preservation is typical for Vendian-Ediacaran Metazoa and descrip-
tion was made by using a zoological terminology. Authors understanding of generic belonging of the
taxon and not well-established views on nature of Beltanelloides are the reasons for combining fossils,
which are so unlike, under one genera name.
Beltanelloides amorphus Menasova, 2003
Pl. I, fig. 4
D e s c r i p t i o n (Menasova, 2003b). Large imprint of umbrella of approximately round shape,
very flexible. Edge is undulating, bent inwards on some areas, forming bulges, divided by furrows.
Furrows are fragmentary, their orientation is sub-concentric. Surface of imprint is almost non-elevated;
elements of radial symmetry are absent; its whole surface has blistered appearance. There is indistinct
depression at the center, probably, related to oral aperture.
C o m p a r i s o n s (A.Sh. Menasova, herein). It differs from B. sorichevae Sokolov in more pro-
nounced peripheral part, sub-concentric direction of wrinkles, presence of bulges.
R e m a r k s (A.Yu. Ivantsov, herein). In paper by A.Sh. Menasova number . 2/49 is provided for
the specimen, the new one KTSNY, specimen no. 1742.
Beltanelloides podolicus A. Istchenko in Gnilovskaya et al., 1988
Pl. X, fig. 1
D e s c r i p t i o n (Istchenko at Gnilovskaya et al., 1988). Phytolaims with round shape, mainly
even edges, flat, of dark brown color, stand out distinctly against the grey background of mudstone.
Organic matter is slightly carbonized to the powder-like friable condition; therefore phytolaims can be
separated from the rock with difficulty, only as small fragments or grains of organic matter. Size of phy-
tolaims is different, ones with diameter 10-20 mm are dominated, but forms with diameter from 5 to 10
mm and bigger forms, with diameter up to 35 mm, also occur. Small rugosity (folding) is clearly seen on
the periphery of large phytolaims. Wrinkles are arranged sub-concentrically, get rumpled, intersect and
overlap each other. At the central part phytolaims are smooth, even, without wrinkles. Some phytolaims
have round clots of organic matter with almost black color at the central part, with diameter 0.5-1 mm.
Clots are arranged irregularly, but on one specimen four bundles are placed in one row, forming a chain.
The edges of phytolaims are mainly even, distinct, but rarely blurred, uneven. Light fringe, 1-2 mm in
width, occurs on the rock at the edge of some phytolaims. It limits phytolaims from darker rock. While
observing such specimens, wet with alcohol or xylene, it is seen under binocular that organic phyto-
laims were surrounded by thinner envelope (probably, mucous), which is preserved in fossil condition
only as faint, barely perceptible imprint on the rock.

Genus Eoholynia Gnilovskaya, 1975

D i a g n o s i s (Gnilovskaya et al., 1988). Thallus is cord-like, intensely branching and centrally
attached. Height of the bush is about 10 mm. Major branches are dividing monopodially, side branches
dichotomize. Thallus is multicellular, formed by tissue similar to parenchyma; sizes of cells are about
3 m. Sporangia are sessile and occur on major and side branches, less commonly on the tops of termi-
nal branches. They are 50-100 m in size.

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 Eoholynia capillaria A. Istchenko in Gnilovskaya et al., 1988
Pl. X, fig. 3
D e s c r i p t i o n (Gnilovskaya et al., 1988). Thallus is thread-like, consisting of numerous thin
piliform filaments, assembled in non-compact bunches, 2-3 cm in length. Filaments are arranged tightly
at the base of a bunch and arranged non-compactly and irregularly in its upper part. Filaments are
subparallel at the lower part of a bunch, subsequently and gradually bending sidewards, twisting and
interlacing. Filaments have approximately equal width along the full length, but more wide ones are
observed at the base of a bunch or thin ones at its top. The width of filaments is 0.02-0.03 mm <>.
C o m p a r i s o n s (Gnilovskaya et al., 1988). It differs from type species in length of bunches and
filaments and their arrangement in bunches. Filaments branching junction is not observed at the base
of a bunch.
Eoholynia fruticulosa A. Istchenko in Gnilovskaya et al., 1988
Pl. X, fig. 7
D e s c r i p t i o n (Gnilovskaya et al., 1988). Thallus is thread-like, numerously irregularly branch-
ing, forming numerous bushes on the substrate, 2-3 mm in height. Spread of thallus sidewards occurs
by filaments branching directly at the base of attachment to the substrate. Separate filaments of the
bush are intensively branching, while growing upwards, and give new bushes. The bedding planes of
mudstones, entirely covered with algal bushes, each of which is connected by algal filaments with the
others, are observed. The width of filaments is 20-30 m <>.
C o m p a r i s o n s (Gnilovskaya et al., 1988). It differs from type species (E. mosquensis Gnilovs-
kaya, 1975 editors note) in sizes and character (intensity) of filaments branching.
Eoholynia longa A. Istchenko in Gnilovskaya et al., 1988
D e s c r i p t i o n (Gnilovskaya et al., 1988). Long, up to 2-3 cm, isolated filaments occur on the
bedding surfaces of mudstones. Filaments are sinuous, tangled, with constant width throughout the
length 0.01-0.02 mm. Thin branches, 0.3-0.5 mm in length, extend from filaments at right angle.
Branches are bent, sinuous, sometimes overlapping main wide filament. Filaments occur on the bedding
planes as single, isolated from each other individuals; rarely 2-3 specimens are observed side-by-side.
C o m p a r i s o n s (Gnilovskaya et al., 1988). According to branching mode new species is similar
to type one, but it differs in length of filaments. Besides, branching point was not detected in new spe-
cies. Structure, similar to holdfast disc, is rarely observed. It represents small round carbonaceous crust
at the end of filament, 1-2 mm in size.
R e m a r k s (A.Yu. Ivantsov, herein). Holotype NMNH, no. 2235/14, pictured by A.A. Istchen-
ko (Gnilovskaya et al., 1988, Pl. IV, fig. 2), has the following reference: Podolia, Ushitsa river, near
Minkovtsy village; Vendian, Mogilev-Podolsky Group, Nagoryany Formation, Kalyus Beds. Fossil
cant be seen on the plate, storaged at NMNH.

Genus Fusosquamula Aseeva, 1976

D i a g n o s i s (Aseeva, 1976). Short spindle-shaped thalli, expanded at central part and tapered
at ends.
Fusosquamula vlasovi Aseeva, 1976
Pl. X, fig. 6
D e s c r i p t i o n (Aseeva, 1976). Thalli have an appearance of thin films with spindle-shaped
outlines. According to the shape they resemble short worms. The edges are smooth. Middle part is
expanded to 0.5-1 mm, edges are tapered. The length is 3-10 mm. Thalli are irregularly located on bed-
ding planes, sometimes superimposed each other, and randomly bent. Color is dark brown. Specimens,
prepared with the use of hydrofluoric acid, are similar to band-like algae by the structure of tissues.

Genus Kalusina A. Istchenko in Gnilovskaya et al., 1988

D i a g n o s i s (Gnilovskaya et al., 1988). Thallus is filamentous, branching, forming bushes of
spherical shape with branching point at the base. Filaments are intensively and frequently branching at
acute angles; branches are closely adjoined to main filaments.

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 C o m p a r i s o n s (Gnilovskaya et al., 1988). It differs from genus Eoholynia Gnilovskaya in bush
shape and mode of filaments branching.
Kalusina compacta A. Istchenko in Gnilovskaya et al., 1988
Pl. X, fig. 5
D e s c r i p t i o n (Gnilovskaya et al., 1988). Numerous round impressions of bushes and their frag-
ments occur on the bedding planes of mudstones. Bushes consist of thin filaments, branching from the
base of a bush. Branching is intensive, irregular, at acute angles to main filament. Branches do not ex-
tend sidewards, but adjoin the main filament, from which they branch out; as a result a small ball-shaped
bush, consisting of numerous thin filaments, is formed. Height of bushes is 1-2 cm, width is the same;
width of filaments is 0.02-0.03 mm. Number of filaments in bushes reaches 5-6 dozens.

Genus Kanilovia A. Istchenko, 1983

D i a g n o s i s (Istchenko, 1983). Thallus is non-mineralized, band-like (cylindrical?), short,
branching monopodially. It is tapered in lower part and club-like expanded upwards. Vegetative flat-
coiled filaments are located in one or two rows in the upper third of a thallus.
C o m p a r i s o n s (Istchenko, 1983). It differs from genus Vendotaenia Gnilovskaya in shape and
sizes of thallus, along with the reproductive organs.
Kanilovia insolita A. Istchenko, 1983
Pl. X, fig. 9
D e s c r i p t i o n (Istchenko, 1983). Thallus is band-like, probably flatten, cylindrical, straight or
bent, short, 3.5 cm in length. The shape is elongated and club-like. Width of thallus in lower part is up
to 1.2-1.4 mm, it is gradually widening up to 2 mm. The top is rounded. Side branch is sometimes ob-
served at right angle in the upper part. Width of branch is slightly smaller than width of thallus at the
branching point. Thin flat-coiled filaments, are horizontally located in one row, or rarely in two rows,
at the upper third of thallus. Thick disc, formed by tightly curled end of filament, occurs at the center of
spiral. Width of filament is 0.03-0.036 mm at disc, and up to 0.07-0.075 mm at extended opposite end
of spiral. Length of filament reaches 0.08-0.09 mm. Diameter of flat spiral is 0.1-0.3 mm (it depends on
compactness of filaments curling). On some specimens filaments are observed as uncoiled ones, on the
others as coiled ones in different extent. Coiled filaments are also observed beyond the thallus.

Genus Pilitela Aseeva, 1976

D i a g n o s i s (Aseeva, 1976). Imprints have an appearance of filamentous fibers, aggregated in
bundles. Bundles of fibers are curving, intersecting, making an impression of branching stalks.
Pilitela composita Aseeva, 1976
Pl. X, fig. 4
D e s c r i p t i o n (Aseeva, 1976). Thalli have an appearance of filamentous fibers, 0.2 mm or less
in thickness, extended in one direction, probably, in a current direction. Fibers are aggregated in bun-
dles, 0.5-1.5 mm in thickness; bundles are curving and intersecting, and therefore they make an impres-
sion of branching stalks. Color is dark brown. Filaments, prepared with the use of hydrofluoric acid,
represent non-compact filamentous thalli.
Genus Redkinia Sokolov, 1977
Redkinia fedonkini Assejeva, 1988
D e s c r i p t i o n (Aseeva, 1988b). Flat, elongated, lamellate structures formed by organic matter
of high optic density (chitin?). One margin is uneven, gradually becoming thinner. Second margin is
scalloped. Relation of teeth sizes is mainly stable in middle fragments. Semicircular crown projections
with five-six slightly curved denticles, up to 5 m in width near the base, are located between bases
of large lanceolate teeth, 150-200 m in length and 20-30 m in width, in the intervals of 50-60 m.
Length of denticles is increasing from 6 to 25 m, while coming closer to the center of a crown. Central
denticle is distinguished on its sizes, reaching 7-8 m in width and 40-45 m in length in the widest
part. Apical fragments, narrowing at the end, possess teeth, reduced both in size and number.

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 C o m p a r i s o n s (Aseeva, 1988b). <> It differs from type species R. spinosa Sokolov in sig-
nificantly smaller sizes and teeth morphology <>.
R e m a r k s (A.Yu. Ivantsov, herein). Location of the holotype: Podolian Transdniestria, borehole
no. 2 Pechera, depth of 320.0 m; Vendian, Mogilev-Podolsky Group, Yaryshev Formation, Zinkov Beds.
Specimen was pictured by E.A. Aseeva (1988b, Pl. XXI, fig. 1-3). Place of its storage is unknown.

Genus Serebrina A. Istchenko in Gnilovskaya et al., 1988

D i a g n o s i s (Gnilovskaya et al., 1988). Non-mineralized procumbent bark-like thallus, with
round shape on early stages of ontogenesis and irregularly polygonal shape with rounded or indented
edges on late stages. Type of morphological structure is close to lamellate. It consists of cells of rounded
or round-polygonal shape, non-compactly and randomly arranged within the plate.
C o m p a r i s o n s (Gnilovskaya et al., 1988). It differs from Vendotaenia Gnilovskaya in bark-like
habitus, shape and arrangement of cells.
Serebrina crustacea A. Istchenko in Gnilovskaya et al., 1988
Pl. X, fig. 2
D e s c r i p t i o n (Gnilovskaya et al., 1988). Thallus is lamellated and has an appearance of thin
carbonaceous black or dark brown crusts on the bedding surfaces of siltstones. Relief of crusts is un-
even, replicating the irregularities of the substrates relief. Numerous depressions of irregular shape
(impressions of grains of siliciclastic rock) occur on outer surface of crusts. Sizes and shapes of thalli
remains are various: from small round ones with smooth edges, 1.5-2 mm in diameter, to large irregu-
larly round ones with indented edges, lateral projections and outgrowths, 2-3 cm in length. Organic mat-
ter is strongly carbonized crumbling when separated from the substrate; it contains inclusions of rock
grains, which bear crusts. Cellular structure of the thallus is clearly visible on the preparations, derived
by method of acid-free maceration in glycerine. Cells of various shape from rounded to polygonally
rounded. They are randomly and non-compactly arranged, forming sort of a spongy matter on the bed-
ding planes of rocks. Inclusions of grains of quartz, mica and feldspar are observed between cells. Cells
walls are dense, dark brown and slightly darker in color than inner parts of cells. Diameter of cells is
from 0.005 to 0.05-0.06 mm, thickness of envelopes is 0.002-0.003 mm.

Genus Tawuia Hofmann, 1979

D i a g n o s i s (Gnilovskaya et al., 1988). Carbonaceous organic envelopes flatten, band-like, up to
8 cm in length and up to 0.6 cm in width; band edges are smooth, even and parallel. Width is constant
along the full length, bands are occasionally tapering at one end. Edges of bands are rounded. Surface
of bands is smooth.
Tawuia dalensis Hofmann, 1979
Pl. X, fig. 8
D e s c r i p t i o n (Gnilovskaya et al., 1988). Band-like carbonaceous phytolaims are short: 0.5 to
2.5 cm (much shorter, than in type specimens, described by Hoffman). Width of bands is from 0.8 to 2-3
mm, constant along the full length. Bands are straight, with smooth edges or U-shape and S-shape bent.
Surface of bands is smooth or punctulated, having impressions of grains of terrigenous rocks.

Genus Tyrasotaenia Gnilovskaya, 1971

D i a g n o s i s (Gnilovskaya, 1971). Narrow, extremely thin and flexible bands, generally twisted
and curved in rock. Surface of bands is smooth.
C o m p a r i s o n s (Gnilovskaya, 1971). Thalli of Tyrasotaenia are preserved in rock as thin film
bands of brown color, similarly to thalli of Vendotaenia. However, in case of Vendotaenia they com-
monly lie in one plane, and are unfolded along the full length and width, and strictly parallel to bedding
plane. In case of Tyrasotaenia bands are almost always curved and twisted, and therefore they seem to
be much narrower, than they were in reality. This is probably due to the fact that thalli of Vendotaenia
were coarser and leather-like, while in case of Tyrasotaenia they were more flexible and thinner. Width
of Tyrasotaenias thallus is approximately 3-4 times smaller, than of Vendotaenia. Surface of thallus

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 of Vendotaenia reveals specific structure in a form of very thin longitudinal filaments. This structure
is clearly visible both on specimens, contained in rock, and on parts of bands that were extracted by
hydrofluoric acid. Some longitudinal striation is also observed on specimens of Tyrasotaenia. However,
only longitudinal contortion folds are visible against the background of general structureless substance
of bands on specimens, extracted from the rock.
Tyrasotaenia podolica Gnilovskaya, 1971
D e s c r i p t i o n (Gnilovskaya, 1971). Thallus is preserved in a form of extremely thin film bands,
curving in a very fanciful manner. They occur as single specimens or several specimens, superimposed
in different directions. Imprints of three bands, stretching quite parallel from some common base, are
seen on one specimen. It is possible that Tyrasotaenia grew in bundles, disintegrated after plants dying.
Bands in most cases are preserved in the rock as twisted, folded and upturned ones. Real width can be
seen only on those sections, which lie parallel to bedding plane. Bands are relatively long and narrow;
length of available fragments reaches 40 mm, width is about 0.5 mm. Thin longitudinal striation is
clearly visible on surfaces of some bands; other bands show no structure. Fragments of bands, extracted
from the rock by hydrofluoric acid, probably have no structure on the surface; but longitudinal contor-
tion folds are clearly visible and numerous.
R e m a r k s (A.Yu. Ivantsov, herein). Specimen NMNH, no. 2236/27, pictured by A.A. Istchenko
(1983, Pl. XIX, fig. 5), has the following reference: Podolia, Ternava river near Kitaygorod village;
Vendian, Kanilovka Group, Studenitsa Formation, Komarovo Beds. Fossil cant be seen on the plate,
storaged at NMNH.

Genus Vendotaenia Gnilovskaya, 1971

D i a g n o s i s (Gnilovskaya et al., 1988). Thallus is ribbon-like, rarely branching; the surface is
longitudinal-fibrous. Margins of bands are distinct and straight; width of single band is constant, it var-
ies from 0.11 to 3.5 mm in various specimens. Thallus is multicellular, multiserial and multilayered. All
cells are arranged in distinct longitudinal rows, forming a tissue sort of parenchyma with cells, about
5x8 m in size, or probably prosenchyma, and an integumentary extremely small-celled tissue (?). Spo-
rangia are 50-90 m in size.
Vendotaenia antiqua Gnilovskaja, 1971
Pl. IX, fig. 3
D e s c r i p t i o n (Aseeva, 1976). Thalli have an appearance of thin brown film-like bands, ar-
ranged parallel to bedding plane. Width of bands is 1.5-2.0, length of fragments is up to 70 mm. Edges
of bands are distinct, smooth and parallel. Thallis structure is longitudinally striated, conditioned by
longitudinal thread-like bulges. Thalli are bending and overlapping each other, creating an impression
of false dichotomization (in Remarks section .. Aseeva states cases of true dichotomization of
Vendotaenia thalli, sometimes repeated editors note).

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Sokolov, B.S. Sketches of Vendian Development. Moscow: KMK Sci. Press Ltd, 1997, pp. 1154 [in Russian].
Sokolov, B.S., Ivanovsky, A.B., (ed.) Vendian System: Historical Geological and Paleontological Foundation, vol.
1 Paleontology, Moscow: Nauka, 1985a, pp. 1221 [in Russian].
Sokolov, B.S., Ivanovsky, A.B., (eds) Vendian System: Historical Geological and Paleontological Foundation, v. 2.
Stratigraphy and geological processes. Moscow: Nauka, 1985b, pp. 1238 [in Russian].
Sprigg, R.C., Early Cambrian ? jellyfishes from the Flinders Ranges, South Australia // Transact. Roy. Soc. South
Australia. 1947, no. 71, pp. 221224.
Sprigg, R.C. Early Cambrian ? jellyfishes of Ediacara, South Australia, and Mouht John, Kimberley District,
Western Australia // Transact. Roy. Soc. South Australia. 1949, no. 73, pp. 72-79.
Stashchuk, M.F. Lithological features of the Early Paleozoic deposits of the Dniester Region, Tr. Inst. Geol. Nauk
Akad. Nauk Ukr. RSR. Ser. Stratigr. Paleontol., 1958, no. 21. pp. 144 [in Ukrainian].
Sokolov, B.S. Vendian of Northern Eurasia // Arctic Geology. Proc. Second Intern. Symp. Arctic Geology. Febr.
14. 1971. San Francisco, Cal., Tulsa, Okla., USA. 1973, pp. 204218.
Velikanov, V.A. Key Section of the Vendian in Podolia // The Vendian System. HistoricalGeological and Pale-
ontological Foundation, V. 2. Stratigraphy and .geological processes. Moscow: Nauka, 1985, pp. 3567 [in
Russian].
Velikanov, V.A. Ukrainian hypostratotype of Vendian System // Geological magazine, 2011, no. 1, pp. 4249 [in
Ukrainian].
Velikanov, V.A., Aseeva, E.A., and Fedonkin, M.A., The Vendian of Ukraine. Excursion Guide. Kiev: Naukova
Dumka, 1983, pp. 1162 [in Russian].
Velikanov, V.A., Korenchuk, L.V., Kirjanov, V.V. et al. Vendian of Podolia: Guidebook for Excursion of III Inter-
national symposium on Cambrian System and Vendian-Cambrian boundary. Kiev, 1990, pp. 1129 [in Rus-
sian].
Voznesensky, O.N. On the origin of mysterious forms of microrelief on the bedding surfaces of Riphean sand-
stones, Geol. Zh. 1956, v. 16, no. 1, pp. 6972 [in Ukrainian].
Walcott, C.D. Fossil medusae // U.S. Geol. Surv. Washington, 1898, v. 30, pp. 1201.
Yatsenko, G.M., Trophimovich, N.A. Problems of Vendian of south-western margin of East-European platform //
Bulletin of Lvov University. Ser. Geology. 2001, issue 15, pp. 135143 [in Ukrainian].

121

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 Zaika-Novatsky, V.S. New problematic imprints from the Upper Precambrian of the Dniester Region // All-Union
Symposium on Paleontology of the Precambrian and Early Cambrian, Novosibirsk: Inst. Geol. Geofiz. Sib.
Otd. Akad. Nauk SSSR, 1965, pp. 9899 [in Russian].
Zaika-Novatsky, V.S. Paleontological and palynological characteristics of the Upper Proterozoic deposits of the
Dniester Region // Stratigraphy of the Ukrainian SSR, vol. 2: RipheanVendian, Kiev: Naukova Dumka,
1971, pp. 158163 [in Ukrainian].
Zaika-Novatsky, V.S., Velikanov, V.A., Koval A.P. First representative of Ediacaran fauna in Vendian of Russian
platform (Upper Precambrian) // Paleontological Journal, 1968, no. 2, pp. 133134 [in Russian].
Zaika-Novatsky, V.S., Paliy, V.M. New data on problematic imprints from the Vendian deposits of Podolia, Pale-
ontol. Sb., 1968, v. 5, no. 1, pp. 130135 [in Russian].
Zaika-Novatsky, V.S., Paliy, V.M. Most ancient fossil organisms in the deposits of Vendian complex of the Dnies-
ter Region, Paleontol. Sb., 1974, v. 11, no. 1, pp. 5965 [in Russian].

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.
1 .

PLATES AND CAPTIONS

Macrofossils of the Upper Vendian and Lower Cambrian

of Middle Dniester area and Volhynia.
Length of the scale bar 1 cm.

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 I
. 1. Beltanella velikania Menasova, 2003; , 17134 (, 2003, ,
. 6.); , . , ; , - , -
, .
. 2. Cyclomedusa serebrina Palij, 1969; , 176 (, 1969, . 112, . 2; 1976, . I,
. 4; ., 1979, . XLVIII, . 4); , . -,
. , . , 800 . ; , - ,
, .
. 3. Gureevella elliptica Menasova, 2003; , 17150 (, 2003, ,
. 5); , . , ; , - , -
, .
. 4. Beltanelloides amorphus Menasova, 2003; , 1742 (, 2003, . 1); ,
. , ; , - , ,
.
. 5. Gritcenia nana Menasova, 2003; , 1768 (, 2003, . 4); -
, . -, . , ; , - ,
, .
. 6. Tirasiana disciformis Palij, 1976; , 1722 (, 1976, . XXII, . 4;
., 1979, . XLIX, . 3); , . (); , - ,
, .
. 7. Tirasiana coniformis Palij, 1976; , 1710 (, 1976, . XXIII, . 3; .,
1979, . XLIX, . 2); - -, . ; , - ,
, .
. 8. Cyclomedusa plana Glaessner et Wade, 1966; . , . .. -, 179 (--
., 1968, . 1; , 1972, . I, . 1; 1997, . XVII, . 4; Sokolov, 1973, fig. 2-1); ,
-, . ; , - , , -
.
Plate I
Fig. 1. Beltanella velikania Menasova, 2003; holotype KTSNY, specimen no. 17134 (Menasova, 2003, Plate
without number, fig. 6); Podolia, left bank of Dniester river, Dnestrovskaya Hydroelectric Power Station; Vendian,
Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 2. Cyclomedusa serebrina Palij, 1969; holotype KTSNY, specimen no. 176 (Palij, 1969, p. 112, fig. 2; Palij,
1976, Pl. I, fig. 4; Palij et al., 1979, Pl. XLVIII, fig. 4); Podolia, vicinity of Mogilev-Podolsky, left bank of Dniester
river, Serebria village, ravine 800 m downstream from mouth of Serebria river; Vendian, Mogilev-Podolsky Group,
Yaryshev Formation, Bernashevka Beds.
Fig. 3. Gureevella elliptica Menasova, 2003; holotype KTSNY, specimen no. 17150 (Menasova, 2003, Plate
without number, fig. 5); Podolia, left bank of Dniester river, Dnestrovskaya Hydroelectric Power Station; Vendian,
Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 4. Beltanelloides amorphus Menasova, 2003; holotype KTSNY, specimen no. 1742 (Menasova, 2003b,
fig. 1); Podolia, left bank of Dniester river, Dnestrovskaya Hydroelectric Power Station; Vendian, Mogilev-Podolsky
Group, Mogilev Formation, Lomozov Beds.
Fig. 5. Gritcenia nana Menasova, 2003; holotype KTSNY, specimen no. 1768 (Menasova, 2003b, fig. 4); Podolia,
Vinnitsa region, town of Mogilev-Podolsky, left bank of Derlo river, Borshchov Yar; Vendian, Mogilev-Podolsky Group,
Yaryshev Formation, Bronnitsa Beds.
Fig. 6. Tirasiana disciformis Palij, 1976; holotype KTSNY, specimen no. 1722 (Palij, 1976, Pl. XXII, fig. 4; Palij
et al., 1979, Pl. XLIX, fig. 3); Moldovian Dniester area, Otach (Ataki) village; Vendian, Mogilev-Podolsky Group, Yary-
shev Formation, Bernashevka Beds.
Fig. 7. Tirasiana coniformis Palij, 1976; holotype KTSNY, specimen no. 1710 (Palij, 1976, Pl. XXIII, fig. 3; Palij
et al., 1979, Pl. XLIX, fig. 2); Mogilev-Podolsky area, Serebria village; Vendian, Mogilev-Podolsky Group, Yaryshev
Formation, Bernashevka Beds.
Fig. 8. Cyclomedusa plana Glaessner et Wade, 1966; specimen KTSNY, collection of V.S. Zaika-Novatsky, speci-
men no. 179 (Zaika-Novatsky et al., 1968, fig. 1; Sokolov, 1972, Pl. I, fig. 1; 1997, Pl. XVII, fig. 4; Sokolov, 1973,
fig. 2-1); Podolia, vicinity of Mogilev-Podolsky, Serebria village; Vendian, Mogilev-Podolsky Group, Yaryshev Forma-
tion, Bernashevka Beds.

124

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 /Plate I

3
1 2

4 5

6 7 8

125

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 II
. 1. Pollukia shulgae Gureev, 1987; , 2127/55 ( ., 1985, . 2 ; , 1987, . 49,
. 20; 1988, . VI, . 6); , . , . 3642; , ,
, .
. 2. Kaisalia levis Gureev, 1987; , 2127/71 (, 1987, . 51, . 21); ,
. , . , ; , - , , -
.
. 3. Vendella haelenicae Gureev, 1987; , 2127/49 (, 1987, . 35, . 12; 1988, . V,
. 3); , . -, . , ; , - -
, , .
. 4. Paliella pateliformis Fedonkin, 1980; . , 3994/5 (, 1983, . XXVIII, . 5; 1984, .
III, . 3; 1985, . X, . 5); , . , ; , -
, , .
. 5. Irridinitus multiradiatus Fedonkin, 1983; , 3994/524 (, 1983. . XXIX, . 10;
1985, . VII, . 5); , . , ; , --
, , .
. 6. Elasenia aseevae Fedonkin, 1983; , 3994/338 (, 1983, . XXXII, . 4; 1984, . V,
. 2; 1985, . VII, . 8; Fedonkin, 1992a, fig. 2 d; , 1997, . XVII, . 3); ,
. , ; , - , , .
. 7. Ediacaria sp.; . , 3994/334 (, 1983, . XXXII, . 6; 1985, . II, . 2);
, . , ; , - , ,
.
. 8. Cyclomedusa cf. davidi Sprigg, 1947; . , 3994/370 (, 1983, . XXVIII, . 3; 1985,
. III, . 10); , . , ; , -
, , .
. 9. Glaessneria imperfecta Gureev, 1987; , 2127/56 (, 1987, . 39, . 15; 1988, .
X, . 7); , . -, . , ; , -
, , .
. 10. Nimbia dniesteri Fedonkin, 1983; , 3994/385- (, 1983, . XXIX, . 4; 1984,
. III, . 1; 1985, . III, . 2; Fedonkin, 1992b, fig. 6; , 1997, . XVI, . 2); ,
. , ; , - , , .
Plate II
Fig. 1. Pollukia shulgae Gureev, 1987; holotype NMNH, specimen no. 2127/55 (Gureev et al., 1985, fig. 2 ; Gureev,
1987, 49 p., fig. 20; Gureev, 1988, Pl. VI, fig. 6); Ternopol region, Novoselka village, borehole no. 3642; Cambrian, Lontova
Horizon, Baltic Group, Zbruch Formation.
Fig. 2. Kaisalia levis Gureev, 1987; holotype NMNH, specimen no. 2127/71 (Gureev, 1987, p. 51, fig. 21); Vinnitsa re-
gion, right bank of Lyadova river, Vinozh village, Popeluhovsky Yar; Vendian, Mogilev-Podolsky Group, Mogilev Formation,
Lomozov Beds.
Fig. 3. Vendella haelenicae Gureev, 1987; holotype NMNH, specimen no. 2127/49 (Gureev, 1987, 35 p., fig. 12; Gureev,
1988, Pl. V, fig. 3); Vinnitsa region, town of Mogilev-Podolsky, left bank of Derlo river, Borshchov Yar; Vendian, Mogilev-
Podolsky Group, Yaryshev Formation, Bronnitsa Beds.
Fig. 4. Paliella pateliformis Fedonkin, 1980; PIN, specimen no. 3994/5 (Fedonkin, 1983, Pl. XXVIII, fig. 5; 1984, Pl. III,
fig. 3; 1985 b, Pl. X, fig. 5); Middle Dniester area, right bank of Dniester river, Dnestrovskaya Hydroelectric Power Station;
Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 5. Irridinitus multiradiatus Fedonkin, 1983; holotype PIN, specimen no. 3994/524 (Fedonkin, 1983.
Pl. XXIX, fig. 10; 1985b, Pl. VII, fig. 5); Middle Dniester area, right bank of Dniester river, Dnestrovskaya Hydroelectric
Power Station; Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 6. Elasenia aseevae Fedonkin, 1983; holotype PIN, specimen no. 3994/338 (Fedonkin, 1983, Pl. XXXII,
fig. 4; 1984, Pl. V, fig. 2; 1985b, Pl. VII, fig. 8; Fedonkin, 1992a, fig. 2 d; Sokolov, 1997, Pl. XVII, fig. 3); Middle Dniester area,
right bank of Dniester river, Dnestrovskaya Hydroelectric Power Station; Vendian, Mogilev-Podolsky Group, Mogilev Forma-
tion, Lomozov Beds.
Fig. 7. Ediacaria sp.; PIN, specimen no. 3994/334 (Fedonkin, 1983, Pl. XXXII, fig. 6; Fedonkin, 1985b, Pl. II, fig. 2);
Middle Dniester area, right bank of Dniester river, Dnestrovskaya Hydroelectric Power Station; Vendian, Mogilev-Podolsky
Group, Mogilev Formation, Lomozov Beds.
Fig. 8. Cyclomedusa cf. davidi Sprigg, 1947; PIN, specimen no. 3994/370 (Fedonkin, 1983, Pl. XXVIII, fig. 3; Fedonkin,
1985b, Pl. III, fig. 10); Middle Dniester area, right bank of Dniester river, Dnestrovskaya Hydroelectric Power Station; Ven-
dian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 9. Glaessneria imperfecta Gureev, 1987; holotype NMNH, specimen no. 2127/56 (Gureev, 1987, p. 39, fig. 15;
Gureev, 1988, Pl. X, fig. 7); Vinnitsa region, town of Mogilev-Podolsky, left bank of Derlo river, Borshchov Yar; Vendian,
Mogilev-Podolsky Group, Yaryshev Formation, Bronnitsa Beds.
Fig. 10. Nimbia dniesteri Fedonkin, 1983; holotype PIN, specimen no. 3994/385- (Fedonkin, 1983, Pl. XXIX, fig. 4;
1984, Pl. III, fig. 1; 1985b, Pl. III, fig. 2; Fedonkin, 1992b, fig. 6; Sokolov, 1997, Pl. XVI, fig. 2); Middle Dniester area, right
bank of Dniester river, Dnestrovskaya Hydroelectric Power Station; Vendian, Mogilev-Podolsky Group, Mogilev Formation,
Lomozov Beds.

126

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 /Plate II

1 3
2

6
4

8 9 10

127

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 III
. 1. Bronicella podolica (Zaika-Novatsky, 1965); , 1724 (, 1976, . XXI, . 1);
., . ; , - , , .
. 2. Nemiana simplex Palij, 1976; , 1739 (, 1976, . XXII, . 1,
., 1979, . XLIX, . 1); (, 1987); , . ; , -
- , , .
. 3. Elasenia zhuravlevae Gureev, 1988; , 2127/65 ( ., 1985, . 1 ; ,
1988, . X, . 5); , . , . ; (?), , -
.
. 4. Medusinites asteroides (Sprigg, 1949); . , 2127/16 (, 1987, . 31, . 10; 1988, .
IX, . 2); , . , . , ; , --
, , .
. 5. Sekwia kaptarenkoe Gureev, 1987; , 2127/11 (, 1987, . 32, . 11; 1988, .
VIII, . 4); , . , . , ; , --
, , .
. 6. Vaveliksia velikanovi Fedonkin, 1983; , 3994/581 (, 1983, . XXXII, .
2; 1985, . XII, . 1); , . , ; , -
, , .
. 7. Medusinites paliji Gureev, 1987; . , 2127/19 (, 1988, . VIII, . 2):
, ( ); , 5
; , - , , .

Plate III
Fig. 1. Bronicella podolica (Zaika-Novatsky, 1965); holotype KTSNY, specimen no. 1724 (Palij, 1976, Pl. XXI,
fig. 1); Vinnitsa region, Yastrebna village; Vendian, Mogilev-Podolsky Group, Yaryshev Formation, Bronnitsa Beds.
Fig. 2. Nemiana simplex Palij, 1976; holotype KTSNY, specimen no. 1739 (Palij, 1976); (Palij, 1976, Pl. XXII,
fig. 1, Palij et al., 1979, Pl. XLIX, fig. 1); lectotype upper imprint (Gureev, 1987); Podolia, Ozarintsy village; Vendian,
Mogilev-Podolsky Group, Mogilev Formation, Yampol Beds.
Fig. 3. Elasenia zhuravlevae Gureev, 1988; holotype NMNH, specimen no. 2127/65 (Gureev et al., 1985, fig. 1
b; Gureev, 1988, Pl. X, fig. 5); Podolia, right bank of Ternava river, Kitaygorod village; Cambrian(?), Rovno Horizon,
Khmelnitsky Formation.
Fig. 4. Medusinites asteroides (Sprigg, 1949); NMNH, specimen no. 2127/16 (Gureev, 1987, p. 31, fig. 10. Gureev,
1988, Pl. IX, fig. 2); Vinnitsa region, right bank of Lyadova river, Vinozh village, Popeluhovsky Yar; Vendian, Mogilev-
Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 5. Sekwia kaptarenkoe Gureev, 1987; holotype NMNH, specimen no. 2127/11 (Gureev, 1987, 32 p., fig. 11;
Gureev, 1988, Pl. VIII, fig. 4); Vinnitsa region, right bank of Lyadova river, Vinozh village, Popeluhovsky Yar; Vendian,
Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 6. Vaveliksia velikanovi Fedonkin, 1983; holotype PIN, specimen no. 3994/581 (Fedonkin, 1983,
Pl. XXXII, fig. 2; 1985b, Pl. XII, fig. 1); Middle Dniester area, right bank of Dniester river, Dnestrovskaya Hydroelectric
Power Station; Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 7. Medusinites paliji Gureev, 1987; NMNH, specimen no. 2127/19 (Gureev, 1988, Pl. VIII, fig. 2):
a general view of the plate, b lectotype (large imprint at the center); Vinnitsa region, right bank of Nemija river,
5 km below the river mouth; Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.

128

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 /Plate III

5
4

7
7

129

Atlas_2015.indd 129 25-04-2015 14:18

 IV
. 1. Conomedusites cf. lobatus Glaessner et Wade, 1966; , . 4158/29,
. .. (, 1983, . XXXI, . 1; 1984, . VI, . 4; 1985, . VIII, . 1; 1987,
VIII; Fedonkin, 1992b, fig. 24; , 1997, . XVII, . 1); , . ,
; , - , , .
. 2. Jampolium wyrzhykowskii Gureev, 1988; , 2127/58 (, 1988, . VI, . 3);
. , ; , - , ,
.
. 3. Vendella larini Gureev, 1987; , 2089/27 (, 1983, , . 2;
1985, . XXXVIII, . 2; 1987, . 35, . 13; 1988, . IV, . 8); , . ,
. ; , , , .
. 4. Nimbia paula Gureev, 1985; , 2089/23 (, 1985, . XXXVIII, . 3; 1988,
. VIII, . 7); , . , . ; , -
, , .
. 5. Kullingia concentrica Glaessner, 1979; . , 2089/19 (, 1985, . XXXIX, . 1; 1988,
. I, . 5; ., 1985, . 1; , 1997, . XVIII, . 1); , . ,
. (. ); (?), , , .
. 6. Cyclomedusa minuta Fedonkin in Palij et al., 1979; . , 2127/27 ( ) ( .,
1985, . 1 ; , 1988, . VII, . 9); ; (?), , .
. 7. Medusinites sokolovi Gureev, 1985; , 2089/24 (, 1985, . XXXVIII, . 4;
1988, . VIII, . 6); , . , . , ; , -
- , , .
. 8. Bessarabia bukatchuki Gureev, 1988; , 2127/60 (, 1988, . X, . 3);
, . 27, . 249 ; , - , ;
.. .
. 9. Protodipleurosoma wardi Sprigg, 1947; . , 2127/72 (, 1988, . XIII, . 1); -
, . , . , ; , - , -
, .
Plate IV
Fig. 1. Conomedusites cf. lobatus Glaessner et Wade, 1966; PIN, specimen no. 4158/29, exposition of Yu.A. Orlov
Museum (Fedonkin, 1983, Pl. XXXI, fig. 1; 1984, Pl. VI, fig. 4; 1985b, Pl. VIII, fig. 1; 1987, image VIII; Fedonkin, 1992b,
fig. 24; Sokolov, 1997, Pl. XVII, fig. 1); Middle Dniester area, right bank of Dniester river, Dnestrovskaya Hydroelectric
Power Station; Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 2. Jampolium wyrzhykowskii Gureev, 1988; holotype NMNH, specimen no. 2127/58 (Gureev, 1988, Pl. VI,
fig. 3); left bank of Dniester river, downstream the dam of Dnestrovskaya Hydroelectric Power Station; Vendian, Mogi-
lev-Podolsky Group, Mogilev Formation, upper Yampol Beds.
Fig. 3. Vendella larini Gureev, 1987; holotype NMNH, specimen no. 2089/27 (Gureev, 1983a, Plate without num-
ber, fig. 2; 1985, Pl. XXXVIII, fig. 2; 1987, 35 p., fig. 13; 1988, Pl. IV, fig. 8); Khmelnitsky region, Sokolets village, right
bank of Ushitsa river; Vendian, Kanilovka Group, Zharnovka Formation, Kuleshovka Beds.
Fig. 4. Nimbia paula Gureev, 1985; holotype NMNH, specimen no. 2089/23 (Gureev, 1985, Pl. XXXVIII, fig. 3.
Gureev, 1988, Pl. VIII, fig. 7); Khmelnitsky region, Pilipy village, right bank of the Danilov Creek; Vendian, Mogilev-
Podolsky Group, Nagoryany Formation, Kalyus Beds.
Fig. 5. Kullingia concentrica Glaessner, 1979; NMNH, specimen no. 2089/19 (Gureev, 1985, Pl. XXXIX, fig. 1. Gureev,
1988, Pl. I, fig. 5; Gureev et al., 1985, fig. 1 a; Sokolov, 1997, Pl. XVIII, fig. 1); Podolia, right bank of Ternava river, Kitaygorod
village (Suhoy log outcrop); Cambrian(?), Rovno Horizon, Baltic Group, Khmelnitsky Formation.
Fig. 6. Cyclomedusa minuta Fedonkin in Palij et al., 1979; NMNH, specimen no. 2127/27 (marked with an arrow) (Gureev
et al., 1985, fig, 1 ; Gureev, 1988, Pl. VII, fig. 9); Podolia; Cambrian(?), Rovno Horizon, Khmelnitsky Formation.
Fig. 7. Medusinites sokolovi Gureev, 1985; holotype NMNH, specimen no. 2089/24 (Gureev, 1985, Pl. XXXVIII,
fig. 4. Gureev, 1988, Pl. VIII, fig. 6); Vinnitsa region, right bank of Lyadova river, Vinozh village, Popeluhovsky Yar;
Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 8. Bessarabia bukatchuki Gureev, 1988; holotype NMNH, specimen no. 2127/60 (Gureev, 1988, Pl. X, fig. 3);
Moldavian Transdniestria, borehole no. 27, depth of 249 m; Vendian, Mogilev-Podolsky Group, Mogilev Formation;
collected by P.D. Bykatchuk.
Fig. 9. Protodipleurosoma wardi Sprigg, 1947; NMNH, specimen no. 2127/72 (Gureev, 1988, Pl. XIII, fig. 1);
Vinnitsa region, right bank of Lyadova river, Vinozh village, Popeluhovsky Yar; Vendian, Mogilev-Podolsky Group,
Mogilev Formation, Lomozov Beds.

130

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 /Plate IV

1
2

6 7

8 9

131

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 V
. 1. Palaeospinther conoideus Menasova, 2003; , 1781 (, 2003, -
, . 2, 3, 4); , . -, . , ; , -
, , .
. 2. Palaeospinther nucis Menasova, 2003; , 17151 (, 2003, ,
. 1); , . -, . , ; , --
, , .
. 3. Tribrachidium heraldicum Glaessner, 1959; . , 4158/26 (, 1983, . XXIX, . 5;
1984, . VI, . 8; 1985, . IX, . 3); , . ,
; , - , , .
. 4. Valdainia plumosa Fedonkin, 1983; , 3994/276 (, 1983, . XXX, . 4;
1985, . X, . 1; , 1997, . XX, . 1); , . , -
; , - , , .
. 5. Podolimirus mirus Fedonkin; , 3994/417 (, 1983, . XXX, . 6; 1985,
. X, . 4); , . , ; , -
, , .
. 6. Vaveliksia svetozarovae Gureev, 1988; , 2127/63 (, 1988, . X, . 2);
, . , . , ; , - ,
, .
. 7. Kamenecia stella Gureev, 1988; , 2127/67 (, 1988, . XI, . 4); ,
. , . ; (?), , , .
. 8. Ternavellus vialovi Gureev, 1988 (=aff. Kimberella sp.); , 2127/66 ( ., 1985,
. 2 ; , 1988, . X, . 8); , . , . , ;
(?), , , .

Plate V
Fig. 1. Palaeospinther conoideus Menasova, 2003; holotype KTSNY, specimen no. 1781 (Menasova, 2003, Plate
without number, fig. 2, 3, 4); Vinnitsa region, town of Mogilev-Podolsky, left bank of Derlo river, Borshchov Yar; Ven-
dian, Mogilev-Podolsky Group, Yaryshev Formation, Bronnitsa Beds.
Fig. 2. Palaeospinther nucis Menasova, 2003; holotype KTSNY, specimen no. 17151 (Menasova, 2003, Plate
without number, fig. 1); Vinnitsa region, town of Mogilev-Podolsky, left bank of Derlo river, Borshchov Yar; Vendian,
Mogilev-Podolsky Group, Yaryshev Formation, Bronnitsa Beds.
Fig. 3. Tribrachidium heraldicum Glaessner, 1959; PIN, specimen no. 4158/26 (Fedonkin, 1983, Pl. XXIX, fig. 5;
1984, Pl. VI, fig. 8; Fedonkin, 1985b, Pl. IX, fig. 3); Middle Dniester area, right bank of Dniester river, Dnestrovskaya
Hydroelectric Power Station; Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 4. Valdainia plumosa Fedonkin, 1983; holotype PIN, specimen no. 3994/276 (Fedonkin, 1983, Pl. XXX, fig. 4;
1985b, Pl. X, fig. 1; Sokolov, 1997, Pl. XX, fig. 1); Middle Dniester area, right bank of Dniester river, Dnestrovskaya Hydro-
electric Power Station; Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 5. Podolimirus mirus Fedonkin, 1983; holotype PIN, specimen no. 3994/417 (Fedonkin, 1983, Pl. XXX, fig.
6; 1985b, Pl. X, fig. 4); Middle Dniester area, right bank of Dniester river, Dnestrovskaya Hydroelectric Power Station;
Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 6. Vaveliksia svetozarovae Gureev, 1988; holotype NMNH, specimen no. 2127/63 (Gureev, 1988, Pl. X,
fig. 2); Vinnitsa region, right bank of Lyadova river, Vinozh village, Popeluhovsky Yar; Vendian, Mogilev-Podolsky
Group, Mogilev Formation, Lomozov Beds.
Fig. 7. Kamenecia stella Gureev, 1988; holotype NMNH, specimen no. 2127/67 (Gureev, 1988, Pl. XI, fig. 4);
Podolia, right bank of Ternava river, Kitaygorod village; Cambrian(?), Rovno Horizon, Baltic Group, Khmelnitsky
Formation.
Fig. 8. Ternavellus vialovi Gureev, 1988 (=aff. Kimberella sp.); holotype NMNH, specimen no. 2127/66 (Gureev
et al., 1985, fig. 2 b; Gureev, 1988, Pl. X, fig. 8); Podolia, right bank of Ternava river, Kitaygorod village, Suhoy log out-
crop; Cambrian(?), Rovno Horizon, Baltic Group, Okunets Formation.

132

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 /Plate V

1 2

7 8

133

Atlas_2015.indd 133 25-04-2015 14:18

 VI
. 1. Dickinsonia costata Sprigg, 1947; . , 3994/564 (, 1983, . XI, . 3); -
, . , ; , - ,
, .
. 2. Dickinsonia cf. tenuis Glaessner et Wade, 1969; . , 3994/418- (, 1983, . XII,
. 5; 1985, . XXXII, . 1; 1985, . XVI, . 4; Fedonkin, 1992b, fig. 36); , -
. , ; , - , , .
. 3. Marnium cristatum Gureev, 1988; , 2127/54 (, 1988, . XI, . 1); -
, . -, . , ; , - ,
, .
. 4. Hiemalora cf. stellaris Fedonkin, 1980; . , 3994/302- (, 1983, . XXIX, . 3;
1984, . V, . 5; Fedonkin, 1992a, fig. 1 c.); , . ,
; , - , , .
. 5. Lomosovis malus Fedonkin, 1983; , 3994/418- (, 1983, . XXXIII, . 1,
2; 1985, . XV, . 1, 2; , 1997, . XVIII, . 4); , . ,
; , - , , .
. 6. Harlaniella podolica Sokolov, 1972; . , 1907/5 (, 1976, . XXIV, . 1;
., 1979, . L, . 1); ., . ; , , . 12639.
. 7. Palaeopascichnus delicatus Palij, 1976; , 1907/7 ( ) (,
1976, . XXIV, . 2; ., 1979, . L, . 4); , .
. , 1 ; , ,
1 .

Plate VI
Fig. 1. Dickinsonia costata Sprigg, 1947; PIN, specimen no. 3994/564 (Fedonkin, 1983, Pl. XI, fig. 3); Middle
Dniester area, right bank of Dniester river, Dnestrovskaya Hydroelectric Power Station; Vendian, Mogilev-Podolsky
Group, Mogilev Formation, Lomozov Beds.
Fig. 2. Dickinsonia cf. tenuis Glaessner et Wade, 1966; PIN, specimen no. 3994/418- (Fedonkin, 1983, Pl. XII,
fig. 5; Fedonkin, 1985a, Pl. XXII, fig. 1; Fedonkin, 1985b, Pl. XVI, fig. 4; Fedonkin, 1992b, fig. 36); Middle Dnie-
ster area, right bank of Dniester river, Dnestrovskaya Hydroelectric Power Station; Vendian, Mogilev-Podolsky Group,
Mogilev Formation, Lomozov Beds.
Fig. 3. Marnium cristatum Gureev, 1988; holotype NMNH, specimen no. 2127/54 (Gureev, 1988, Pl. XI, fig. 1);
Vinnitsa region, town of Mogilev-Podolsky, left bank of Derlo river, Borshchov Yar; Vendian, Mogilev-Podolsky Group,
Yaryshev Formation, Bronnitsa Beds.
Fig. 4. Hiemalora cf. stellaris Fedonkin, 1980; PIN, specimen no. 3994/302- (Fedonkin, 1983, Pl. XXIX, fig. 3;
1984, Pl. V, fig. 5; Fedonkin, 1992a, fig. 1 c); Middle Dniester area, right bank of Dniester river, Dnestrov-skaya Hydro-
electric Power Station; Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 5. Lomosovis malus Fedonkin, 1983; holotype PIN, specimen no. 3994/418- (Fedonkin, 1983, Pl. XXXIII,
fig. 1, 2; 1985b, Pl. XV, fig. 1, 2; Sokolov, 1997, Pl. XVIII, fig. 4); Middle Dniester area, right bank of Dniester river,
Dnestrovskaya Hydroelectric Power Station; Vendian, Mogilev-Podolsky Group, Mogilev Formation, Lomozov Beds.
Fig. 6. Harlaniella podolica Sokolov, 1972; NMNH, specimen no. 1907/5 (Palij, 1976, Pl. XXIV, fig. 1; Palij et al.,
1979, Pl. L, fig. 1); Khmelnitsky region, Chovguzov village; Vendian, Kanilovka Group, borehole no. 12639.
Fig. 7. Palaeopascichnus delicatus Palij, 1976; holotype NMNH, specimen no. 1907/7 (marked with an arrow)
(Palij, 1976, Pl. XXIV, fig. 2; Palij et al., 1979, Pl. L, fig. 4), Middle Dniester area, right bank of Dniester river, Molodovo
village, gully in the right slope of ravine, 1 km southwards the church; Vendian, Kanilovka Group, Komarovo Beds,
1 m from the base of Ordovician deposits.

134

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 /Plate VI

7
6

135

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 VII
. 1. Treptichnus bifurcus Miller, 1889; . , 4158/32 (, 1983, . XXXIV, . 6; 1985,
. XXII, . 5); , . . ; (?), -
, , .
. 2. Treptichnus triplex Palij, 1976; , 1907/13 (, 1968, . VI, . 3; ,
1976, . XXIV, . 3; ., 1979, . LI, . 4); , . . -; (?),
, , .
. 3. Cochlichnus sp.; . , 1907/55 (, 1976, . XXVI, . 5; ., 1979, .
LIV, . 3); , . . ; (?),
, , .
. 4. Planispiralichnus rarus Menasova, 2003; , 1741 (, 2003, . 3): ,
; , . , . ; (?), , , -
.
. 5. Bergaueria major Palij, 1976; , 1907/67 (, 1976, . XXVIII, . 1, ; -
., 1979, . LV, . 2, ); , . . ; (?),
, , .
. 6. Didymaulichnus tirasensis Palij, 1974; , 1831/7 (, 1974, . 500, . 1;
., 1979, . LII, . 1, 2); , . . ; (?), -
, , .
. 7. Phycodes pedum Seilacher, 1955; . , 3994/26 (, 1983, . XXXIV, . 3; 1985,
. XXII, . 4); , . . ; (?), -
, , .

Plate VII
Fig. 1. Treptichnus bifurcus Miller, 1889; PIN, specimen no. 4158/32 (Fedonkin, 1983, Pl. XXXIV, fig. 6; 1985b,
Pl. XXII, fig. 5); Middle Dniester area, right bank of Dniester river, opposite the mouth of Ternava river; Cambrian(?),
Rovno Horizon, Baltic Group, Khmelnitsky Formation.
Fig. 2. Treptichnus triplex Palij, 1976; holotype NMNH, specimen no. 1907/13 (Kirjanov, 1968, Pl. VI, fig. 3; Palij,
1976, Pl. XXIV, fig. 3; Palij et al., 1979, Pl. LI, fig. 4); Podolia, borehole in the town of Kamenets-Podolsky; Cambrian(?),
Rovno Horizon, Baltic Group, Khmelnitsky Formation.
Fig. 3. Cochlichnus isp.; NMNH, specimen no. 1907/55 (Palij, 1976, Pl. XXVI, fig. 5; Palij et al., 1979, Pl. LIV,
fig. 3); Middle Dniester area, right bank of Dniester river, opposite the mouth of Ternava river; Cambrian(?), Rovno
Horizon, Baltic Group, Khmelnitsky Formation.
Fig. 4. Planispiralichnus rarus Menasova, 2003; holotype KTSNY, specimen no. 1741 (Menasova, 2003b,
fig. 3): detail, b general view; Podolia, Ternava river, Kitaygorod village; Cambrian(?), Rovno Horizon, Baltic
Group, Khmelnitsky Formation.
Fig. 5. Bergaueria major Palij, 1976; holotype NMNH, specimen no. 1907/67 (Palij, 1976, Pl. XXVIII, fig. 1a, ;
Palij et al., 1979, Pl. LV, fig. 2a, ); Middle Dniester area, right bank of Dniester river, near Suboch village; Cambrian(?),
Rovno Horizon, Baltic Group, Khmelnitsky Formation.
Fig. 6. Didymaulichnus tirasensis Palij, 1974; holotype NMNH, specimen no. 1831/7 (Palij, 1974, 500 p., fig. 1;
Palij et al., 1979, Pl. LII, fig. 1, 2); Middle Dniester area, right bank of Dniester river, near Suboch village; Cambrian(?),
Rovno Horizon, Baltic Group, Khmelnitsky Formation.
Fig. 7. Phycodes pedum Seilacher, 1955; PIN, specimen no. 3994/26 (Fedonkin, 1983, Pl. XXXIV, fig. 3; Fedonkin,
1985, Pl. XXII, fig. 4); Middle Dniester area, right bank of Dniester river, opposite the mouth of Ternava river; Cam-
brian(?), Rovno Horizon, Baltic Group, Khmelnitsky Formation.

136

Atlas_2015.indd 136 25-04-2015 14:18

 /Plate VII

1 4

2
3

6 7

137

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 VIII
. 1. Podoliina crassa Gureev, 1988; , 2127/64 (, 1988, . XII, . 3);
, . -, . , ; , -
, , .
. 2. Veprina cf. undosa Fedonkin; . , 2525/103; , . --
, . , ; , - , ,
.
. 3. Aviculaichnus gureevi Gritsenko, 2009; , 2525/93 (, 2009, . 33, . 1,
. I, . 1-3); , . -, . , ; ,
- , , .
. 4. Pseudohiemaloraichnus podolica Gritsenko, 2009; , 2480/10 (, 2009,
. 34, . 2, . II, . 1, 2); , . -, . ,
; , - , , .
. 5. Studenicia galeiforma Gureev, 1983 =Monocraterion sp.; , 2088/9 (,
1983, , . 1; 1984, . 1 ); , . . ; ,
, , .
. 6. ; . , 1714 (-
, 1976, . XXXIII, . 4, ., 1979, . LI, . 1); ,
. , . (); , - , , .

Plate VIII
Fig. 1. Podoliina crassa Gureev, 1988; holotype NMNH, specimen no. 2127/64 (Gureev, 1988, Pl. XII,
fig. 3); Vinnitsa region, town of Mogilev-Podolsky, left bank of Derlo river, Borshchov Yar; Vendian, Mogilev-
Podolsky Group, Yaryshev Formation, Bronnitsa Beds.
Fig. 2. Veprina cf. undosa Fedonkin; NMNH, specimen no. 2525/103; Vinnitsa region, town of Mogilev-
Podolsky, left bank of Derlo river, Borshchov Yar; Vendian, Mogilev-Podolsky Group, Yaryshev Formation,
lower part of Bronnitsa Beds.
Fig. 3. Aviculaichnus gureevi Gritsenko, 2009; holotype NMNH, specimen no. 2525/93 (Gritsenko, 2009,
33 p., fig. 1, Pl. I, fig. 1-3); Vinnitsa region, town of Mogilev-Podolsky, left bank of Derlo river, Borshchov Yar;
Vendian, Mogilev-Podolsky Group, Yaryshev Formation, lower part of Bronnitsa Beds.
Fig. 4. Pseudohiemaloraichnus podolica Gritsenko, 2009; holotype NMNH, specimen no. 2480/10 (Grit-
senko, 2009, 34 p., fig. 2, Pl. II, fig. 1, 2); Vinnitsa region, town of Mogilev-Podolsky, left bank of Derlo river,
Borshchov Yar; Vendian, Mogilev-Podolsky Group, Yaryshev Formation, lower part of Bronnitsa Beds.
Fig. 5. Studenicia galeiforma Gureev, 1983 =Monocraterion isp.; NMNH, specimen no. 2088/9 (Gureev,
1983a, Plate without number, fig. 1; 1984, fig. 1 b); Podolia, left bank of Dniester river, near Studenitsa village;
Vendian, Kanilovka Group, Studenitsa Formation, Komarovo Beds.
Fig. 6. Discoidal casts with adjoining worm-shaped body; KTSNY, specimen no. 1714 (Palij, 1976,
Pl. XXXIII, fig. 4, Palij et al., 1979, Pl. LI, fig. 1); Moldovian Dniester area, right bank of Dniester river, Otach
(Ataki) village; Vendian, Mogilev-Podolsky Group, Yaryshev Formation, Bernashevka Beds.

138

Atlas_2015.indd 138 25-04-2015 14:18

 /Plate VIII

5 6

139

Atlas_2015.indd 139 25-04-2015 14:18

 IX
. 1. Propaleolina vendiensis Menasova, 2003; , 1782 (, 2003, . 2):
, ; , . -, . , ;
, - , , .
. 2. Platysolenites antiquissimus Eichwald, 1860; . , 1731/4 (, 1968, . V,
. 21); ., - , . . , . 5, . 192,9 ; , -
, , ( ).
. 3. Vendotaenia antiqua Gnilovskaja, 1971; . , 1915/7 (, 1976, . XIX,
. 2); , . , . 350, . 63,7-71,3 ; , , , -
.
. 4. Onuphionella agglutinata Kirjanov, 1968; , 1731/6(1) (, 1968, .
VI, . 1): , ; ., , . , . 17,
. 149,0-151,7 ; , , , (
).
. 5. Sabellidites cambriensis Yanischevskii, 1926; , . 1731/9 (, 1968, . III,
. 3): , ; ., . , . 11673, . 344,9-346,9 ; -
(?), , .

Plate IX
Fig. 1. Propaleolina vendiensis Menasova, 2003; holotype KTSNY, specimen no. 1782 (Menasova, 2003b,
fig. 2): general view, b detail; Vinnitsa region, town of Mogilev-Podolsky, left bank of Derlo river, Borsh-
chov Yar; Vendian, Mogilev-Podolsky Group, Yaryshev Formation, Bronnitsa Beds.
Fig. 2. Platysolenites antiquissimus Eichwald, 1860; NMNH, specimen no. 1731/4 (Kirjanov, 1968, Pl. V,
fig. 21); Volhynia region, Kamen-Kashirsky area, B. Obzyr village, borehole no. 5, depth of 192.9 m; Cambrian,
Lontova Horizon, Baltic Group, Stokhod Beds (age analogue of Zbruch Formation of Dniester area).
Fig. 3. Vendotaenia antiqua Gnilovskaja, 1971; NMNH, specimen no. 1915/7 (Aseeva, 1976, Pl. XIX, fig. 2);
Volhynia, Grabov village, borehole no. 350, depth of 63.7-71.3 m; Vendian, Kanilovka Group, Studenitsa Forma-
tion, Komarovo Beds.
Fig. 4. Onuphionella agglutinata Kirjanov, 1968; holotype NMNH, specimen no. 1731/6(1) (Kirjanov,
1968, Pl. VI, fig. 1): detail, b general view; Volhynia region, Manevich area, Berezhnitsa village, borehole
no. 17, depth of 149.0-151.7 m; Cambrian, Lontova Horizon, Baltic Group, Stokhod Beds (age analogue of Zbruch
Formation of Dniester area).
Fig. 5. Sabellidites cambriensis Yanischevskii, 1926; NMNH, specimen no. 1731/9 (Kirjanov, 1968, Pl. III,
fig. 3): detail, b general view; Ternopol region, Gusjatin village, borehole no. 11673, depth of 344.9-346.9 m;
Cambrian(?), Roven Horizon, Baltic Group.

140

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 /Plate IX

2
4

141

Atlas_2015.indd 141 25-04-2015 14:18

 X
. 1. Beltanelloides podolicus A. Istchenko in Gnilovskaya et al., 1988; , 2235/3
( ., 1988, . I, . 3); , . -, . -
, ; , - , , .
. 2. Serebrina crustacea A. Istchenko in Gnilovskaya et al., 1988; , 2235/23 (-
., 1988, . XV, . 5); , . . ; , - -
, , .
. 3. Eoholynia capillaria A. Istchenko in Gnilovskaya et al., 1988; , 2235/15 (-
., 1988, . IV, . 3); , . . ; , - ,
, .
. 4. Pilitela composita Aseeva, 1976; , 1915/10 (, 1976, . XX, . 4;
1988, . XV, . 1); , . , . ; , --
, , .
. 5. Kalusina compacta A. Istchenko in Gnilovskaya et al., 1988;
, 2235/16
(
-
., 1988, . IV, . 4); , . . ; , - ,
, .
. 6. Fusosquamula vlasovi Aseeva, 1976; , 1915/8 (, 1976, . XX, . 1,
2; 1983, . I, . 6; 1988, . XV, . 5); , . , . 350, . 65; ,
.
. 7. Eoholynia fruticulosa A. Istchenko in Gnilovskaya et al., 1988;
, 2235/13
(
-
., 1988, . IV, . 1); , . -, . ,
; , - , , .
. 8. Tawuia dalensis Hofmann, 1979; . , 2235/9 ( ., 1988, . II, . 1);
, . . ; , - , , .
. 9. Kanilovia insolita A. Istchenko, 1983; , 2236/28 (, 1983, . XVII,
. 1); ., . ; , , , .

Plate X
Fig. 1. Beltanelloides podolicus A. Istchenko in Gnilovskaya et al., 1988; holotype NMNH, specimen
no. 2235/3 (Gnilovskaya et al., 1988, Pl. I, fig. 3); Vinnitsa region, town of Mogilev-Podolsky, left bank of Derlo
river, Borshchov Yar; Vendian, Mogilev-Podolsky Group, Yaryshev Formation, Bronnitsa Beds.
Fig. 2. Serebrina crustacea A. Istchenko in Gnilovskaya et al., 1988; holotype NMNH, specimen
no. 2235/23 (Gnilovskaya et al., 1988, Pl. XV, fig. 5); Podolia, Serebria river, near Serebria village; Vendian,
Mogilev-Podolsky Group, Yaryshev Formation, Lyadov Beds.
Fig. 3. Eoholynia capillaria A. Istchenko in Gnilovskaya et al., 1988; holotype NMNH, specimen
no. 2235/15 (Gnilovskaya et al., 1988, Pl. IV, fig. 3); Podolia, Ushitsa river, near Minkovtsy village; Vendian,
Mogilev-Podolsky Group, Nagoryany Formation, Kalyus Beds.
Fig. 4. Pilitela composita Aseeva, 1976; holotype NMNH, specimen no. 1915/10 (Aseeva, 1976, Pl. XX,
fig. 4; 1988a, Pl. XV, fig. 1); Middle Dniester area, right bank of Dniester river, Naslavche village; Vendian,
Mogilev-Podolsky Group, Nagoryany Formation, Kalyus Beds.
Fig. 5. Kalusina compacta A. Istchenko in Gnilovskaya et al., 1988; holotype NMNH, specimen no. 2235/16
(Gnilovskaya et al., 1988, Pl. IV, fig. 4); Podolia, Ushitsa river, near Minkovtsy village; Vendian, Mogilev-Podol-
sky Group, Nagoryany Formation, Kalyus Beds.
Fig. 6. Fusosquamula vlasovi Aseeva, 1976; holotype NMNH, specimen no. 1915/8 (Aseeva, 1976, Pl. XX,
fig. 1, 2; 1983, Pl. I, fig. 6; 1988, Pl. XV, fig. 5); Volhynia, Grabov village, borehole no. 350, depth of 65 m; Ven-
dian, Kanilovka Group.
Fig. 7. Eoholynia fruticulosa A. Istchenko in Gnilovskaya et al., 1988; holotype NMNH, specimen
no. 2235/13 (Gnilovskaya et al., 1988, Pl. IV, fig. 1); Vinnitsa region, town of Mogilev-Podolsky, left bank of Derlo
river, Borshchov Yar; Vendian, Mogilev-Podolsky Group, Yaryshev Formation, Bronnitsa Beds.
Fig. 8. Tawuia dalensis Hofmann, 1979; NMNH, specimen no. 2235/9 (Gnilovskaya et al., 1988, Pl. II,
fig. 1); Podolia, Ushitsa river, near Minkovtsy village; Vendian, Mogilev-Podolsky Group, Nagoryany Forma-
tion, Kalyus Beds.
Fig. 9. Kanilovia insolita A. Istchenko, 1983; holotype NMNH, specimen no. 2236/28 (Istchenko, 1983,
Pl. XVII, fig. 1); Chernovtsy region, Kuleshovka village; Vendian, Kanilovka Group, Zharnovka Formation,
Kuleshovka Beds.

142

Atlas_2015.indd 142 25-04-2015 14:18

 /Plate X

1 2

5
6

8 9

143

Atlas_2015.indd 143 25-04-2015 14:18

 144

Atlas_2015.indd 144 25-04-2015 14:18



. ..

.
/ .. . - .: , 2015.

..

: .. , ..

: ..
: ..

24 2015 .
6084/8. . . .
15 .-. ., 17 . . . 200 .
. .. ( )
, ., 123

, . , . 20, . 8
245

145

Atlas_2015.indd 145 25-04-2015 14:18

 146

Atlas_2015.indd 146 25-04-2015 14:18